treatments-xml/data/0C/7A/87/0C7A87DECB78FF98FF10FA67FF361C64.xml
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<document id="9B46847A7AFAFDF535069BA4FCCCD043" ID-CLB-Dataset="3683" ID-DOI="10.11646/zootaxa.5060.4.9" ID-GBIF-Dataset="31872afa-34e2-48c8-800a-43acaeaf071c" ID-ISSN="1175-5326" ID-Zenodo-Dep="5638064" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635837831356" checkinUser="plazi" docAuthor="Amaral, André P., Bello-González, Orestes C. &amp; Pinho, Luiz Carlos" docDate="2021" docId="0C7A87DECB78FF98FF10FA67FF361C64" docLanguage="en" docName="zootaxa.5060.4.9.pdf" docOrigin="Zootaxa 5060 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Corethrella (Corethrella) stenostyla Amaral &amp; Bello-González &amp; Pinho 2021, sp. n." docType="treatment" docUuid="1BEA6C85-D4BC-4EE7-B59E-7112B57D4EE8" docUuidSource="ZooBank" docVersion="7" lastPageNumber="598" masterDocId="F043FFA6CB78FF9BFF87FFE2FFE21F3F" masterDocTitle="Corethrella stenostyla sp. n., a new frog-biting midge (Diptera: Corethrellidae) from Cuba" masterLastPageNumber="598" masterPageNumber="595" pageNumber="595" updateTime="1699467431541" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2B3500BDFEFE9BA5EBAB730F3627FAF6">Corethrella stenostyla sp. n., a new frog-biting midge (Diptera: Corethrellidae) from Cuba</mods:title>
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<mods:namePart id="3180EE209C5B8FDF6F7AA3B44BA7F4F1">Amaral, André P.</mods:namePart>
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<mods:namePart id="DB3DA71CA84CA5D3AF19889630F5C85F">Bello-González, Orestes C.</mods:namePart>
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<mods:namePart id="3F1533BB075C9CF312D57B43A008E7DA">Pinho, Luiz Carlos</mods:namePart>
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<treatment id="0C7A87DECB78FF98FF10FA67FF361C64" ID-DOI="http://doi.org/10.5281/zenodo.5638062" ID-GBIF-Taxon="190002181" ID-Zenodo-Dep="5638062" LSID="urn:lsid:zoobank.org:act:1BEA6C85-D4BC-4EE7-B59E-7112B57D4EE8" httpUri="http://treatment.plazi.org/id/0C7A87DECB78FF98FF10FA67FF361C64" lastPageId="3" lastPageNumber="598" pageId="0" pageNumber="595">
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<paragraph id="846C36C8CB78FF9BFF10FA67FD6D1A9F" blockId="0.[151,1437,1413,2016]" box="[151,655,1413,1440]" pageId="0" pageNumber="595">
<heading id="DF2481A4CB78FF9BFF10FA67FD6D1A9F" bold="true" box="[151,655,1413,1440]" fontSize="11" level="1" pageId="0" pageNumber="595" reason="1">
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<taxonomicName id="43D34D4BCB78FF9BFF10FA67FDA71A9F" ID-CoL="9HYLZ" authority="Amaral &amp; Bello-González &amp; Pinho, 2021" authorityName="Amaral &amp; Bello-González &amp; Pinho" authorityYear="2021" box="[151,581,1413,1440]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="595" phylum="Arthropoda" rank="species" species="stenostyla" status="sp. nov." subGenus="Corethrella">
<emphasis id="B6A7EADACB78FF9BFF10FA67FEC01A9F" bold="true" box="[151,290,1413,1440]" italics="true" pageId="0" pageNumber="595">Corethrella</emphasis>
(
<emphasis id="B6A7EADACB78FF9BFEB4FA67FE5C1A9F" bold="true" box="[307,446,1413,1440]" italics="true" pageId="0" pageNumber="595">Corethrella</emphasis>
)
<emphasis id="B6A7EADACB78FF9BFE49FA67FDA71A9F" bold="true" box="[462,581,1413,1440]" italics="true" pageId="0" pageNumber="595">stenostyla</emphasis>
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<taxonomicNameLabel id="AD9457A1CB78FF9BFDCAFA64FD6D1A9F" box="[589,655,1414,1440]" pageId="0" pageNumber="595" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="846C36C8CB78FF9BFF10FA4EFEC31AFB" blockId="0.[151,1437,1413,2016]" box="[151,289,1452,1476]" pageId="0" pageNumber="595">
(
<figureCitation id="1CE82A4DCB78FF9BFF18FA4EFF0A1AFB" box="[159,232,1452,1476]" captionStart="FIGURE 1" captionStartId="1.[151,250,1152,1176]" captionTargetBox="[151,1436,271,1127]" captionTargetId="figure-66@1.[151,1436,271,1127]" captionTargetPageId="1" captionText="FIGURE 1. Collection site in La Bayamesa National Park, Cuba." figureDoi="http://doi.org/10.5281/zenodo.5638066" httpUri="https://zenodo.org/record/5638066/files/figure.png" pageId="0" pageNumber="595">Figs. 1</figureCitation>
&amp;
<figureCitation id="1CE82A4DCB78FF9BFE8CFA4EFEFB1AFB" box="[267,281,1452,1476]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="0" pageNumber="595">2</figureCitation>
)
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urn:lsid:zoobank.org:act:
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<subSubSection id="CCC96543CB78FF9BFF10FA16FD43196B" pageId="0" pageNumber="595" type="materials_examined">
<paragraph id="846C36C8CB78FF9BFF10FA16FD43196B" blockId="0.[151,1437,1413,2016]" pageId="0" pageNumber="595">
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<emphasis id="B6A7EADACB78FF9BFF10FA16FED51934" bold="true" box="[151,311,1523,1548]" pageId="0" pageNumber="595">Type material</emphasis>
.
