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<document id="5D84D5D6C0E542007103232CC9ED7B1D" ID-DOI="10.5281/zenodo.5415636" ID-ISSN="0253-116X" ID-Zenodo-Dep="5415636" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630358179373" checkinUser="marcus" docAuthor="Smetana, Aleš" docDate="2015" docId="6824690DF805FFD7FF6FFA0D3EBD803A" docLanguage="en" docName="LinzerbiolBeitr.2015.47.1.905-924.pdf" docOrigin="Linzer biologische Beiträge 47 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Quedius (Distichalius) causarius Smetana 2015, nov.sp." docType="treatment" docVersion="3" lastPageNumber="911" masterDocId="941D1175F800FFD1FFDDFF803F6D860D" masterDocTitle="Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 51. Genus Quedius STEPHENS, 1829. Subgenus Distichalius CASEY, 1915. Section 4" masterLastPageNumber="924" masterPageNumber="905" pageNumber="910" updateTime="1699246506670" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="4C225AF3B92B4CD134097BE2466A2762" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4ABF3768043733FB73312ABF1E733EE8">
<mods:title id="D90F36F2548651BD1DA8B5C3E9D59416">Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 51. Genus Quedius STEPHENS, 1829. Subgenus Distichalius CASEY, 1915. Section 4</mods:title>
</mods:titleInfo>
<mods:name id="E55DEC420097A4BC9FB88384D2C03AA0" type="personal">
<mods:role id="FC31F5E222F13950CA0E100F7060ADF1">
<mods:roleTerm id="A073C32C5C7C43FFDEFC4477F478834A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8592A1C4638F9DF85556A4F8E95E3D14">Smetana, Aleš</mods:namePart>
</mods:name>
<mods:typeOfResource id="5370CD17C237C7D972CBCEF13A253C41">text</mods:typeOfResource>
<mods:relatedItem id="121512CE2695320F79E1ED3402D0DBBF" type="host">
<mods:titleInfo id="5965604870574D83C9F15E59B2CB793E">
<mods:title id="E76C366936ADD284F9C3C618601E8DE2">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="4DEA926BDA43BF12A7B827E7C601009F">
<mods:date id="FE3853825713C8EF749176454DD28B51">2015</mods:date>
<mods:detail id="18B28669F224632E60483783AD9162D8" type="pubDate">
<mods:number id="0B4349378F81021C8A8F7DF01B5A51BA">2015-07-31</mods:number>
</mods:detail>
<mods:detail id="38B82B1AF509FBB52AE276B0D1961C24" type="volume">
<mods:number id="98EC04C97ED63E0D6E453FB4A3C65345">47</mods:number>
</mods:detail>
<mods:detail id="801747CBD2591854CB118A8BB025B55D" type="issue">
<mods:number id="81A8ABF2ED837E69457BCF04CE93D6F1">1</mods:number>
</mods:detail>
<mods:extent id="B187F398877C821411AD75E739741913" unit="page">
<mods:start id="5CB1388718C53867FA947129117ADF9C">905</mods:start>
<mods:end id="B075D49608D8E4DDF6D02C72C98A91CF">924</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="ADD682696F930F2A5F519DD23C372EC6">journal article</mods:classification>
<mods:identifier id="34CBC3D591031BEA4DA483F5286E2691" type="DOI">10.5281/zenodo.5415636</mods:identifier>
<mods:identifier id="E970F4D7BA5D1B8CC1F18C411765B803" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="3352B3336F42C632532030A439DE4A79" type="Zenodo-Dep">5415636</mods:identifier>
</mods:mods>
<treatment id="6824690DF805FFD7FF6FFA0D3EBD803A" LSID="urn:lsid:plazi:treatment:6824690DF805FFD7FF6FFA0D3EBD803A" httpUri="http://treatment.plazi.org/id/6824690DF805FFD7FF6FFA0D3EBD803A" lastPageId="6" lastPageNumber="911" pageId="5" pageNumber="910">
<subSubSection id="A8978B90F805FFD4FF6FFA0D3B5383C5" pageId="5" pageNumber="910" type="nomenclature">
<paragraph id="E032D81BF805FFD4FF6FFA0D3D8C83A9" blockId="5.[178,737,1421,1444]" box="[178,737,1421,1444]" pageId="5" pageNumber="910">
<heading id="BB7A6F77F805FFD4FF6FFA0D3D8C83A9" box="[178,737,1421,1444]" fontSize="8" level="2" pageId="5" pageNumber="910" reason="2">
<emphasis id="D2F90409F805FFD4FF6FFA0D3D8C83A9" bold="true" box="[178,737,1421,1444]" pageId="5" pageNumber="910">
<taxonomicName id="278DA398F805FFD4FF6FFA0D3D6483AE" ID-CoL="4R2FG" authority="Smetana, 2015" authorityName="Smetana" authorityYear="2015" box="[178,521,1421,1444]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="910" phylum="Arthropoda" rank="species" species="causarius" status="sp. nov." subGenus="Distichalius">
