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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1088.72910" ID-GBIF-Dataset="06fde24f-d1bf-4b09-a390-e01ee47e2ec3" ID-PMC="PMC8930931" ID-Pensoft-Pub="1313-2970-1088-129" ID-Pensoft-UUID="A9678B3356615CE9BE576ADD2001E4A0" ID-PubMed="35437367" ID-ZooBank="015929874A644BA89D830B4A2F162E27" ModsDocID="1313-2970-1088-129" checkinTime="1647039168473" checkinUser="pensoft" docAuthor="Szumik, Claudia, Pereyra, Veronica, Szumik, Victoria E. Goloboff, Costa-Pinto, Paula Jessica &amp; Juarez, Maria Laura" docDate="2022" docId="203627CC7A7C5EEB8B3867B07761B320" docLanguage="en" docName="ZooKeys 1088: 129-171" docOrigin="ZooKeys 1088" docPubDate="2022-03-10" docSource="http://dx.doi.org/10.3897/zookeys.1088.72910" docTitle="Archembia Ross 1971" docType="treatment" docVersion="4" id="A9678B3356615CE9BE576ADD2001E4A0" lastPageNumber="129" masterDocId="A9678B3356615CE9BE576ADD2001E4A0" masterDocTitle="Embioptera (Insecta) from Brazil: New species and a taxonomic update" masterLastPageNumber="171" masterPageNumber="129" pageNumber="129" updateTime="1668151519134" updateUser="ExternalLinkService">
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<mods:title>Embioptera (Insecta) from Brazil: New species and a taxonomic update</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Szumik, Claudia</mods:namePart>
<mods:affiliation>Unidad Ejecutora Lillo, Consejo Nacional de Investigaciones Cientificas y Tecnicas, Fundacion Miguel Lillo, Miguel Lillo 251, 4000, S. M. de Tucuman, Argentina</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pereyra, Veronica</mods:namePart>
<mods:affiliation>Unidad Ejecutora Lillo, Consejo Nacional de Investigaciones Cientificas y Tecnicas, Fundacion Miguel Lillo, Miguel Lillo 251, 4000, S. M. de Tucuman, Argentina</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Szumik, Victoria E. Goloboff</mods:namePart>
<mods:affiliation>Unidad Ejecutora Lillo, Consejo Nacional de Investigaciones Cientificas y Tecnicas, Fundacion Miguel Lillo, Miguel Lillo 251, 4000, S. M. de Tucuman, Argentina &amp; Facultad de Ciencias Naturales e Instituto Miguel Lillo, Universidad Nacional de Tucuman, Argentina</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Costa-Pinto, Paula Jessica</mods:namePart>
<mods:affiliation>Instituto Nacional de Pesquisas da Amazonia, INPA, Manaus, Amazonas, Brazil &amp; Programa de Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Juarez, Maria Laura</mods:namePart>
<mods:affiliation>Unidad Ejecutora Lillo, Consejo Nacional de Investigaciones Cientificas y Tecnicas, Fundacion Miguel Lillo, Miguel Lillo 251, 4000, S. M. de Tucuman, Argentina</mods:affiliation>
<mods:nameIdentifier type="email">lau_zoo@yahoo.com.ar</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-10</mods:number>
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<mods:number>1088</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1088.72910</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1088.72910</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1088-129</mods:identifier>
<mods:identifier type="ZooBank">015929874A644BA89D830B4A2F162E27</mods:identifier>
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<treatment ID-GBIF-Taxon="193943946" LSID="urn:lsid:plazi:treatment:203627CC7A7C5EEB8B3867B07761B320" httpUri="http://treatment.plazi.org/id/203627CC7A7C5EEB8B3867B07761B320" lastPageNumber="129" pageId="0" pageNumber="129">
<subSubSection pageId="0" pageNumber="129" type="nomenclature">
<paragraph pageId="0" pageNumber="129">
Genus
<taxonomicName LSID="203627CC-7A7C-5EEB-8B38-67B07761B320" authority="Ross, 1971" authorityName="Ross" authorityYear="1971" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Archembia Ross, 1971</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="reference_group">
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Ross" authorityYear="1971" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">Archembia</taxonomicName>
Ross, 1971: 30, type species
