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<mods:title id="20F593D8D8AB43C8AC146B10FD023D24">Late Silurian palynomorphs from the Precordillera of San Juan, Argentina: Diversity, palaeoenvironmental and palaeogeographic significance</mods:title>
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<mods:namePart id="CAE36121BB76920E20C516A03D3DD29B">Rubinstein, Claudia V.</mods:namePart>
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<treatment id="88508783FD4CFF84FFBBF8CC09B887C6" ID-DOI="http://doi.org/10.5281/zenodo.10999461" ID-Zenodo-Dep="10999461" LSID="urn:lsid:plazi:treatment:88508783FD4CFF84FFBBF8CC09B887C6" httpUri="http://treatment.plazi.org/id/88508783FD4CFF84FFBBF8CC09B887C6" lastPageNumber="46" pageId="5" pageNumber="46">
<subSubSection id="48E3651EFD4CFF84FFBBF8CC0E7986F3" box="[98,520,1893,1921]" pageId="5" pageNumber="46" type="nomenclature">
<paragraph id="00463695FD4CFF84FFBBF8CC0E7986F3" blockId="5.[98,520,1893,1921]" box="[98,520,1893,1921]" pageId="5" pageNumber="46">
<heading id="5B0E81F9FD4CFF84FFBBF8CC0E7986F3" box="[98,520,1893,1921]" fontSize="12" level="2" pageId="5" pageNumber="46" reason="2">
<taxonomicName id="C7F94D16FD4CFF84FFBBF8CC0D4A86F3" authorityName="McGregor" authorityYear="1961" box="[98,315,1893,1921]" family="Zygnemataceae" genus="Emphanisporites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Zygnematales" pageId="5" pageNumber="46" phylum="Bryophyta" rank="genus">
<emphasis id="328DEA87FD4CFF84FFBBF8CC0D4A86F3" box="[98,315,1893,1921]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
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?
<emphasis id="328DEA87FD4CFF84FE88F8CC0DEC86F3" box="[337,413,1893,1921]" italics="true" pageId="5" pageNumber="46">tenuis</emphasis>
sp. nov.
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</paragraph>
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<subSubSection id="48E3651EFD4CFF84FFBBF8380CA986D4" box="[98,216,1937,1958]" pageId="5" pageNumber="46" type="description">
<paragraph id="00463695FD4CFF84FFBBF8380CA986D4" blockId="5.[98,216,1937,1958]" box="[98,216,1937,1958]" pageId="5" pageNumber="46">
<figureCitation id="98C22A10FD4CFF84FFBBF8380CA286D4" box="[98,211,1937,1958]" captionStart="Fig" captionStartId="12.[151,184,1473,1493]" captionTargetBox="[236,1457,159,1444]" captionTargetId="figure-83@12.[230,1458,159,1448]" captionTargetPageId="12" captionText="Fig. 11. Palynomorphs from the studied sections of the Los Espejos Formation of Argentina, Quebrada Ancha, Přídolí (A, F, K, M, N), Cerro La Chilca, Gorstian?Ludfordian? (B, L), Quebrada Ancha, Ludfordian (C, E, H), Cerro La Chilca, Ludfordian (D, G, I, J). A. Tylotopalla caelamenicutis Loeblich, 1970; MPLP 14-60982 (E32/3). B. Tylotopalla deerlijkianum (Martin, 1973) Martin, 1978; MPLP 2-61057 (K42/2). C. Tylotopalla robustispinosa (Downie, 1959) Mullins, 2001; MPLP 2-60704 (L42/2). D. Veryhachium trispinosum Group Servais, Vecoli, Li, Molyneux, Raevskaya, and Rubinstein, 2007; MPLP 4-61059 (E33). E. Vestitusdyadus sp.; MPLP 8-60677 (G31/1). F. Villosacapsula setosapellicula (Loeblich, 1970) Loeblich and Tappan, 1976; MPLP 9-60664 (Q30). G. Villosacapsula sp.; MPLP 5-61060 (U32/2). H. Visbysphaera cf. pirifera (Eisenack, 1954) Kiryanov, 1978; MPLP 8-60677 (T33). I. Indeterminate palynomorphs; MPLP 4-61059 (G35/4) (I1, I2 are different focus). J. cf. Veryhachium sp., with abnormal processes, distally expanded; MPLP 4-61059 (P25/3). KN. Emphanisporites? tenuis sp. nov. K. MPLP 13-60984 (N30/2). L. MPLP 2-61057 (Q42). M. MPLP 10- 60673 (J23). N. MPLP 12-60986 (R34). Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10626559" httpUri="https://zenodo.org/record/10626559/files/figure.png" pageId="5" pageNumber="46">Fig. 11KN</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="48E3651EFD4CFF84FCF1FF0908F38013" pageId="5" pageNumber="46" type="reference_group">