<emphasis id="D2F90409F805FFD4FF6FFA0D3E6483AE" bold="true" box="[178,265,1421,1443]" italics="true" pageId="5" pageNumber="910">Quedius</emphasis>
(
<emphasis id="D2F90409F805FFD4FEC5FA0D3EF883AE" bold="true" box="[280,405,1421,1443]" italics="true" pageId="5" pageNumber="910">Distichalius</emphasis>
)
<emphasis id="D2F90409F805FFD4FE79FA0E3D6483AE" bold="true" box="[420,521,1422,1443]" italics="true" pageId="5" pageNumber="910">causarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9CAB972F805FFD4FDD2FA0E3D3683AE" box="[527,603,1422,1443]" pageId="5" pageNumber="910" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="78B6C49EF805FFD4FDB5FA0E3DB483A9" box="[616,729,1422,1444]" captionStart-0="Figs 11-19" captionStart-1="Figs 20-30" captionStartId-0="17.[178,218,1473,1495]" captionStartId-1="18.[178,218,1446,1468]" captionTargetBox-0="[179,1082,213,1461]" captionTargetBox-1="[197,1066,218,1437]" captionTargetId-0="figure-1@17.[178,1084,212,1464]" captionTargetId-1="figure-1@18.[197,1067,212,1437]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figs 11-19: 11-14: Quedius bipictus sp. nov.: (11) aedoeagus, ventral view; (12) apical portion of aedoeagus, lateral view; (13) apical portion of underside of paramere with sensory peg setae; (14) tergite 10 of female genital segment. Quedius gynaikos sp. nov.: (15) tergite 10 of female genital segment. 16-19: Quedius causarius sp. nov.: (16) apical portion of male sternite 8; (17) tergite 10 of male genital segment; (18) sternite 9 of male genital segment; (19) aedoeagus, ventral view." captionText-1="Figs 20-30: 20-23: Quedius causarius sp. nov.: (20) apical portion of aedoeagus, lateral view; (21) apical portion of median lobe, paramere removed; (22) apical portion of underside of paramere with sensory peg setae; (23) tergite 10 of female genital segment. 24-29: Quedius laetepictus sp. nov.: (24) apical portion of male sternite 8; (25) tergite 10 of male genital segment; (26) sternite 9 of male genital segment; (27) aedoeagus, ventral view; (28) apical portion of median lobe, paramere removed; (29) apical portion of underside of paramere with sensory peg setae. Quedius iaculifer sp. nov.: (30) apical portion of male sternite 8." figureDoi-0="http://doi.org/10.5281/zenodo.5415644" figureDoi-1="http://doi.org/10.5281/zenodo.5415648" httpUri-0="https://zenodo.org/record/5415644/files/figure.png" httpUri-1="https://zenodo.org/record/5415648/files/figure.png" pageId="5" pageNumber="910">Figs 16-23</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="E032D81BF805FFD4FF6FFA333B5383C5" blockId="5.[178,1086,1459,1611]" box="[178,1086,1459,1481]" pageId="5" pageNumber="910">
T y p e l o c a l i t y
<collectingCountry id="989A988BF805FFD4FE78FA333EBA83C4" box="[421,471,1459,1481]" name="China" pageId="5" pageNumber="910">China</collectingCountry>
:
<collectingRegion id="224916F9F805FFD4FE36FA333D5B83C4" box="[491,566,1459,1481]" country="China" name="Qinghai" pageId="5" pageNumber="910">Qinghai</collectingRegion>
, Lang Shi Dang Jing Qu (Park),
<quantity id="277575FEF805FFD4FCA4FA333CDB83C4" box="[889,950,1459,1481]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="910" unit="km" value="75.0">75 km</quantity>
NW Honggu,
</paragraph>
</subSubSection>
<subSubSection id="A8978B90F805FFD7FF6FFA493EBD803A" lastPageId="6" lastPageNumber="911" pageId="5" pageNumber="910" type="description">
<paragraph id="E032D81BF805FFD4FF6FFA493D7D83D2" blockId="5.[178,1086,1459,1611]" box="[178,528,1481,1503]" pageId="5" pageNumber="910">
<geoCoordinate id="85B9BEDCF805FFD4FF6FFA493E5F83D2" box="[178,306,1481,1503]" degrees="36" direction="north" minutes="54" orientation="latitude" pageId="5" pageNumber="910" precision="1" seconds="05.2" value="36.901447">36°54’05.2&quot;N</geoCoordinate>
102°21&quot;
<geoCoordinate id="85B9BEDCF805FFD4FE5CFA4A3ED783D2" box="[385,442,1482,1503]" degrees="07.1" direction="east" orientation="longitude" pageId="5" pageNumber="910" precision="5555" value="7.1">07.1E</geoCoordinate>
,
<quantity id="277575FEF805FFD4FE19FA493D6683D2" box="[452,523,1481,1503]" metricMagnitude="3" metricUnit="m" metricValue="2.925" pageId="5" pageNumber="910" unit="m" value="2925.0">2925 m</quantity>
.