<taxonomicName authorityName="Ross" authorityYear="1971" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia lacombea" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="lacombea">Archembia lacombea</taxonomicName>
Ross by original designation;
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-0031.1996.tb00192.x" author="Szumik, C" journalOrPublisher="Cladistics" pageId="0" pageNumber="129" pagination="41 - 64" refId="B44" refString="Szumik, C, 1996. The higher classification of the order Embioptera: a cladistic analysis. Cladistics 12: 41 - 64, DOI: https://doi.org/10.1111/j.1096-0031.1996.tb00192.x" title="The higher classification of the order Embioptera: a cladistic analysis." url="https://doi.org/10.1111/j.1096-0031.1996.tb00192.x" volume="12" year="1996">Szumik 1996</bibRefCitation>
: 51, phylogenetic analysis;
<bibRefCitation author="Szumik, C" journalOrPublisher="Journal of the New York Entomological Society" pageId="0" pageNumber="129" pagination="140 - 153" refId="B45" refString="Szumik, C, 1997. Two new neotropical genera of Embiidae (Embioptera, Insecta). Journal of the New York Entomological Society 105: 140 - 153" title="Two new neotropical genera of Embiidae (Embioptera, Insecta)." volume="105" year="1997">Szumik 1997</bibRefCitation>
: 141, phylogenetic relationships;
<bibRefCitation author="Szumik, C" editor="Coscaron, S" journalOrPublisher="Ediciones del Sur, Argentina" pageId="0" pageNumber="129" pagination="32 - 37" refId="B46" refString="Szumik, C, 1998a. Capitulo 4: Embioptera. In: Coscaron, S, Morrone, J, Eds., Biodiversidad de artropodos argentinos. Ediciones del Sur, Argentina: 32 - 37" title="Capitulo 4: Embioptera." volumeTitle="Biodiversidad de artropodos argentinos." year="1998 a">Szumik 1998a</bibRefCitation>
: 34, new record for the genus in Argentina;
<bibRefCitation author="Ross, ES" journalOrPublisher="Occasional Papers of the California Academy of Sciences" pageId="0" pageNumber="129" pagination="1 - 86" refId="B39" refString="Ross, ES, 2001. Embia. Contributions to the Biosystematics of the Insect Order Embiidina. Part 3. The Embiidae of the Americas (Order Embiidina). Occasional Papers of the California Academy of Sciences 150: 1 - 86" title="Embia. Contributions to the Biosystematics of the Insect Order Embiidina. Part 3. The Embiidae of the Americas (Order Embiidina)." volume="150" year="2001">Ross 2001</bibRefCitation>
: 4, diagnosis and redescription;
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
: 222, phylogenetic analysis, diagnosis and delimitation of the genus, type species
<taxonomicName baseAuthorityName="Navas" baseAuthorityYear="1925" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia kotzbaueri" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="kotzbaueri">Archembia kotzbaueri</taxonomicName>
(Navas, 1925), senior synonym of
<taxonomicName family="Archembiidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="A. lacombea" order="Embioptera" pageId="0" pageNumber="129" rank="species" species="lacombea">A. lacombea</taxonomicName>
;
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-0031.2008.00228.x" author="Szumik, C" journalOrPublisher="Cladistics" pageId="0" pageNumber="129" pagination="993 - 1005" refId="B52" refString="Szumik, C, Edgerly, J, Hayashi, C, 2008. Phylogeny of Embiopterans (Insecta). Cladistics 24: 993 - 1005, DOI: https://doi.org/10.1111/j.1096-0031.2008.00228.x" title="Phylogeny of Embiopterans (Insecta)." url="https://doi.org/10.1111/j.1096-0031.2008.00228.x" volume="24" year="2008">Szumik et al. 2008</bibRefCitation>
: 1003, phylogenetic analysis;
<bibRefCitation author="Szumik, C" editor="Rafael, JA" journalOrPublisher="Holos Editora, Ribeirao Preto" pageId="0" pageNumber="129" pagination="348 - 354" refId="B50" refString="Szumik, C, 2012. Capitulo 20. Embioptera. In: Rafael, JA, Melo, GAR, Carvalho, CJB, Casari, SAC, Constatino, R, Eds., Insetos do Brasil, Diversidade e Taxonomia. Holos Editora, Ribeirao Preto: 348 - 354" title="Capitulo 20. Embioptera." volumeTitle="Insetos do Brasil, Diversidade e Taxonomia." year="2012">Szumik 2012</bibRefCitation>
: 352, composition;