<paragraph id="00463695FD4CFF84FCF1FF0909EF81C7" blockId="5.[808,1481,160,353]" box="[808,1438,160,181]" pageId="5" pageNumber="46">
<treatmentCitationGroup id="20E911BBFD4CFF84FCF1FF0909EF81C7" box="[808,1438,160,181]" pageId="5" pageNumber="46">
1981
<taxonomicName id="C7F94D16FD4CFF84FC87FF09084D81C7" box="[862,1084,160,181]" genus="Emphanisporites" kingdom="Plantae" pageId="5" pageNumber="46" phylum="Bryophyta" rank="species" species="undefined-D">
<emphasis id="328DEA87FD4CFF84FC87FF09087081C7" box="[862,1025,160,181]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
sp. D
</taxonomicName>
;
<treatmentCitation id="81581084FD4CFF84FB9FFF09092081C7" author="Richardson, J. B. &amp; Rasul, S. M. &amp; Al-Ameri, T." box="[1094,1361,160,181]" page="215" pageId="5" pageNumber="46" year="1981">
<bibRefCitation id="64684B64FD4CFF84FB9FFF09092081C7" author="Richardson, J. B. &amp; Rasul, S. M. &amp; Al-Ameri, T." box="[1094,1361,160,181]" pageId="5" pageNumber="46" pagination="209 - 224" refId="ref15311" refString="Richardson, J. B., Rasul, S. M., and Al-Ameri, T. 1981. Acritarchs, miospores and correlation of the Ludlovian - Downtonian and Silurian - Devonian boundaries. Review of Palaeobotany and Palynology 34: 209 - 224." type="journal article" year="1981">Richardson et al. 1981: 215</bibRefCitation>
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, pl. 1.3.
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</paragraph>
<paragraph id="00463695FD4CFF84FCF1FF1508B7819C" blockId="5.[808,1481,160,353]" pageId="5" pageNumber="46">
<treatmentCitationGroup id="20E911BBFD4CFF84FCF1FF1508B7819C" pageId="5" pageNumber="46">
2008
<taxonomicName id="C7F94D16FD4CFF84FC86FF15084A81A3" box="[863,1083,188,209]" genus="Emphanisporites" kingdom="Plantae" pageId="5" pageNumber="46" phylum="Bryophyta" rank="species" species="undefined-D">
<emphasis id="328DEA87FD4CFF84FC86FF15087381A3" box="[863,1026,188,209]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
sp. D
</taxonomicName>
<bibRefCitation id="64684B64FD4CFF84FB9BFF1509B481A3" author="Richardson, J. B. &amp; Rasul, S. M. &amp; Al-Ameri, T." box="[1090,1477,188,209]" pageId="5" pageNumber="46" pagination="209 - 224" refId="ref15311" refString="Richardson, J. B., Rasul, S. M., and Al-Ameri, T. 1981. Acritarchs, miospores and correlation of the Ludlovian - Downtonian and Silurian - Devonian boundaries. Review of Palaeobotany and Palynology 34: 209 - 224." type="journal article" year="1981">Richardson, Rasul, and Al-Ameri, 1981</bibRefCitation>
;
<treatmentCitation id="81581084FD4CFF84FC85FF700827819C" author="Steemans, P. &amp; Rubinstein, C. V. &amp; de Melo, J. H. G." box="[860,1110,217,238]" page="274" pageId="5" pageNumber="46" year="2008">
<bibRefCitation id="64684B64FD4CFF84FC85FF700827819C" author="Steemans, P. &amp; Rubinstein, C. V. &amp; de Melo, J. H. G." box="[860,1110,217,238]" pageId="5" pageNumber="46" pagination="263 - 282" refId="ref16499" refString="Steemans, P., Rubinstein, C. V., and de Melo, J. H. G. 2008. Siluro-Devonian miospore biostratigraphy of the Urubu River area, western Amazon Basin, northern Brazil. Geobios 41: 263 - 282." type="journal article" year="2008">Steemans et al. 2008: 274</bibRefCitation>
</treatmentCitation>
, figs. 810.