</paragraph>
<paragraph id="E032D81BF805FFD7FF6FFA6A3EC58738" blockId="5.[178,1086,1459,1611]" lastBlockId="6.[178,1087,214,374]" lastPageId="6" lastPageNumber="911" pageId="5" pageNumber="910">
<materialsCitation id="50E5D246F805FFD4FF6FFA6A3C848022" collectingDate="2011-06-28" collectorName="Schulke" country="China" elevation="2925" latitude="36.901447" location="Park" longLatPrecision="4442" longitude="7.1" municipality="Lang Shi Dang Jing Qu" pageId="5" pageNumber="910" specimenCount="1" stateProvince="Qinghai" typeStatus="holotype">
T y p e m a t e r i a l
<typeStatus id="3F3666B9F805FFD4FE71FA693E9083F2" box="[428,509,1513,1535]" pageId="5" pageNumber="910" type="holotype">Holotype</typeStatus>
(3):
<collectingCountry id="989A988BF805FFD4FDEBFA693D1283F3" box="[566,639,1513,1534]" name="China" pageId="5" pageNumber="910">CHINA</collectingCountry>
: &quot;
<collectingCountry id="989A988BF805FFD4FD4FFA693DB083F3" box="[658,733,1513,1534]" name="China" pageId="5" pageNumber="910">CHINA</collectingCountry>
:
<collectingRegion id="224916F9F805FFD4FD35FA693C0A83F2" box="[744,871,1513,1535]" country="China" name="Qinghai" pageId="5" pageNumber="910">Qinghai Prov.</collectingRegion>
[CH 11-11a]
<collectingMunicipality id="00564261F805FFD4FC37FA6A3E0F801B" pageId="5" pageNumber="910">Lang Shi Dang Jing Qu</collectingMunicipality>
(
<location id="E5528EC0F805FFD4FEAEF9813EF2801A" LSID="urn:lsid:plazi:treatment:6824690DF805FFD7FF6FFA0D3EBD803A:E5528EC0F805FFD4FEAEF9813EF2801A" box="[371,415,1537,1559]" country="China" latitude="36.901447" longLatPrecision="4442" longitude="7.1" municipality="Lang Shi Dang Jing Qu" name="Park" pageId="5" pageNumber="910" stateProvince="Qinghai">Park</location>
),
<locationDeviation id="F180C139F805FFD4FE6FF9813D1C801A" box="[434,625,1537,1559]" location="Honggu" pageId="5" pageNumber="910">
<quantity id="277575FEF805FFD4FE6FF9813E82801A" box="[434,495,1537,1559]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="910" unit="km" value="75.0">75 km</quantity>
NW Honggu
</locationDeviation>
,
<quantity id="277575FEF805FFD4FDA3F9813DAA801A" box="[638,711,1537,1559]" metricMagnitude="3" metricUnit="m" metricValue="2.925" pageId="5" pageNumber="910" unit="m" value="2925.0">
<elevation id="6BA03F28F805FFD4FDA3F9813DAA801A" box="[638,711,1537,1559]" metricMagnitude="3" metricUnit="m" metricValue="2.925" pageId="5" pageNumber="910" unit="m" value="2925.0">2925 m</elevation>
</quantity>
,
<geoCoordinate id="85B9BEDCF805FFD4FD08F9813C22801A" box="[725,847,1537,1559]" degrees="36" direction="north" minutes="54" orientation="latitude" pageId="5" pageNumber="910" precision="1" seconds="05.2" value="36.901447">36°54’05.2&quot;N</geoCoordinate>
102° 21&quot;
<geoCoordinate id="85B9BEDCF805FFD4FC77F9813C8C801B" box="[938,993,1537,1558]" degrees="07.1" direction="east" orientation="longitude" pageId="5" pageNumber="910" precision="5555" value="7.1">07.1E</geoCoordinate>
, creek on overgrown scree, Populus and Betula for. with Rhodod., litter,
<date id="9433FEDBF805FFD4FCD9F9993C048023" box="[772,873,1561,1582]" pageId="5" pageNumber="910" value="2011-06-28">
<collectingDate id="84770733F805FFD4FCD9F9993C048023" box="[772,873,1561,1582]" pageId="5" pageNumber="910" value="2011-06-28">28.VI.2011</collectingDate>
</date>
, leg.
<collectorName id="4D78BDCDF805FFD4FC45F9993C8F8022" box="[920,994,1561,1583]" pageId="5" pageNumber="910">Schülke</collectorName>
&quot;
</materialsCitation>
.
<materialsCitation id="50E5D246F805FFD4FC2CF9993EB5804A" collectingDate="2011-06-28" collectorName="Schulke" country="China" elevation="2925" latitude="36.901447" location="Park" longLatPrecision="4442" longitude="7.1" municipality="Lang Shi Dang Jing Qu" pageId="5" pageNumber="910" specimenCount="1" specimenCount-female="1" stateProvince="Qinghai" typeStatus="allotype">
<typeStatus id="3F3666B9F805FFD4FC2CF9993B508022" box="[1009,1085,1561,1583]" pageId="5" pageNumber="910" type="allotype">Allotype</typeStatus>
(
<specimenCount id="F68B1392F805FFD4FF3BF9B63F948046" box="[230,249,1590,1611]" pageId="5" pageNumber="910" type="female">♀</specimenCount>
): samed data as holotype
</materialsCitation>
.