<bibRefCitation DOI="https://doi.org/10.1206/3933.1" author="Szumik, C" journalOrPublisher="Novitates" pageId="0" pageNumber="129" pagination="1 - 31" refId="B54" refString="Szumik, C, Juarez, ML, Ramirez, MJ, Goloboff, P, Pereyra, V, 2019. Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera. Novitates 3933: 1 - 31, DOI: https://doi.org/10.1206/3933.1" title="Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera." url="https://doi.org/10.1206/3933.1" volume="3933" year="2019">Szumik et al. 2019</bibRefCitation>
: 22, tympanal hearing, silk ejectors, leg chaetotaxy, phylogeny.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="diagnosis">
<paragraph pageId="0" pageNumber="129">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="129">
<taxonomicName authorityName="Ross" authorityYear="1971" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia</emphasis>
</taxonomicName>
differs from the close related genera
<taxonomicName authorityName="Enderlein" authorityYear="1909" class="Insecta" family="Archembiidae" genus="Calamoclostes" kingdom="Animalia" lsidName="Calamoclostes" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Calamoclostes</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Szumik" authorityYear="2004" class="Insecta" family="Scelembiidae" genus="Ecuadembia" kingdom="Animalia" lsidName="Ecuadembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Ecuadembia</emphasis>
</taxonomicName>
by having mandibles with incisive teeth concentrated in the apex, anterior edge of Sm diffuse; apical cerci longer than basal cerci, and medial position of LC1dp (
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="distribution">
<paragraph pageId="0" pageNumber="129">Composition and distribution.</paragraph>
<paragraph pageId="0" pageNumber="129">
According to
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik (2004)</bibRefCitation>
<taxonomicName authorityName="Ross" authorityYear="1971" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia</emphasis>
</taxonomicName>
includes six species, one is known exclusively from Bolivia,
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia boliviana" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia boliviana</emphasis>
</taxonomicName>
Ross, 2001 and the other five originally described from Brazil, but recorded in other countries:
<taxonomicName baseAuthorityName="Navas" baseAuthorityYear="1925" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia kotzbaueri" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="kotzbaueri">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia kotzbaueri</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Navás">Navas</normalizedToken>
, 1925),
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia bahia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="bahia">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia bahia</emphasis>
</taxonomicName>
Ross, 2001,
<taxonomicName baseAuthorityName="McLachlan" baseAuthorityYear="1877" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia batesi" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="batesi">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia batesi</emphasis>
</taxonomicName>
McLachlan, 1877 also present in Peru,
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia dilata" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="dilata">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia dilata</emphasis>
</taxonomicName>
Ross, 2001 also present in Argentina, and
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia paranae" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="paranae">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia paranae</emphasis>
</taxonomicName>
Ross, 2001, exclusively from Brazil (see Catalog). Here, one new species is described from Brazil. Almost all the species of the genus were described by
<bibRefCitation author="Ross, ES" journalOrPublisher="The Wasmann Journal of Biology" pageId="0" pageNumber="129" pagination="29 - 36" refId="B34" refString="Ross, ES, 1971. A new neotropical genus and species of Embioptera. The Wasmann Journal of Biology 29: 29 - 36" title="A new neotropical genus and species of Embioptera." volume="29" year="1971">Ross (1971</bibRefCitation>
