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<mods:title id="3EDAE3396C25F6B2792225C12563B4D7">First Cretaceous teleostean otolith assemblage (Arkadelphia Formation, upper Maastrichtian) from Arkansas, USA, early Gadiformes, and the Western Interior Seaway</mods:title>
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<treatment id="2B4F878B2571FF8BFFB9FA71FBE1FDE4" ID-DOI="http://doi.org/10.5281/zenodo.11474219" ID-GBIF-Taxon="232134926" ID-Zenodo-Dep="11474219" LSID="urn:lsid:plazi:treatment:2B4F878B2571FF8BFFB9FA71FBE1FDE4" httpUri="http://treatment.plazi.org/id/2B4F878B2571FF8BFFB9FA71FBE1FDE4" lastPageNumber="16" pageId="17" pageNumber="16">
<subSubSection id="EBFC65162571FF8BFFB9FA71FE18FA65" pageId="17" pageNumber="16" type="nomenclature">
<paragraph id="A359369D2571FF8BFFB9FA71FCEBFA44" blockId="17.[114,799,1380,1506]" box="[114,799,1411,1438]" pageId="17" pageNumber="16">
<heading id="F81181F12571FF8BFFB9FA71FCEBFA44" allCaps="true" box="[114,799,1411,1438]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="16" reason="2">
<taxonomicName id="64E64D1E2571FF8BFFB9FA71FEEEFA47" authorityName="RATH" authorityYear="1859" box="[114,282,1413,1438]" family="Merlucciidae" genus="Palaeogadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="17" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="9192EA8F2571FF8BFFB9FA71FEEEFA47" box="[114,282,1413,1438]" italics="true" pageId="17" pageNumber="16">PALAEOGADUS</emphasis>
</taxonomicName>
CF.
<taxonomicName id="64E64D1E2571FF8BFE8DFA77FCEEFA44" authority="SCHWARZHANS AND STRINGER" authorityName="Schwarzhans and Stringer" authorityYear="2020" box="[326,794,1411,1438]" family="Merlucciidae" genus="Palaeogadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="17" pageNumber="17" phylum="Chordata" rank="species" species="weltoni">
<emphasis id="9192EA8F2571FF8BFE8DFA77FE3DFA47" box="[326,457,1411,1438]" italics="true" pageId="17" pageNumber="16">P. WELTONI</emphasis>
SCHWARZHANS AND STRINGER
</taxonomicName>
,
</heading>
</paragraph>
<paragraph id="A359369D2571FF8BFE6FFA51FE18FA65" blockId="17.[114,799,1380,1506]" box="[420,492,1445,1469]" pageId="17" pageNumber="16">2020a</paragraph>
</subSubSection>
<subSubSection id="EBFC65162571FF8BFE6AFA3EFE1BFA3B" box="[417,495,1482,1506]" pageId="17" pageNumber="16" type="description">
<paragraph id="A359369D2571FF8BFE6AFA3EFE1BFA3B" blockId="17.[114,799,1380,1506]" box="[417,495,1482,1506]" pageId="17" pageNumber="16">
<figureCitation id="3BDD2A182571FF8BFE6AFA3EFE1BFA3B" box="[417,495,1482,1506]" captionStart="Figure 8" captionStartId="18.[115,190,1732,1756]" captionTargetBox="[243,1396,192,1709]" captionTargetId="figure-15@18.