<typeStatus id="3F3666B9F805FFD4FE02F9B13D5D804A" box="[479,560,1585,1607]" pageId="5" pageNumber="910" type="holotype">Holotype</typeStatus>
in MSC,
<typeStatus id="3F3666B9F805FFD4FD59F9B13DA7804A" box="[644,714,1585,1607]" pageId="5" pageNumber="910" type="allotype">allotype</typeStatus>
in ASC.
<materialsCitation id="50E5D246F805FFD7FCC6F9B23EA586E0" collectingDate="2011-06-28" collectionCode="ASC, MSC" collectorName="Schulke" country="China" elevation="2925" lastPageId="6" lastPageNumber="911" latitude="36.901447" location="Park" longLatPrecision="4442" longitude="7.1" municipality="Lang Shi Dang Jing Qu" pageId="5" pageNumber="910" specimenCount="2" specimenCount-female="2" stateProvince="Qinghai" typeStatus="paratype">
<typeStatus id="3F3666B9F805FFD4FCC6F9B23C02804A" box="[795,879,1586,1607]" pageId="5" pageNumber="910" type="paratype">Paratypes</typeStatus>
: same data as holotype, 333,
<specimenCount id="F68B1392F806FFD7FEC6FF573E2686FC" box="[283,331,215,241]" pageId="6" pageNumber="911" type="female">2♀♀</specimenCount>
(
<collectionCode id="869C40DEF806FFD7FE85FF573EEB86E1" box="[344,390,215,236]" pageId="6" pageNumber="911">ASC</collectionCode>
,
<collectionCode id="869C40DEF806FFD7FE4DFF573EAF86E0" box="[400,450,215,237]" pageId="6" pageNumber="911">MSC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="50E5D246F806FFD7FE0FFF573D458708" collectingDate="2011-06-28" collectionCode="ASC, MSC" collectorName="Schulke" country="Switzerland" elevation="2896" latitude="36.902447" location="CH" longLatPrecision="1" longitude="102.35469" pageId="6" pageNumber="911" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
same, but
<quantity id="277575FEF806FFD7FDF0FF563D1E86E1" box="[557,627,214,236]" metricMagnitude="3" metricUnit="m" metricValue="2.896" pageId="6" pageNumber="911" unit="m" value="2896.0">
<elevation id="6BA03F28F806FFD7FDF0FF563D1E86E1" box="[557,627,214,236]" metricMagnitude="3" metricUnit="m" metricValue="2.896" pageId="6" pageNumber="911" unit="m" value="2896.0">2896 m</elevation>
</quantity>
,
<geoCoordinate id="85B9BEDCF806FFD7FDA0FF563D9B86E1" box="[637,758,214,236]" degrees="36" direction="north" minutes="54" orientation="latitude" pageId="6" pageNumber="911" precision="1" seconds="08.8" value="36.902447">36°54’08.8&quot;N</geoCoordinate>
<geoCoordinate id="85B9BEDCF806FFD7FD20FF563CEF86E1" box="[765,898,214,236]" degrees="102" direction="east" minutes="21" orientation="longitude" pageId="6" pageNumber="911" precision="1" seconds="16.9" value="102.35469">102°21’16.9&quot;E</geoCoordinate>
,
<date id="9433FEDBF806FFD7FC51FF573C9C86E1" box="[908,1009,215,236]" pageId="6" pageNumber="911" value="2011-06-28">
<collectingDate id="84770733F806FFD7FC51FF573C9C86E1" box="[908,1009,215,236]" pageId="6" pageNumber="911" value="2011-06-28">28.VI.2011</collectingDate>
</date>
, [
<collectingCountry id="989A988BF806FFD7FBDDFF573B4E86E1" box="[1024,1059,215,236]" name="Switzerland" pageId="6" pageNumber="911">CH</collectingCountry>
11- 11], leg.
<collectorName id="4D78BDCDF806FFD7FEF1FF6E3E198709" box="[300,372,238,260]" pageId="6" pageNumber="911">Schülke</collectorName>
,
<specimenCount id="F68B1392F806FFD7FEA0FF6F3EC08704" box="[381,429,239,265]" pageId="6" pageNumber="911" type="female">4♀♀</specimenCount>
(
<collectionCode id="869C40DEF806FFD7FE64FF6F3E8A8709" box="[441,487,239,260]" pageId="6" pageNumber="911">ASC</collectionCode>
,
<collectionCode id="869C40DEF806FFD7FE32FF6F3D4C8708" box="[495,545,239,261]" pageId="6" pageNumber="911">MSC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="50E5D246F806FFD7FDECFF6F3C188708" box="[561,885,238,265]" collectionCode="MSC" country="Switzerland" location="Wrase" pageId="6" pageNumber="911" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
same, but D.W.