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<materialsCitation id="34BB3C95CB78FF9BFEC0FA16FD43196B" ID-GBIF-Occurrence="3398380301" collectingDate="2015-12" collectionCode="CEMHS" collectorName="Orestes C. Bello-Gonzalez" country="Cuba" county="Guisa" location="Pico La Bayamesa'" municipality="La Bayamesa National Park" pageId="0" pageNumber="595" specimenCount="1" specimenCount-male="1" stateProvince="Granma" typeStatus="holotype">
<typeStatus id="5B68886ACB78FF9BFEC0FA16FE4C1933" box="[327,430,1524,1548]" pageId="0" pageNumber="595" type="holotype">Holotype</typeStatus>
: male adult, slide mounted: “HOLOTYPE
<taxonomicName id="43D34D4BCB78FF9BFC03FA16FF37190F" authority="Amaral, Bello-Gonzalez &amp; Pinho" authorityName="Amaral, Bello-Gonzalez &amp; Pinho" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="595" phylum="Arthropoda" rank="species" species="stenostyla">
<emphasis id="B6A7EADACB78FF9BFC03FA16FB901933" box="[900,1138,1524,1548]" italics="true" pageId="0" pageNumber="595">Corethrella stenostyla</emphasis>
Amaral, Bello-González &amp; Pinho
</taxonomicName>
<specimenCount id="92D5FD41CB78FF9BFF5AF9FAFF111910" box="[221,243,1560,1583]" pageId="0" pageNumber="595" type="male"></specimenCount>
,
<collectingCountry id="FCC47658CB78FF9BFF78F9FAFEAE190F" box="[255,332,1560,1584]" name="Cuba" pageId="0" pageNumber="595">CUBA</collectingCountry>
,
<collectingRegion id="4617F82ACB78FF9BFED0F9FAFDF4190F" box="[343,534,1560,1584]" country="Cuba" name="Granma" pageId="0" pageNumber="595">Granma Province</collectingRegion>
,
<collectingCounty id="6D0D4E44CB78FF9BFDA7F9FAFD83190F" box="[544,609,1560,1584]" pageId="0" pageNumber="595">Guisa</collectingCounty>
,
<collectingMunicipality id="6408ACB2CB78FF9BFDEBF9FAFC7E190F" box="[620,924,1560,1584]" pageId="0" pageNumber="595">La Bayamesa National Park</collectingMunicipality>
,
<quantity id="432B9B2DCB78FF9BFC21F9FAFC34190F" box="[934,982,1560,1584]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="595" unit="km" value="1.0">1km</quantity>
of
<location id="810C6013CB78FF9BFC78F9FAFB34190F" LSID="urn:lsid:plazi:treatment:0C7A87DECB78FF98FF10FA67FF361C64:810C6013CB78FF9BFC78F9FAFB34190F" box="[1023,1238,1560,1584]" country="Cuba" county="Guisa" municipality="La Bayamesa National Park" name="Pico La Bayamesa'" pageId="0" pageNumber="595" stateProvince="Granma">Pico La Bayamesa</location>
,
<date id="F06D1008CB78FF9BFB65F9FAFADE190F" box="[1250,1340,1560,1584]" pageId="0" pageNumber="595" value="2015-12">
<collectingDate id="E029E9E0CB78FF9BFB65F9FAFADE190F" box="[1250,1340,1560,1584]" pageId="0" pageNumber="595" value="2015-12">xii.2015</collectingDate>
</date>
,
<collectingMethod id="5D924EDFCB78FF9BFACFF9FAFF21196C" pageId="0" pageNumber="595">Malaise trap</collectingMethod>
,
<collectorName id="2926531ECB78FF9BFF49F9DEFE09196B" box="[206,491,1596,1620]" pageId="0" pageNumber="595">Orestes C. Bello-González</collectorName>
leg.” (
<collectionCode id="E2C2AE0DCB78FF9BFDB4F9DEFD77196B" box="[563,661,1596,1620]" pageId="0" pageNumber="595">CEMHS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CCC96543CB78FF9BFF40F982FB6319DB" pageId="0" pageNumber="595" type="etymology">
<paragraph id="846C36C8CB78FF9BFF40F982FADF1947" blockId="0.[151,1437,1413,2016]" box="[199,1341,1632,1656]" pageId="0" pageNumber="595">
<emphasis id="B6A7EADACB78FF9BFF40F982FEAB1948" box="[199,329,1632,1656]" inLineHeading="true" pageId="0" pageNumber="595" reason="1">
<emphasis id="B6A7EADACB78FF9BFF40F982FEA01947" bold="true" box="[199,322,1632,1656]" pageId="0" pageNumber="595">Etymology</emphasis>
.