[243,1396,192,1709]" captionTargetPageId="18" captionText="Figure 8. Otoliths from the Cretaceous Arkadelphia Formation. All specimens unless otherwise noted are inner views of right sagittae. Lapilli are macular views. Length in mm. A. Palaeogadus cf. P. weltoni Schwarzhans and Stringer (2020a), DMNH 2021-09- 23, 1.46 mm. B. Gadiformes indeterminate, DMNH 2021-09-24, 1.56 mm. C. Tippaha mythica Schwarzhans and Stringer (2020a), DMNH 2021-09-25, 3.85 mm. D. Eutawichthys maastrichtiensis Nolf and Stringer (1996), DMNH 2021-09-26, 3.93 mm. E. Eutawichthys zideki Nolf and Stringer (1996), DMNH 2021-09-27, 1.42 mm. F.Eutawichthys cf. E. stringeri Schwarzhans, Huddleston, and Takeuchi (2018b), DMNH 2021-09-28, 1.85 mm. G. Ampheristus cf. A. americanus Schwarzhans and Stringer (2020a), DMNH 2021-09-29, 1.58 mm. H. Protobythities brzobohatyi Schwarzhans (2010), DMNH 2021-09-30, 1.68 mm. I. Lapillus type 1, DMNH 2021-09-31, 2.98 mm." figureDoi="http://doi.org/10.5281/zenodo.10913436" httpUri="https://zenodo.org/record/10913436/files/figure.png" pageId="17" pageNumber="16">FIG. 8A</figureCitation>
</paragraph>
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<paragraph id="A359369D2571FF8BFF5BFA0FFEFFF9E0" blockId="17.[112,801,1531,1945]" pageId="17" pageNumber="16">
<emphasis id="9192EA8F2571FF8BFF5BFA0FFEE8F9CC" bold="true" box="[144,284,1531,1557]" pageId="17" pageNumber="16">Material—</emphasis>
one small,slightly eroded specimen,
<collectionCode id="C5F7AE582571FF8BFD05FA08FCD5F9CF" box="[718,801,1532,1558]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34940" lsid="urn:lsid:biocol.org:col:34940" name="Delaware Museum of Natural History" pageId="17" pageNumber="16" type="Museum">DMNH</collectionCode>
<date id="D758105D2571FF8BFFBBF9EBFEFFF9E0" box="[112,267,1567,1593]" pageId="17" pageNumber="16" value="2021-09-23">2021-09-23.</date>
</paragraph>
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<subSubSection id="EBFC65162571FF8BFF5BF9B5FBE1FDE4" pageId="17" pageNumber="16" type="description">
<paragraph id="A359369D2571FF8BFF5BF9B5FBE1FDE4" blockId="17.[112,801,1531,1945]" lastBlockId="17.[832,1520,195,573]" pageId="17" pageNumber="16">
<emphasis id="9192EA8F2571FF8BFF5BF9B5FDE4F985" bold="true" box="[144,528,1601,1628]" pageId="17" pageNumber="16">Description and Remarks—</emphasis>
The one Arkadelphia Formation specimen assigned to
<taxonomicName id="64E64D1E2571FF8BFDC2F991FCF2F959" authorityName="Schwarzhans and Stringer" authorityYear="2020" box="[521,774,1636,1665]" family="Merlucciidae" genus="Palaeogadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="17" pageNumber="16" phylum="Chordata" rank="species" species="weltoni">
<emphasis id="9192EA8F2571FF8BFDC2F991FCF2F959" box="[521,774,1636,1665]" italics="true" pageId="17" pageNumber="16">Palaeogadus weltoni</emphasis>
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is very small and eroded. However, it was felt that it possessed enough gadid features to compare it to
<taxonomicName id="64E64D1E2571FF8BFD6EF958FF6EF930" authorityName="RATH" authorityYear="1859" family="Merlucciidae" genus="Palaeogadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="17" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="9192EA8F2571FF8BFD6EF958FF6EF930" italics="true" pageId="17" pageNumber="16">Palaeogadus</emphasis>
</taxonomicName>
cf.