,
<bibRefCitation author="Ross, ES" journalOrPublisher="Occasional Papers of the California Academy of Sciences" pageId="0" pageNumber="129" pagination="1 - 86" refId="B39" refString="Ross, ES, 2001. Embia. Contributions to the Biosystematics of the Insect Order Embiidina. Part 3. The Embiidae of the Americas (Order Embiidina). Occasional Papers of the California Academy of Sciences 150: 1 - 86" title="Embia. Contributions to the Biosystematics of the Insect Order Embiidina. Part 3. The Embiidae of the Americas (Order Embiidina)." volume="150" year="2001">2001</bibRefCitation>
), and many of them were not illustrated. In some cases, the criteria used for the creation of a new species were differences on coloration, and a few of these species are now designated junior synonyms (
<taxonomicName family="Archembiidae" kingdom="Animalia" lsidName="A. peruviana" order="Embioptera" pageId="0" pageNumber="129" rank="species" species="peruviana">
<emphasis italics="true" pageId="0" pageNumber="129">A. peruviana</emphasis>
</taxonomicName>
Ross, 2001 and
<taxonomicName family="Archembiidae" kingdom="Animalia" lsidName="A. lacombea" order="Embioptera" pageId="0" pageNumber="129" rank="species" species="lacombea">
<emphasis italics="true" pageId="0" pageNumber="129">A. lacombea</emphasis>
</taxonomicName>
Ross, 1971; see Catalog) or transferred to another genus (
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia arida" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="arida">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia arida</emphasis>
</taxonomicName>
Ross, 2001 from Ecuador now
<taxonomicName baseAuthorityName="Ross" baseAuthorityYear="2001" class="Insecta" family="Scelembiidae" genus="Ecuadembia" kingdom="Animalia" lsidName="Ecuadembia arida" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="species" species="arida">
<emphasis italics="true" pageId="0" pageNumber="129">Ecuadembia arida</emphasis>
</taxonomicName>
(Ross); see
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="129">
Thanks to the observations on
<normalizedToken originalValue="Rosss">Ross's</normalizedToken>
collection at CAS as well as material deposited at INPA, MZUSP, MCZ, and USNM we have a better understanding of the distribution of this genus, with one species being present in the Amazon basin (
<taxonomicName family="Archembiidae" kingdom="Animalia" lsidName="A. batesi" order="Embioptera" pageId="0" pageNumber="129" rank="species" species="batesi">
<emphasis italics="true" pageId="0" pageNumber="129">A. batesi</emphasis>
</taxonomicName>
), four species present in the Atlantic Forest (
<taxonomicName family="Archembiidae" kingdom="Animalia" lsidName="A. kotzbaueri" order="Embioptera" pageId="0" pageNumber="129" rank="species" species="kotzbaueri">
<emphasis italics="true" pageId="0" pageNumber="129">A. kotzbaueri</emphasis>
</taxonomicName>
,
<taxonomicName family="Archembiidae" kingdom="Animalia" lsidName="A. dilata" order="Embioptera" pageId="0" pageNumber="129" rank="species" species="dilata">
<emphasis italics="true" pageId="0" pageNumber="129">A. dilata</emphasis>
</taxonomicName>
,
<taxonomicName family="Archembiidae" kingdom="Animalia" lsidName="A. bahia" order="Embioptera" pageId="0" pageNumber="129" rank="species" species="bahia">
<emphasis italics="true" pageId="0" pageNumber="129">A. bahia</emphasis>
</taxonomicName>
,
<taxonomicName family="Archembiidae" kingdom="Animalia" lsidName="A. paranae" order="Embioptera" pageId="0" pageNumber="129" rank="species" species="paranae">
<emphasis italics="true" pageId="0" pageNumber="129">A. paranae</emphasis>
</taxonomicName>
), and a new species described from the Cerrado and Pantanal ecoregion.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="distribution">
<paragraph pageId="0" pageNumber="129">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="129">South America.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="129" type="relationships">