</emphasis>
The specific epithet refers to the slender shape of the gonostylus, also present in other species.
</paragraph>
<paragraph id="846C36C8CB78FF9BFF40F961FB6319DB" blockId="0.[151,1437,1413,2016]" pageId="0" pageNumber="595">
<emphasis id="B6A7EADACB78FF9BFF40F961FE2519A4" box="[199,455,1667,1692]" inLineHeading="true" pageId="0" pageNumber="595" reason="1">
<emphasis id="B6A7EADACB78FF9BFF40F961FE2319A3" bold="true" box="[199,449,1667,1692]" pageId="0" pageNumber="595">Diagnostic characters</emphasis>
.
</emphasis>
Male adults: only extant species in the New World with a plain wing; ventral portion of posterior anepisternum triangular, without a thickened anterodorsal margin; thorax brown, ventral portion of anepimeron, trochanters and bases of hind and midfemora pale; foreleg third tarsomere shorter than fourth.
</paragraph>
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<subSubSection id="CCC96543CB78FF98FF40F90DFDD81F90" lastPageId="3" lastPageNumber="598" pageId="0" pageNumber="595" type="description">
<paragraph id="846C36C8CB78FF9BFF40F90DFDF61837" blockId="0.[151,1437,1413,2016]" box="[199,532,1775,1800]" pageId="0" pageNumber="595">
<emphasis id="B6A7EADACB78FF9BFF40F90DFEB01838" box="[199,338,1775,1799]" inLineHeading="true" pageId="0" pageNumber="595" reason="1">
<emphasis id="B6A7EADACB78FF9BFF40F90DFEA91838" bold="true" box="[199,331,1775,1799]" pageId="0" pageNumber="595">Description</emphasis>
.
</emphasis>
Male adult (n=1).
</paragraph>
<paragraph id="846C36C8CB78FF9AFF40F8F6FE8D1FC8" blockId="0.[151,1437,1413,2016]" lastBlockId="1.[151,1437,151,248]" lastPageId="1" lastPageNumber="596" pageId="0" pageNumber="595">
<emphasis id="B6A7EADACB78FF9BFF40F8F6FEE31813" box="[199,257,1812,1836]" italics="true" pageId="0" pageNumber="595">Head</emphasis>
(
<figureCitation id="1CE82A4DCB78FF9BFE95F8F6FE881813" box="[274,362,1812,1836]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="0" pageNumber="595">Fig. 2A</figureCitation>
). Medium to dark brown. Outline in anterior view moderately laterally elongate, 1.43 times wider than long. Coronal suture complete, reaching ventral margin of interocular space. Clypeus (
<figureCitation id="1CE82A4DCB78FF9BFBEEF8DAFB59186F" box="[1129,1211,1848,1872]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="0" pageNumber="595">Fig. 2B</figureCitation>
) square, 1.08x wider than long, with 3 setae. Palpus (
<figureCitation id="1CE82A4DCB78FF9BFE6EF8BEFD84184B" box="[489,614,1884,1908]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="0" pageNumber="595">Figs. 2A, C</figureCitation>
) slightly less pigmented than head, third segment with constant width, length of fifth 0.83. Antenna (
<figureCitation id="1CE82A4DCB78FF9BFE16F862FE0A18A7" box="[401,488,1920,1944]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="0" pageNumber="595">Fig. 2D</figureCitation>
): pedicel dark brown, flagellum medium to dark brown with very base of flagellomeres IIXIII pale; flagellomeres (
<figureCitation id="1CE82A4DCB78FF9BFE4CF846FDFC1883" box="[459,542,1956,1980]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="0" pageNumber="595">Fig. 2E</figureCitation>
) I, III elongate, II somewhat short; flagellomere XIII apically bifurcated. Sensilla coeloconica distribution: 1(IXXIII), 4(I).