</treatmentCitationGroup>
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FF5F08458055" blockId="5.[808,1481,160,353]" pageId="5" pageNumber="46">
<treatmentCitationGroup id="20E911BBFD4CFF84FCF1FF5F08458055" pageId="5" pageNumber="46">
2014
<taxonomicName id="C7F94D16FD4CFF84FCB9FF5F084E8079" box="[864,1087,246,267]" genus="Emphanisporites" kingdom="Plantae" pageId="5" pageNumber="46" phylum="Bryophyta" rank="species" species="undefined-D">
<emphasis id="328DEA87FD4CFF84FCB9FF5F08728079" box="[864,1027,246,267]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
sp. D
</taxonomicName>
<bibRefCitation id="64684B64FD4CFF84FB9EFF5F09568079" author="Richardson, J. B. &amp; Rasul, S. M. &amp; Al-Ameri, T." box="[1095,1319,246,267]" pageId="5" pageNumber="46" pagination="209 - 224" refId="ref15311" refString="Richardson, J. B., Rasul, S. M., and Al-Ameri, T. 1981. Acritarchs, miospores and correlation of the Ludlovian - Downtonian and Silurian - Devonian boundaries. Review of Palaeobotany and Palynology 34: 209 - 224." type="journal article" year="1981">Richardson et al. 1981</bibRefCitation>
;
<treatmentCitation id="81581084FD4CFF84FAEAFF5F0F998055" author="Garcia Muro, V. J. &amp; Rubinstein, C. V. &amp; Steemans, P." page="485" pageId="5" pageNumber="46" year="2014">
<bibRefCitation id="64684B64FD4CFF84FAEAFF5F0F998055" author="Garcia Muro, V. J. &amp; Rubinstein, C. V. &amp; Steemans, P." pageId="5" pageNumber="46" pagination="472 - 490" refId="ref12965" refString="Garcia Muro, V. J., Rubinstein, C. V., and Steemans, P. 2014 b. Upper Silurian miospores from the Precordillera Argentina: biostratigraphic, palaeonvironmental and palaeogeographic implications. Geological Magazine 151: 472 - 490." type="journal article" year="2014">García Muro et al. 2014b: 485</bibRefCitation>
</treatmentCitation>
, fig.
<specimenCount id="16FFFD1CFD4CFF84FBC0FEBB08458055" box="[1049,1076,274,295]" pageId="5" pageNumber="46" type="generic">8f.</specimenCount>
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</paragraph>
<paragraph id="00463695FD4CFF84FCF1FE8608F38013" blockId="5.[808,1481,160,353]" pageId="5" pageNumber="46">
<treatmentCitationGroup id="20E911BBFD4CFF84FCF1FE8608F38013" pageId="5" pageNumber="46">
2015
<taxonomicName id="C7F94D16FD4CFF84FC87FE8608488036" box="[862,1081,303,324]" genus="Emphanisporites" kingdom="Plantae" pageId="5" pageNumber="46" phylum="Bryophyta" rank="species" species="undefined-D">
<emphasis id="328DEA87FD4CFF84FC87FE8608708036" box="[862,1025,303,324]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
sp. D
</taxonomicName>
<bibRefCitation id="64684B64FD4CFF84FB99FE86096A8036" author="Richardson, J. B. &amp; Rasul, S. M. &amp; Al-Ameri, T." box="[1088,1307,303,324]" pageId="5" pageNumber="46" pagination="209 - 224" refId="ref15311" refString="Richardson, J. B., Rasul, S. M., and Al-Ameri, T. 1981. Acritarchs, miospores and correlation of the Ludlovian - Downtonian and Silurian - Devonian boundaries. Review of Palaeobotany and Palynology 34: 209 - 224." type="journal article" year="1981">Richardson et al. 1981</bibRefCitation>
;
<treatmentCitation id="81581084FD4CFF84FAFCFE86085C8013" author="Garcia Muro, V. J. &amp; Rubinstein, C. V." page="272" pageId="5" pageNumber="46" year="2015">
<bibRefCitation id="64684B64FD4CFF84FAFCFE86085C8013" author="Garcia Muro, V. J. &amp; Rubinstein, C. V." pageId="5" pageNumber="46" pagination="265 - 285" refId="ref12808" refString="Garcia Muro, V. J. and Rubinstein, C. V. 2015. New biostratigraphic proposal for the Lower Palaeozoic Tucunuco Group (San Juan Precordillera, Argentina) based on marine and terrestrial palynomorphs. Ameghiniana 52: 265 - 285." type="journal article" year="2015">García Muro and Rubinstein 2015: 272</bibRefCitation>
</treatmentCitation>
, fig. 3.7.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="48E3651EFD4CFF84FCF1FEC709B887C6" pageId="5" pageNumber="46" type="etymology">
<paragraph id="00463695FD4CFF84FCF1FEC70FD780D2" blockId="5.[808,1481,366,416]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FEC70FFE80F1" box="[808,911,366,387]" italics="true" pageId="5" pageNumber="46">Etymology</emphasis>
: From Latin
<emphasis id="328DEA87FD4CFF84FBD5FEC7083480F1" box="[1036,1093,366,387]" italics="true" pageId="5" pageNumber="46">tenuis</emphasis>
, tenuous; in references to the characteristic thin ribs.