<paragraph pageId="0" pageNumber="129">Relationships.</paragraph>
<paragraph pageId="0" pageNumber="129">
Several cladistic analyses suggest that
<taxonomicName authorityName="Ross" authorityYear="1971" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia</emphasis>
</taxonomicName>
is a well-supported genus (
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-0031.2008.00228.x" author="Szumik, C" journalOrPublisher="Cladistics" pageId="0" pageNumber="129" pagination="993 - 1005" refId="B52" refString="Szumik, C, Edgerly, J, Hayashi, C, 2008. Phylogeny of Embiopterans (Insecta). Cladistics 24: 993 - 1005, DOI: https://doi.org/10.1111/j.1096-0031.2008.00228.x" title="Phylogeny of Embiopterans (Insecta)." url="https://doi.org/10.1111/j.1096-0031.2008.00228.x" volume="24" year="2008">Szumik et al. 2008</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1206/3933.1" author="Szumik, C" journalOrPublisher="Novitates" pageId="0" pageNumber="129" pagination="1 - 31" refId="B54" refString="Szumik, C, Juarez, ML, Ramirez, MJ, Goloboff, P, Pereyra, V, 2019. Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera. Novitates 3933: 1 - 31, DOI: https://doi.org/10.1206/3933.1" title="Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera." url="https://doi.org/10.1206/3933.1" volume="3933" year="2019">2019</bibRefCitation>
).
<taxonomicName authorityName="Ross" authorityYear="1971" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia</emphasis>
</taxonomicName>
is one of the basal genera of
<taxonomicName authorityName="Ross" authorityYear="2001" class="Insecta" family="Archembiidae" kingdom="Animalia" lsidName="" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="family">Archembiidae</taxonomicName>
. The genera
<taxonomicName authorityName="Szumik" authorityYear="2004" class="Insecta" family="Scelembiidae" genus="Ecuadembia" kingdom="Animalia" lsidName="Ecuadembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Ecuadembia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Enderlein" authorityYear="1909" class="Insecta" family="Archembiidae" genus="Calamoclostes" kingdom="Animalia" lsidName="Calamoclostes" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Calamoclostes</emphasis>
</taxonomicName>
are the sister group of
<taxonomicName authorityName="Ross" authorityYear="1971" class="Insecta" family="Archembiidae" genus="Archembia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archembia" order="Embioptera" pageId="0" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="129">Archembia</emphasis>
</taxonomicName>
they share the type B vein origin, a large anal area, the straight 10Lp1 with a spatulate apex and the Ep fused to the 10Rp1 (
<bibRefCitation DOI="https://doi.org/10.1111/j.0307-6970.2004.00239.x" author="Szumik, C" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="129" pagination="215 - 237" refId="B49" refString="Szumik, C, 2004. Phylogenetic systematics of Archembiidae (Embiidina, Insecta). Systematic Entomology 29: 215 - 237, DOI: https://doi.org/10.1111/j.0307-6970.2004.00239.x" title="Phylogenetic systematics of Archembiidae (Embiidina, Insecta)." url="https://doi.org/10.1111/j.0307-6970.2004.00239.x" volume="29" year="2004">Szumik 2004</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1206/3933.1" author="Szumik, C" journalOrPublisher="Novitates" pageId="0" pageNumber="129" pagination="1 - 31" refId="B54" refString="Szumik, C, Juarez, ML, Ramirez, MJ, Goloboff, P, Pereyra, V, 2019. Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera. Novitates 3933: 1 - 31, DOI: https://doi.org/10.1206/3933.1" title="Implications of the tympanal hearing organ and ultrastructure of chaetotaxy for the higher classification of Embioptera." url="https://doi.org/10.1206/3933.1" volume="3933" year="2019">Szumik et al. 2019</bibRefCitation>
); the three genera conform the subfamily
<taxonomicName authorityName="Ross" authorityYear="2001" lsidName="" pageId="0" pageNumber="129" rank="subFamily" subFamily="Archembiinae">Archembiinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>