<taxonomicName id="64E64D1E2571FF8BFF0FF939FECCF930" authorityName="Schwarzhans and Stringer" authorityYear="2020" box="[196,312,1741,1769]" family="Merlucciidae" genus="Palaeogadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="17" pageNumber="16" phylum="Chordata" rank="species" species="weltoni">
<emphasis id="9192EA8F2571FF8BFF0FF939FECCF930" box="[196,312,1741,1769]" italics="true" pageId="17" pageNumber="16">P. weltoni</emphasis>
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. Similarities include the overall oblong shape (
<emphasis id="9192EA8F2571FF8BFF02F904FEF8F8D5" box="[201,268,1776,1804]" italics="true" pageId="17" pageNumber="16">sensu</emphasis>
<bibRefCitation id="C7774B6C2571FF8BFEDAF906FE14F8D4" author="Smale, M. &amp; G. Watson &amp; T. Hecht" box="[273,480,1778,1805]" pageId="17" pageNumber="16" pagination="1 - 253" refId="ref30467" refString="Smale, M., G. Watson, and T. Hecht. 1995. Otolith atlas of southern African marine fishes. Ichthyological Monographs of the J. L. B. Smith Institute of Ichthyology 1: 1 - 253." type="journal article" year="1995">Smale et al. 1995</bibRefCitation>
), the slightly convex inner face; the tapered and rounded anterior and posterior margins that are almost alike; the homosulcoid-type sulcus; a broadly, gently arched dorsal margin; a very shallow ventral margin that approaches horizontal; and a prominent ventral furrow extending from under the anterior of the ostium to near the posterior of the cauda; the anterior and posterior ends of the ventral furrow appear to turn upwards. It compares very well to the
<specimenCount id="B5E0FD142571FF8BFA09FED9FC43FEB3" count="1" pageId="17" pageNumber="16" type="generic">one specimen</specimenCount>
of
<taxonomicName id="64E64D1E2571FF8BFC29FEBAFBADFEB3" authorityName="Schwarzhans and Stringer" authorityYear="2020" box="[994,1113,334,362]" family="Merlucciidae" genus="Palaeogadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="17" pageNumber="16" phylum="Chordata" rank="species" species="weltoni">
<emphasis id="9192EA8F2571FF8BFC29FEBAFBADFEB3" box="[994,1113,334,362]" italics="true" pageId="17" pageNumber="16">P. weltoni</emphasis>
</taxonomicName>
illustrated by
<bibRefCitation id="C7774B6C2571FF8BFAD8FEA4FBE5FE54" author="Schwarzhans, W. &amp; G. Stringer" pageId="17" pageNumber="16" pagination="395 - 446" refId="ref30045" refString="Schwarzhans, W., and G. Stringer. 2020 a. Fish otoliths from the late Maastrichtian Kemp Clay (Texas, USA) and the early Danian Clayton Formation (Arkansas, USA) and an assessment of extinction and survival of teleost lineages across the K-Pg boundary based on otoliths. Rivista Italiana di Paleontologia e Stratigrafia 126 (2): 395 - 446." type="journal article" year="2020">Schwarzhans and Stringer (2020a)</bibRefCitation>
from the Kemp Clay in
<collectingRegion id="6122F87F2571FF8BFAE1FE87FA84FE54" box="[1322,1392,371,397]" country="United States of America" name="Texas" pageId="17" pageNumber="16">Texas</collectingRegion>
and to the specimens shown by
<bibRefCitation id="C7774B6C2571FF8BFB81FE62FA04FE68" author="Stringer, G. &amp; W. Schwarzhans" box="[1098,1520,406,433]" pageId="17" pageNumber="16" pagination="104867" refId="ref31141" refString="Stringer, G., and W. Schwarzhans. 2021. Upper Cretaceous teleosteans otoliths from the Severn Formation (Maastrichtian) of Maryland, USA, with an unusual occurrence of Silurifomes and Beryciformes and the oldest Atlantic coast Gadiformes. Cretaceous Research 125 (2021) 104867." type="journal article" year="2021">Stringer and Schwarzhans (2021)</bibRefCitation>
from the Severn Formation in
<collectingRegion id="6122F87F2571FF8BFB7EFE4DFADBFE0A" box="[1205,1327,441,467]" country="United States of America" name="Maryland" pageId="17" pageNumber="16">Maryland</collectingRegion>
.
<taxonomicName id="64E64D1E2571FF8BFAF2FE4EFA21FE0F" authorityName="RATH" authorityYear="1859" box="[1337,1493,442,470]" family="Merlucciidae" genus="Palaeogadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="17" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="9192EA8F2571FF8BFAF2FE4EFA21FE0F" box="[1337,1493,442,470]" italics="true" pageId="17" pageNumber="16">Palaeogadus</emphasis>
</taxonomicName>
is an extinct genus that is known from otoliths and skeletons from the early Paleogene of Europe (
<bibRefCitation id="C7774B6C2571FF8BFA80FDF4FC75FDE4" author="Schwarzhans, W." pageId="17" pageNumber="16" pagination="1 - 94" refId="ref29703" refString="Schwarzhans, W. 2003. Fish otoliths from the Paleocene of Denmark. Geological Survey of Denmark and Greenland Bulletin 2: 1 - 94." type="journal article" year="2003">Schwarzhans 2003</bibRefCitation>
, fig. 20AI).
</paragraph>
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