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FE0409CE80AC" blockId="5.[808,1481,429,478]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FE040FDF80B0" box="[808,942,429,450]" italics="true" pageId="5" pageNumber="46">Type material</emphasis>
:
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<typeStatus id="DF428837FD4CFF84FC64FE04086B80B0" box="[957,1050,429,450]" pageId="5" pageNumber="46">Holotype</typeStatus>
:
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(
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/2) (
<figureCitation id="98C22A10FD4CFF84FAF0FE0409F280B0" box="[1321,1411,429,450]" captionStart="Fig" captionStartId="12.[151,184,1473,1493]" captionTargetBox="[236,1457,159,1444]" captionTargetId="figure-83@12.[230,1458,159,1448]" captionTargetPageId="12" captionText="Fig. 11. Palynomorphs from the studied sections of the Los Espejos Formation of Argentina, Quebrada Ancha, Přídolí (A, F, K, M, N), Cerro La Chilca, Gorstian?Ludfordian? (B, L), Quebrada Ancha, Ludfordian (C, E, H), Cerro La Chilca, Ludfordian (D, G, I, J). A. Tylotopalla caelamenicutis Loeblich, 1970; MPLP 14-60982 (E32/3). B. Tylotopalla deerlijkianum (Martin, 1973) Martin, 1978; MPLP 2-61057 (K42/2). C. Tylotopalla robustispinosa (Downie, 1959) Mullins, 2001; MPLP 2-60704 (L42/2). D. Veryhachium trispinosum Group Servais, Vecoli, Li, Molyneux, Raevskaya, and Rubinstein, 2007; MPLP 4-61059 (E33). E. Vestitusdyadus sp.; MPLP 8-60677 (G31/1). F. Villosacapsula setosapellicula (Loeblich, 1970) Loeblich and Tappan, 1976; MPLP 9-60664 (Q30). G. Villosacapsula sp.; MPLP 5-61060 (U32/2). H. Visbysphaera cf. pirifera (Eisenack, 1954) Kiryanov, 1978; MPLP 8-60677 (T33). I. Indeterminate palynomorphs; MPLP 4-61059 (G35/4) (I1, I2 are different focus). J. cf. Veryhachium sp., with abnormal processes, distally expanded; MPLP 4-61059 (P25/3). KN. Emphanisporites? tenuis sp. nov. K. MPLP 13-60984 (N30/2). L. MPLP 2-61057 (Q42). M. MPLP 10- 60673 (J23). N. MPLP 12-60986 (R34). Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10626559" httpUri="https://zenodo.org/record/10626559/files/figure.png" pageId="5" pageNumber="46">Fig. 11K</figureCitation>
)
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.