<location id="E5528EC0F806FFD7FD63FF6F3D958709" LSID="urn:lsid:plazi:treatment:6824690DF805FFD7FF6FFA0D3EBD803A:E5528EC0F806FFD7FD63FF6F3D958709" box="[702,760,239,260]" country="Switzerland" name="Wrase" pageId="6" pageNumber="911">Wrase</location>
,
<specimenCount id="F68B1392F806FFD7FCDCFF6F3C5C8704" box="[769,817,239,265]" pageId="6" pageNumber="911" type="female">2♀♀</specimenCount>
(
<collectionCode id="869C40DEF806FFD7FCE1FF6F3C028708" box="[828,879,239,261]" pageId="6" pageNumber="911">MSC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="50E5D246F806FFD7FCA3FF6E3EC98738" collectingDate="2011-07-05" collectionCode="MSC" collectorName="M. Schulke" country="Switzerland" elevation="2704" location="Men Yuan" longitude="102.034996" municipality="Qinghai" pageId="6" pageNumber="911" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingMunicipality id="00564261F806FFD7FCA3FF6E3CAB8709" box="[894,966,238,260]" pageId="6" pageNumber="911">Qinghai</collectingMunicipality>
, road
<quantity id="277575FEF806FFD7FC26FF6F3B538709" box="[1019,1086,238,260]" metricMagnitude="5" metricUnit="m" metricValue="3.01" pageId="6" pageNumber="911" unit="km" value="301.0">301 km</quantity>
180, 43 km ESE
<location id="E5528EC0F806FFD7FEA3FE873E8D8711" LSID="urn:lsid:plazi:treatment:6824690DF805FFD7FF6FFA0D3EBD803A:E5528EC0F806FFD7FEA3FE873E8D8711" box="[382,480,263,284]" country="Switzerland" longitude="102.034996" municipality="Qinghai" name="Men Yuan" pageId="6" pageNumber="911">Men Yuan</location>
, 37°09.32.6N
<geoCoordinate id="85B9BEDCF806FFD7FDBFFE873D8A8711" box="[610,743,262,284]" degrees="102" direction="east" minutes="02" orientation="longitude" pageId="6" pageNumber="911" precision="1" seconds="06.0" value="102.034996">102°02’06.0&quot;E</geoCoordinate>
,
<quantity id="277575FEF806FFD7FD2FFE873C558711" box="[754,824,263,284]" metricMagnitude="3" metricUnit="m" metricValue="2.7039999999999997" pageId="6" pageNumber="911" unit="m" value="2704.0">
<elevation id="6BA03F28F806FFD7FD2FFE873C558711" box="[754,824,263,284]" metricMagnitude="3" metricUnit="m" metricValue="2.7039999999999997" pageId="6" pageNumber="911" unit="m" value="2704.0">2704 m</elevation>
</quantity>
,
<date id="9433FEDBF806FFD7FC99FE863CC88711" box="[836,933,262,284]" pageId="6" pageNumber="911" value="2011-07-05">
<collectingDate id="84770733F806FFD7FC99FE863CC88711" box="[836,933,262,284]" pageId="6" pageNumber="911" value="2011-07-05">5.VII.2011</collectingDate>
</date>
, [
<collectingCountry id="989A988BF806FFD7FC68FE873CBB8711" box="[949,982,263,284]" name="Switzerland" pageId="6" pageNumber="911">CH</collectingCountry>
11-19],
<collectorName id="4D78BDCDF806FFD7FBF9FE873E4A8739" pageId="6" pageNumber="911">M. Schülke</collectorName>
,
<specimenCount id="F68B1392F806FFD7FEEDFE9F3E0D8734" box="[304,352,287,313]" pageId="6" pageNumber="911" type="female">2♀♀</specimenCount>
(
<collectionCode id="869C40DEF806FFD7FEB6FE9F3EF38738" box="[363,414,287,309]" pageId="6" pageNumber="911">MSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFEC33CA98778" blockId="6.[178,1087,214,374]" pageId="6" pageNumber="911">
D e s c r i p t i o n: In all characters similar to
<taxonomicName id="278DA398F806FFD7FD78FEC33CBC8755" authority="SMETANA, 2008" authorityName="SMETANA" authorityYear="2008" box="[677,977,323,344]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="911" phylum="Arthropoda" rank="species" species="menippus">
<emphasis id="D2F90409F806FFD7FD78FEC33C4B8755" box="[677,806,323,344]" italics="true" pageId="6" pageNumber="911">Q. menippus</emphasis>
SMETANA, 2008
</taxonomicName>
and different only by the different aedoeagus and tergite 10 of female genital segment.