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<typeStatus id="DF428837FD4CFF84FA4FFE040F2480AC" pageId="5" pageNumber="46">Paratype</typeStatus>
:
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(
<specimenCode id="505F9EEEFD4CFF84FC26FE60085A80AC" box="[1023,1067,457,478]" collectionCode="Q" country="Ecuador" lsid="urn:lsid:biocol.org:col:14273" name="Universidad Central" pageId="5" pageNumber="46" type="Herbarium">Q42</specimenCode>
) (
<figureCitation id="98C22A10FD4CFF84FBE7FE6008E380AC" box="[1086,1170,457,478]" captionStart="Fig" captionStartId="12.[151,184,1473,1493]" captionTargetBox="[236,1457,159,1444]" captionTargetId="figure-83@12.[230,1458,159,1448]" captionTargetPageId="12" captionText="Fig. 11. Palynomorphs from the studied sections of the Los Espejos Formation of Argentina, Quebrada Ancha, Přídolí (A, F, K, M, N), Cerro La Chilca, Gorstian?Ludfordian? (B, L), Quebrada Ancha, Ludfordian (C, E, H), Cerro La Chilca, Ludfordian (D, G, I, J). A. Tylotopalla caelamenicutis Loeblich, 1970; MPLP 14-60982 (E32/3). B. Tylotopalla deerlijkianum (Martin, 1973) Martin, 1978; MPLP 2-61057 (K42/2). C. Tylotopalla robustispinosa (Downie, 1959) Mullins, 2001; MPLP 2-60704 (L42/2). D. Veryhachium trispinosum Group Servais, Vecoli, Li, Molyneux, Raevskaya, and Rubinstein, 2007; MPLP 4-61059 (E33). E. Vestitusdyadus sp.; MPLP 8-60677 (G31/1). F. Villosacapsula setosapellicula (Loeblich, 1970) Loeblich and Tappan, 1976; MPLP 9-60664 (Q30). G. Villosacapsula sp.; MPLP 5-61060 (U32/2). H. Visbysphaera cf. pirifera (Eisenack, 1954) Kiryanov, 1978; MPLP 8-60677 (T33). I. Indeterminate palynomorphs; MPLP 4-61059 (G35/4) (I1, I2 are different focus). J. cf. Veryhachium sp., with abnormal processes, distally expanded; MPLP 4-61059 (P25/3). KN. Emphanisporites? tenuis sp. nov. K. MPLP 13-60984 (N30/2). L. MPLP 2-61057 (Q42). M. MPLP 10- 60673 (J23). N. MPLP 12-60986 (R34). Scale bars 20 μm." figureDoi="http://doi.org/10.5281/zenodo.10626559" httpUri="https://zenodo.org/record/10626559/files/figure.png" pageId="5" pageNumber="46">Fig. 11L</figureCitation>
)
</materialsCitation>
<materialsCitation id="B0913CC8FD4CFF84FB46FE6009CA80AC" box="[1183,1467,457,478]" collectionCode="Q" location="Cerro La Chilca" pageId="5" pageNumber="46" specimenCount="1" typeStatus="paratype">
from
<location id="0526604EFD4CFF84FB0DFE60091E80AC" LSID="urn:lsid:plazi:treatment:88508783FD4CFF84FFBBF8CC09B887C6:0526604EFD4CFF84FB0DFE60091E80AC" box="[1236,1391,457,478]" name="Cerro La Chilca" pageId="5" pageNumber="46">Cerro La Chilca</location>
section
</materialsCitation>
.
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FE420817836F" blockId="5.[808,1481,491,541]" pageId="5" pageNumber="46">
<materialsCitation id="B0913CC8FD4CFF84FCF1FE420813836F" country="Argentina" location="Quebrada Ancha" municipality="Los Espejos Formation" pageId="5" pageNumber="46" specimenCount="1" stateProvince="San Juan" typeStatus="holotype">
<emphasis id="328DEA87FD4CFF84FCF1FE420FD28372" box="[808,931,491,512]" italics="true" pageId="5" pageNumber="46">
<typeStatus id="DF428837FD4CFF84FCF1FE420F248372" box="[808,853,491,512]" pageId="5" pageNumber="46">Type</typeStatus>
locality
</emphasis>
:
<location id="0526604EFD4CFF84FC69FE4208208372" LSID="urn:lsid:plazi:treatment:88508783FD4CFF84FFBBF8CC09B887C6:0526604EFD4CFF84FC69FE4208208372" box="[944,1105,491,512]" country="Argentina" municipality="Los Espejos Formation" name="Quebrada Ancha" pageId="5" pageNumber="46" stateProvince="San Juan">Quebrada Ancha</location>
upper section,
<collectingMunicipality id="E022ACEFFD4CFF84FB3DFE4209B78372" box="[1252,1478,491,512]" pageId="5" pageNumber="46">Los Espejos Formation</collectingMunicipality>
,
<collectingRegion id="C23DF877FD4CFF84FCF1FDA10FF1836F" box="[808,896,520,541]" country="Argentina" name="San Juan" pageId="5" pageNumber="46">San Juan</collectingRegion>
,
<collectingCountry id="78EE7605FD4CFF84FC50FDA10F98836F" box="[905,1001,520,541]" name="Argentina" pageId="5" pageNumber="46">Argentina</collectingCountry>