</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFE053EC8856E" blockId="6.[178,1086,389,867]" pageId="6" pageNumber="911">
Male. First four segments of front tarsus markedly dilated, almost patellate, each densely covered with tenent setae ventrally; segment 2 markedly wider than apex of tibia (ratio 1.40), segment 4 markedly narrower than preceding segments. Sternite 8 rather densely setose, with two long setae on each side; with wide and deep medioapical emargination, small triangular area before emargination flattened and smooth (
<figureCitation id="78B6C49EF806FFD7FC87FE773CC58400" box="[858,936,503,525]" captionStart="Figs 11-19" captionStartId="17.[178,218,1473,1495]" captionTargetBox="[179,1082,213,1461]" captionTargetId="figure-1@17.[178,1084,212,1464]" captionTargetPageId="17" captionText="Figs 11-19: 11-14: Quedius bipictus sp. nov.: (11) aedoeagus, ventral view; (12) apical portion of aedoeagus, lateral view; (13) apical portion of underside of paramere with sensory peg setae; (14) tergite 10 of female genital segment. Quedius gynaikos sp. nov.: (15) tergite 10 of female genital segment. 16-19: Quedius causarius sp. nov.: (16) apical portion of male sternite 8; (17) tergite 10 of male genital segment; (18) sternite 9 of male genital segment; (19) aedoeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5415644" httpUri="https://zenodo.org/record/5415644/files/figure.png" pageId="6" pageNumber="911">Fig. 16</figureCitation>
). Genital segment with tergite 10 rather narrow, narrowed toward arcuate apex, with three or four long setae at apex, otherwise only sparingly setose (
<figureCitation id="78B6C49EF806FFD7FD04FDB03C4A844B" box="[729,807,560,582]" captionStart="Figs 11-19" captionStartId="17.[178,218,1473,1495]" captionTargetBox="[179,1082,213,1461]" captionTargetId="figure-1@17.[178,1084,212,1464]" captionTargetPageId="17" captionText="Figs 11-19: 11-14: Quedius bipictus sp. nov.: (11) aedoeagus, ventral view; (12) apical portion of aedoeagus, lateral view; (13) apical portion of underside of paramere with sensory peg setae; (14) tergite 10 of female genital segment. Quedius gynaikos sp. nov.: (15) tergite 10 of female genital segment. 16-19: Quedius causarius sp. nov.: (16) apical portion of male sternite 8; (17) tergite 10 of male genital segment; (18) sternite 9 of male genital segment; (19) aedoeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5415644" httpUri="https://zenodo.org/record/5415644/files/figure.png" pageId="6" pageNumber="911">Fig. 17</figureCitation>
); sternite 9 with wide and short basal portion, apical portion widely arcuate apically, with two slightly differentiated apical setae, otherwise finely setose (
<figureCitation id="78B6C49EF806FFD7FDBDFDE93DC38472" box="[608,686,617,639]" captionStart="Figs 11-19" captionStartId="17.[178,218,1473,1495]" captionTargetBox="[179,1082,213,1461]" captionTargetId="figure-1@17.[178,1084,212,1464]" captionTargetPageId="17" captionText="Figs 11-19: 11-14: Quedius bipictus sp. nov.: (11) aedoeagus, ventral view; (12) apical portion of aedoeagus, lateral view; (13) apical portion of underside of paramere with sensory peg setae; (14) tergite 10 of female genital segment. Quedius gynaikos sp. nov.: (15) tergite 10 of female genital segment. 16-19: Quedius causarius sp. nov.: (16) apical portion of male sternite 8; (17) tergite 10 of male genital segment; (18) sternite 9 of male genital segment; (19) aedoeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5415644" httpUri="https://zenodo.org/record/5415644/files/figure.png" pageId="6" pageNumber="911">Fig. 18</figureCitation>
). Aedoeagus (
<figureCitation id="78B6C49EF806FFD7FC99FDE93CDB8472" box="[836,950,617,639]" captionStart-0="Figs 11-19" captionStart-1="Figs 20-30" captionStartId-0="17.[178,218,1473,1495]" captionStartId-1="18.[178,218,1446,1468]" captionTargetBox-0="[179,1082,213,1461]" captionTargetBox-1="[197,1066,218,1437]" captionTargetId-0="figure-1@17.[178,1084,212,1464]" captionTargetId-1="figure-1@18.[197,1067,212,1437]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figs 11-19: 11-14: Quedius bipictus sp. nov.: (11) aedoeagus, ventral view; (12) apical portion of aedoeagus, lateral view; (13) apical portion of underside of paramere with sensory peg setae; (14) tergite 10 of female genital segment. Quedius gynaikos sp. nov.: (15) tergite 10 of female genital segment. 16-19: Quedius causarius sp. nov.: (16) apical portion of male sternite 8; (17) tergite 10 of male genital segment; (18) sternite 9 of male genital segment; (19) aedoeagus, ventral view." captionText-1="Figs 20-30: 20-23: Quedius causarius sp. nov.: (20) apical portion of aedoeagus, lateral view; (21) apical portion of median lobe, paramere removed; (22) apical portion of underside of paramere with sensory peg setae; (23) tergite 10 of female genital segment. 24-29: Quedius laetepictus sp. nov.: (24) apical portion of male sternite 8; (25) tergite 10 of male genital segment; (26) sternite 9 of male genital segment; (27) aedoeagus, ventral view; (28) apical portion of median lobe, paramere removed; (29) apical portion of underside of paramere with sensory peg setae. Quedius iaculifer sp. nov.: (30) apical portion of male sternite 8." figureDoi-0="http://doi.org/10.5281/zenodo.5415644" figureDoi-1="http://doi.org/10.5281/zenodo.5415648" httpUri-0="https://zenodo.org/record/5415644/files/figure.png" httpUri-1="https://zenodo.org/record/5415648/files/figure.png" pageId="6" pageNumber="911">Figs 19-22</figureCitation>
) rather large and robust; median lobe markedly, almost conically narrowed into narrowly arcuate apex, on face adjacent to paramere with short complex carina below apex; paramere large, shaped as in
<figureCitation id="78B6C49EF806FFD7FEA2FD3E3EA384DE" box="[383,462,702,724]" captionStart="Figs 11-19" captionStartId="17.[178,218,1473,1495]" captionTargetBox="[179,1082,213,1461]" captionTargetId="figure-1@17.[178,1084,212,1464]" captionTargetPageId="17" captionText="Figs 11-19: 11-14: Quedius bipictus sp. nov.: (11) aedoeagus, ventral view; (12) apical portion of aedoeagus, lateral view; (13) apical portion of underside of paramere with sensory peg setae; (14) tergite 10 of female genital segment. Quedius gynaikos sp. nov.: (15) tergite 10 of female genital segment. 16-19: Quedius causarius sp. nov.: (16) apical portion of male sternite 8; (17) tergite 10 of male genital segment; (18) sternite 9 of male genital segment; (19) aedoeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5415644" httpUri="https://zenodo.org/record/5415644/files/figure.png" pageId="6" pageNumber="911">Figs 19</figureCitation>
,
<figureCitation id="78B6C49EF806FFD7FE01FD3E3E9584DE" box="[476,504,702,723]" captionStart="Figs 20-30" captionStartId="18.[178,218,1446,1468]" captionTargetBox="[197,1066,218,1437]" captionTargetId="figure-1@18.[197,1067,212,1437]" captionTargetPageId="18" captionText="Figs 20-30: 20-23: Quedius causarius sp. nov.: (20) apical portion of aedoeagus, lateral view; (21) apical portion of median lobe, paramere removed; (22) apical portion of underside of paramere with sensory peg setae; (23) tergite 10 of female genital segment. 24-29: Quedius laetepictus sp. nov.: (24) apical portion of male sternite 8; (25) tergite 10 of male genital segment; (26) sternite 9 of male genital segment; (27) aedoeagus, ventral view; (28) apical portion of median lobe, paramere removed; (29) apical portion of underside of paramere with sensory peg setae. Quedius iaculifer sp. nov.: (30) apical portion of male sternite 8." figureDoi="http://doi.org/10.5281/zenodo.5415648" httpUri="https://zenodo.org/record/5415648/files/figure.png" pageId="6" pageNumber="911">22</figureCitation>
, with narrowly arcuate apex slightly exceeding apex of median lobe, with face adjacent to median lobe markedly excavate; two apical setae, one minute seta at each lateral margin below apex and two similar setae at each lateral margin still farther posteriad; sensory peg setae on underside quite small and few, usually three setae below apex and two or one on each side below apex, shifted quite close to lateral margin (
<figureCitation id="78B6C49EF806FFD7FE90FCCC3EF7856E" box="[333,410,844,867]" captionStart="Figs 20-30" captionStartId="18.[178,218,1446,1468]" captionTargetBox="[197,1066,218,1437]" captionTargetId="figure-1@18.[197,1067,212,1437]" captionTargetPageId="18" captionText="Figs 20-30: 20-23: Quedius causarius sp. nov.: (20) apical portion of aedoeagus, lateral view; (21) apical portion of median lobe, paramere removed; (22) apical portion of underside of paramere with sensory peg setae; (23) tergite 10 of female genital segment. 24-29: Quedius laetepictus sp. nov.: (24) apical portion of male sternite 8; (25) tergite 10 of male genital segment; (26) sternite 9 of male genital segment; (27) aedoeagus, ventral view; (28) apical portion of median lobe, paramere removed; (29) apical portion of underside of paramere with sensory peg setae. Quedius iaculifer sp. nov.: (30) apical portion of male sternite 8." figureDoi="http://doi.org/10.5281/zenodo.5415648" httpUri="https://zenodo.org/record/5415648/files/figure.png" pageId="6" pageNumber="911">Fig. 22</figureCitation>
).
</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFCF13E7F85F5" blockId="6.[178,1086,881,1016]" pageId="6" pageNumber="911">
Female. First three segments of front tarsus similar to those of male, but considerably less dilated; segment 2 about as wide as apex of tibia. Tergite 10 of genital segment quite characteristic, rather narrow with large pigmented middle portion and with distinct, setose medioapical carina and with numerous setae on apical portion, otherwise asetose (
<figureCitation id="78B6C49EF806FFD7FF67FC623E6B85F5" box="[186,262,994,1016]" captionStart="Figs 20-30" captionStartId="18.[178,218,1446,1468]" captionTargetBox="[197,1066,218,1437]" captionTargetId="figure-1@18.[197,1067,212,1437]" captionTargetPageId="18" captionText="Figs 20-30: 20-23: Quedius causarius sp. nov.: (20) apical portion of aedoeagus, lateral view; (21) apical portion of median lobe, paramere removed; (22) apical portion of underside of paramere with sensory peg setae; (23) tergite 10 of female genital segment. 24-29: Quedius laetepictus sp. nov.: (24) apical portion of male sternite 8; (25) tergite 10 of male genital segment; (26) sternite 9 of male genital segment; (27) aedoeagus, ventral view; (28) apical portion of median lobe, paramere removed; (29) apical portion of underside of paramere with sensory peg setae. Quedius iaculifer sp. nov.: (30) apical portion of male sternite 8." figureDoi="http://doi.org/10.5281/zenodo.5415648" httpUri="https://zenodo.org/record/5415648/files/figure.png" pageId="6" pageNumber="911">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFB873E118210" blockId="6.[178,380,1031,1053]" box="[178,380,1031,1053]" pageId="6" pageNumber="911">Length 6,0-7.0 mm.</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFBAD3E508271" blockId="6.[178,1085,1068,1149]" pageId="6" pageNumber="911">
E t y m o l o g y: The specific epithet is the Latin adjective
<emphasis id="D2F90409F806FFD7FCF3FBAD3CFD824F" box="[814,912,1069,1090]" italics="true" pageId="6" pageNumber="911">causarius</emphasis>
, -
<emphasis id="D2F90409F806FFD7FC7AFBAD3C87824F" box="[935,1002,1069,1090]" italics="true" pageId="6" pageNumber="911">a, -um</emphasis>
(sickly, invalided). It refers to the drastic reduction of the sensory peg setae on the underside of the paramere.