(
<figureCitation id="98C22A10FD4CFF84FC2CFDA10830836F" box="[1013,1089,520,541]" captionStart="Fig" captionStartId="1.[98,131,1412,1432]" captionTargetId="figure-306@1.[101,1479,165,1382]" captionTargetPageId="1" captionText="Fig. 1. A, B. Geographic location of study areas. C, D. Geologic maps showing the fossil localities (asterisks). The northern and middle (C) and southern D) part of the SilurianDevonian Basin. 1, Río Jáchal; 2, Río de las Chacritas; 3, Cerro La Chilca (this paper); 4, Quebrada Ancha (this paper)." figureDoi="http://doi.org/10.5281/zenodo.10999443" httpUri="https://zenodo.org/record/10999443/files/figure.png" pageId="5" pageNumber="46">Fig. 1D</figureCitation>
,
<figureCitation id="98C22A10FD4CFF84FB95FDA1082B836F" box="[1100,1114,520,541]" captionStart="Fig" captionStartId="4.[862,895,921,941]" captionTargetBox="[838,1532,161,911]" captionTargetId="figure-658@4.[864,1533,161,829]" captionTargetPageId="4" captionText="Fig. 4. LudlowPřídolí (424 My) phytoplankton and miospore assemblages plotted on a palaoegeographic reconstruction using BugPlate software (Torsvik 2009). 1, UK (Richardson and Lister 1969; Lister 1970; Richardson et al. 1981; Burgess and Richardson 1995; Wellman and Richardson 1996; Richards and Mullins 2003; Mullins 2001, 2004); 2, Gotland, Sweeden (Stricanne et al. 2004; Stricanne et al. 2006); 3, Allenport, USA (Beck and Strother 2008); 4, Nova Scotia, Canada (Beck and Strother 2001); 5, Ontario, Canada (McGregor and Camfield 1976); 6, Sweden (Mehlqvist et al. 2012; Mehlqvist et al. 2014); 7, Libya (Richardson and Ioannides 1973; Rubinstein and Steemans 2002; Le Hérissé 2002); 8, Saudia Arabia (Le Hérissé et al. 1995; Wellman et al. 2000; Breuer et al. 2017); 9, Poland (Porębska et al. 2004); 10, Turkey (Steemans et al. 1996; Lakova and Göncüoğlu 2005); 11, Iraq (Al-Almeri 1984; 2010); 12, Maranhão Basin, Brazil (Brito 1967); 13, Bolivia (Cramer et al. 1974; Kimyai 1983; Racheboeuf et al. 2012); 14, Spain and West France (Rodríguez González 1978; 1983; Steemans 1989; Cramer 1964a; Richardson et al. 2001); 15, Amazon Basin, Brazil (Cardoso 2005; Steemans et al. 2008); 16, Santiago del Estero, Argentina (Pöthe de Baldis 1974a); 17, Precordillera Argentina (Rubinstein 1992a, b, 1995; García Muro et al. 2014b; this contribution). References discussed in the papers are also included." figureDoi="http://doi.org/10.5281/zenodo.10626543" httpUri="https://zenodo.org/record/10626543/files/figure.png" pageId="5" pageNumber="46">4</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FD830840832E" blockId="5.[808,1481,554,604]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FD830FD9834D" box="[808,936,554,575]" italics="true" pageId="5" pageNumber="46">
<typeStatus id="DF428837FD4CFF84FCF1FD830F24834D" box="[808,853,554,575]" pageId="5" pageNumber="46">Type</typeStatus>
horizon
</emphasis>
: Storm-dominated inner shelf environment of the Los Espejos Formation, Přídolí.
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FDC609198279" blockId="5.[808,1481,623,779]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FDC60FDD83FA" box="[808,940,623,648]" italics="true" pageId="5" pageNumber="46">Material.—</emphasis>
Seven specimens measured. Samples MPLP 1-61065 (O28/3); MPLP 2-61057 (O47/1; Q42/1, U27); MPLP 2-60704 (K38/4), MPLP 9-60664 (S28), MPLP 10-60673 (J23/1). Wenlock?Přídolí of
<collectingCountry id="78EE7605FD4CFF84FBB0FD7B08AC8399" box="[1129,1245,722,747]" name="Argentina" pageId="5" pageNumber="46">Argentina</collectingCountry>
, Cerro La Chilca and Quebrada Ancha localities, Los Espejos Formation.
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FCB7083B8225" blockId="5.[808,1481,798,855]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FCB70FEB8245" box="[808,922,798,823]" italics="true" pageId="5" pageNumber="46">Diagnosis</emphasis>
.—Trilete patinate spore with numerous tenuous ribs on the proximal face.