</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFB0C3EB882CD" blockId="6.[178,1085,1164,1216]" pageId="6" pageNumber="911">
G e o g r a p h i c a l d i s t r i b u t i o n: The species is at present known from two localities in eastern
<collectingRegion id="224916F9F806FFD7FEA1FB2A3EBC82CD" box="[380,465,1194,1216]" country="China" name="Qinghai" pageId="6" pageNumber="911">Qinghai</collectingRegion>
.
</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFB4F3C2A8330" blockId="6.[178,1091,1231,1341]" pageId="6" pageNumber="911">
B i o n o m i c s:Specimensoftheoriginalseriesweretakenonovergrownscreewith
<emphasis id="D2F90409F806FFD7FF6FFB6C3E68830C" box="[178,261,1260,1281]" italics="true" pageId="6" pageNumber="911">Populus</emphasis>
and
<emphasis id="D2F90409F806FFD7FE9EFB6C3EE8830C" box="[323,389,1260,1281]" italics="true" pageId="6" pageNumber="911">Betula</emphasis>
forest with rhododendrons by sifting forest floor litter, and on another overgrown scree with
<emphasis id="D2F90409F806FFD7FE30FA893D2D8313" box="[493,576,1289,1310]" italics="true" pageId="6" pageNumber="911">Populus</emphasis>
and
<emphasis id="D2F90409F806FFD7FDA8FA893DDA8313" box="[629,695,1289,1310]" italics="true" pageId="6" pageNumber="911">Betula</emphasis>
forest by sifting litter between rocks. Overgrown screes seem to be the preferred habitat of the species.
</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFACC3B4783DA" blockId="6.[178,1085,1356,1495]" pageId="6" pageNumber="911">
R e c o g n i t i o n a n d c o m m e n t s
<taxonomicName id="278DA398F806FFD7FD78FACC3C07836F" authorityName="Smetana" authorityYear="2015" box="[677,874,1356,1378]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="911" phylum="Arthropoda" rank="species" species="causarius">
<emphasis id="D2F90409F806FFD7FD78FACC3C07836F" box="[677,874,1356,1378]" italics="true" pageId="6" pageNumber="911">Quedius causarius</emphasis>
</taxonomicName>
may be positively recognized only by the characteristic shape of the aedoeagus, including the location and the drastic reduction of both the number and size of the sensory peg setae on the underside of the paramere (
<figureCitation id="78B6C49EF806FFD7FE4AFA243E8983B7" box="[407,484,1444,1466]" captionStart="Figs 20-30" captionStartId="18.[178,218,1446,1468]" captionTargetBox="[197,1066,218,1437]" captionTargetId="figure-1@18.[197,1067,212,1437]" captionTargetPageId="18" captionText="Figs 20-30: 20-23: Quedius causarius sp. nov.: (20) apical portion of aedoeagus, lateral view; (21) apical portion of median lobe, paramere removed; (22) apical portion of underside of paramere with sensory peg setae; (23) tergite 10 of female genital segment. 24-29: Quedius laetepictus sp. nov.: (24) apical portion of male sternite 8; (25) tergite 10 of male genital segment; (26) sternite 9 of male genital segment; (27) aedoeagus, ventral view; (28) apical portion of median lobe, paramere removed; (29) apical portion of underside of paramere with sensory peg setae. Quedius iaculifer sp. nov.: (30) apical portion of male sternite 8." figureDoi="http://doi.org/10.5281/zenodo.5415648" httpUri="https://zenodo.org/record/5415648/files/figure.png" pageId="6" pageNumber="911">Fig. 22</figureCitation>
), a character state unique among the
<taxonomicName id="278DA398F806FFD7FCB6FA243C8883B4" authorityName="CASEY" authorityYear="1915" box="[875,997,1444,1465]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="920" phylum="Arthropoda" rank="subGenus" subGenus="Distichalius">
<emphasis id="D2F90409F806FFD7FCB6FA243C8883B4" box="[875,997,1444,1465]" italics="true" pageId="6" pageNumber="911">Distichalius</emphasis>
</taxonomicName>
-species, as well as by the characteristic development of tergite 10 of the female genital segment.
</paragraph>
<paragraph id="E032D81BF806FFD7FF6FFA663EBD803A" blockId="6.[178,1085,1510,1591]" pageId="6" pageNumber="911">The location and reduction of the number and size of the sensory peg setae on the paramere is apparently caused by the pronounced excavation of the face of the paramere adjacent to the median lobe.</paragraph>
</subSubSection>
</treatment>
</document>
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