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FCC3090E8554" blockId="5.[808,1481,874,1062]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FCC30FDF82F1" box="[808,942,874,899]" italics="true" pageId="5" pageNumber="46">Description</emphasis>
.—Trilete patinate spore with subtriangular to subcircular amb. The thickness of the patina is variable. Laesure straight, extends to the equator, accompanied by labra about 24 μm in overall width. Equatorially thickened, 24 μm wide. Proximally, 1826 ribs of less than 1μm of width per interradial area, extending to the patina.
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FB9009E48520" blockId="5.[808,1429,1081,1106]" box="[808,1429,1081,1106]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FB900FDC8520" box="[808,941,1081,1106]" italics="true" pageId="5" pageNumber="46">Dimensions</emphasis>
.—42(50)55μm (
<specimenCount id="16FFFD1CFD4CFF84FB5FFB9009608520" box="[1158,1297,1081,1106]" pageId="5" pageNumber="46" type="generic">7 specimens</specimenCount>
measured).
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FBCD086284B6" blockId="5.[808,1481,1124,1476]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FBCD0FFB850F" box="[808,906,1124,1149]" italics="true" pageId="5" pageNumber="46">Remarks</emphasis>
.—Other species of
<taxonomicName id="C7F94D16FD4CFF84FBB6FBCD095E850F" authorityName="McGregor" authorityYear="1961" box="[1135,1327,1124,1149]" genus="Emphanisporites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="5" pageNumber="46" phylum="Bryophyta" rank="genus">
<emphasis id="328DEA87FD4CFF84FBB6FBCD095E850F" box="[1135,1327,1124,1149]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
</taxonomicName>
differ in presenting broader radial ridges and having no patina.
<taxonomicName id="C7F94D16FD4CFF84FA56FB2C08FD85AD" authority="Breuer, Al-Ghazi, Al-Ruwaili, Higgs, Steemans, and Wellman, 2007" authorityName="Breuer, Al-Ghazi, Al-Ruwaili, Higgs, Steemans, and Wellman" authorityYear="2007" genus="Artemopyra" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="5" pageNumber="46" rank="species" species="inconspicua">
<emphasis id="328DEA87FD4CFF84FA56FB2C086185CC" italics="true" pageId="5" pageNumber="46">Artemopyra inconspicua</emphasis>
Breuer, Al-Ghazi, Al-Ruwaili, Higgs,
<bibRefCitation id="64684B64FD4CFF84FCF1FB6F08FD85AD" author="Steemans, P. &amp; Wellman, C. H. &amp; Filatoff, J." box="[808,1164,1222,1247]" pageId="5" pageNumber="46" pagination="237 - 254" refId="ref16546" refString="Steemans, P., Wellman, C. H., and Filatoff, J. 2007. Palaeophytogeographical and palaeoecological implications of a miospore assemblage of earliest Devonian (Lochkovian) age from Saudi Arabia. Palaeogeography, Palaeoclimatology, Palaeoecology 250: 237 - 254." type="journal article" year="2007">Steemans, and Wellman, 2007</bibRefCitation>
</taxonomicName>
is ornamented by similar thin radial ribs in the hilum but lacks the trilete mark and the patina. According to the original diagnosis of
<bibRefCitation id="64684B64FD4CFF84FA8DFAA10F1E8433" author="McGregor, D. C." pageId="5" pageNumber="46" pagination="1 - 11" refId="ref14327" refString="McGregor, D. C. 1961. Spores with proximal radial pattern from the Devoni- an of Canada. Geological Survey of Canada Bulletin 76: 1 - 11." type="journal article" year="1961">McGregor (1961)</bibRefCitation>
, the genus
<taxonomicName id="C7F94D16FD4CFF84FC20FA8108C88433" authorityName="McGregor" authorityYear="1961" box="[1017,1209,1320,1345]" genus="Emphanisporites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="5" pageNumber="46" phylum="Bryophyta" rank="genus">
<emphasis id="328DEA87FD4CFF84FC20FA8108C88433" box="[1017,1209,1320,1345]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
</taxonomicName>
does not comprise patinate forms. Possibly, a new genus should be considered for the specimens included in
<taxonomicName id="C7F94D16FD4CFF84FB51FAC3093684F1" authorityName="McGregor" authorityYear="1961" box="[1160,1351,1386,1411]" genus="Emphanisporites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="5" pageNumber="46" phylum="Bryophyta" rank="genus">
<emphasis id="328DEA87FD4CFF84FB51FAC3093684F1" box="[1160,1351,1386,1411]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
</taxonomicName>
?
<emphasis id="328DEA87FD4CFF84FA87FAC309D384F1" box="[1374,1442,1386,1411]" italics="true" pageId="5" pageNumber="46">tenuis</emphasis>
although the specimens are not enough to create it in the present contribution.
</paragraph>
<paragraph id="00463695FD4CFF84FCF1FA7F09B887C6" blockId="5.[808,1482,1494,1716]" pageId="5" pageNumber="46">
<emphasis id="328DEA87FD4CFF84FCF1FA7F0888849D" box="[808,1273,1494,1519]" italics="true" pageId="5" pageNumber="46">Stratigraphic and geographic range.—</emphasis>
<bibRefCitation id="64684B64FD4CFF84FB23FA7F0F1C8762" author="Richardson, J. B. &amp; Rasul, S. M. &amp; Al-Ameri, T." pageId="5" pageNumber="46" pagination="209 - 224" refId="ref15311" refString="Richardson, J. B., Rasul, S. M., and Al-Ameri, T. 1981. Acritarchs, miospores and correlation of the Ludlovian - Downtonian and Silurian - Devonian boundaries. Review of Palaeobotany and Palynology 34: 209 - 224." type="journal article" year="1981">Richardson et al. (1981)</bibRefCitation>
recorded
<taxonomicName id="C7F94D16FD4CFF84FC03FA5E08A68762" box="[986,1239,1527,1552]" genus="Emphanisporites" kingdom="Plantae" pageId="5" pageNumber="46" phylum="Bryophyta" rank="species" species="undefined-D">
<emphasis id="328DEA87FD4CFF84FC03FA5E08E78762" box="[986,1174,1527,1552]" italics="true" pageId="5" pageNumber="46">Emphanisporites</emphasis>
sp. D
</taxonomicName>
from the late Ludlow of
<collectingCountry id="78EE7605FD4CFF84FC9EF9B10FF68743" box="[839,903,1560,1585]" name="Libya" pageId="5" pageNumber="46">Libya</collectingCountry>
and the early Přídolí of
<collectingCountry id="78EE7605FD4CFF84FB43F9B109038743" box="[1178,1394,1560,1585]" name="United Kingdom" pageId="5" pageNumber="46">England and Wales</collectingCountry>
but did not describe the species and the latter was not even illustrated. Lochkovian of the Amazon Basin,
<collectingCountry id="78EE7605FD4CFF84FB1BF9F009738700" box="[1218,1282,1625,1650]" name="Brazil" pageId="5" pageNumber="46">Brazil</collectingCountry>
(
<bibRefCitation id="64684B64FD4CFF84FAC1F9F00F2E87E1" author="Steemans, P. &amp; Rubinstein, C. V. &amp; de Melo, J. H. G." pageId="5" pageNumber="46" pagination="263 - 282" refId="ref16499" refString="Steemans, P., Rubinstein, C. V., and de Melo, J. H. G. 2008. Siluro-Devonian miospore biostratigraphy of the Urubu River area, western Amazon Basin, northern Brazil. Geobios 41: 263 - 282." type="journal article" year="2008">Steemans et al. 2008</bibRefCitation>
). Wenlock? to Přídolí of the Precordillera,
<collectingCountry id="78EE7605FD4CFF84FA80F9D309B887E1" box="[1369,1481,1658,1683]" name="Argentina" pageId="5" pageNumber="46">Argentina</collectingCountry>
(
<bibRefCitation id="64684B64FD4CFF84FCF6F932084E87C6" author="Garcia Muro, V. J. &amp; Rubinstein, C. V. &amp; Steemans, P." box="[815,1087,1691,1716]" pageId="5" pageNumber="46" pagination="472 - 490" refId="ref12965" refString="Garcia Muro, V. J., Rubinstein, C. V., and Steemans, P. 2014 b. Upper Silurian miospores from the Precordillera Argentina: biostratigraphic, palaeonvironmental and palaeogeographic implications. Geological Magazine 151: 472 - 490." type="journal article" year="2014">García Muro et al. 2014b</bibRefCitation>
;
<bibRefCitation id="64684B64FD4CFF84FB91F93209CF87C6" author="Garcia Muro, V. J. &amp; Rubinstein, C. V." box="[1096,1470,1691,1716]" pageId="5" pageNumber="46" pagination="265 - 285" refId="ref12808" refString="Garcia Muro, V. J. and Rubinstein, C. V. 2015. New biostratigraphic proposal for the Lower Palaeozoic Tucunuco Group (San Juan Precordillera, Argentina) based on marine and terrestrial palynomorphs. Ameghiniana 52: 265 - 285." type="journal article" year="2015">García Muro and Rubinstein 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>