<emphasis id="B6A7EADACB78FF9BFDEAF825FD2618E0" bold="true" box="[621,708,1991,2015]" pageId="0" pageNumber="595">Sensilla</emphasis>
(
<figureCitation id="1CE82A4DCB78FF9BFD50F82AFCCD18DF" box="[727,815,1992,2016]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="0" pageNumber="595">Fig. 2B</figureCitation>
): Ocular row with 1 thick offset seta at ventral part, 12 more dorsally, extending a little beyond vertex. With subocular row not well-defined, many slender setae on vertex and interocular space. Postgenal row with 10 intermediate setae, contiguous to ocular row, extending to ventral margin. With 1 ventromedial seta.
</paragraph>
<caption id="D0AC6640CB79FF9AFF10FB62FCD01BA7" ID-DOI="http://doi.org/10.5281/zenodo.5638066" ID-Zenodo-Dep="5638066" box="[151,818,1152,1176]" httpUri="https://zenodo.org/record/5638066/files/figure.png" pageId="1" pageNumber="596" startId="1.[151,250,1152,1176]" targetBox="[151,1436,271,1127]" targetPageId="1">
<paragraph id="846C36C8CB79FF9AFF10FB62FCD01BA7" blockId="1.[151,818,1152,1176]" box="[151,818,1152,1176]" pageId="1" pageNumber="596">
<emphasis id="B6A7EADACB79FF9AFF10FB62FEF11BA7" bold="true" box="[151,275,1152,1176]" pageId="1" pageNumber="596">FIGURE 1.</emphasis>
Collection site in La Bayamesa National Park, Cuba.
</paragraph>
</caption>
<paragraph id="846C36C8CB79FF9AFF40FB2AFE88191C" blockId="1.[151,1437,1224,2040]" pageId="1" pageNumber="596">
<emphasis id="B6A7EADACB79FF9AFF40FB2AFEF61BDF" box="[199,276,1224,1248]" italics="true" pageId="1" pageNumber="596">Thorax</emphasis>
(
<figureCitation id="1CE82A4DCB79FF9AFEA2FB2AFE461BDF" box="[293,420,1224,1248]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="1" pageNumber="596">Figs. 2F, G</figureCitation>
). Medium to dark brown; ventral portion of anepimeron and hind coxa pale, sclerites around wing light brown. Prescutal suture moderately elongate, fading dorsally. Anterior anepisternum divided longitudinally by sinuous suture. Posterior anepisternum triangular, with anterodorsal margin not thick.
<emphasis id="B6A7EADACB79FF9AFBABFAF2FB611A17" bold="true" box="[1068,1155,1296,1320]" pageId="1" pageNumber="596">Sensilla</emphasis>
(
<figureCitation id="1CE82A4DCB79FF9AFB15FAF2FB051A17" box="[1170,1255,1296,1320]" captionStart="FIGURE 1" captionStartId="1.[151,250,1152,1176]" captionTargetBox="[151,1436,271,1127]" captionTargetId="figure-66@1.[151,1436,271,1127]" captionTargetPageId="1" captionText="FIGURE 1. Collection site in La Bayamesa National Park, Cuba." figureDoi="http://doi.org/10.5281/zenodo.5638066" httpUri="https://zenodo.org/record/5638066/files/figure.png" pageId="1" pageNumber="596">Fig. 1H</figureCitation>
): Antepronotum, 13 intermediate setae on anterior and lateral portions. Postpronotum with 1 thick, dorsal seta and 3 intermediate, more posterior setae. Scutum, prescutal area with 8 intermediate and slender setae located anteriorly, 2 thick and 1 intermediate near prescutal suture. Antealar area with 3 thick setae and 1 slender seta near prescutal suture, 13 thick and intermediate setae spread posteriorly on ventral portion. Supraalar area with 2 thick setae on posterior portion, 6 slender and 4 intermediate ones more anteriorly. Dorsocentral row, posterior part with 4 thick offset setae, about 26 thick and intermediate, 13 slender setae filling the row. Scutellum with 12 thick setae. Posterior anepisternum bare. Anepimeron with 3 slender posterodorsal setae.
</paragraph>
<paragraph id="846C36C8CB79FF9AFF40F9D3FE4E19AF" blockId="1.[151,1437,1224,2040]" pageId="1" pageNumber="596">
<emphasis id="B6A7EADACB79FF9AFF40F9D3FF1C1977" box="[199,254,1585,1608]" italics="true" pageId="1" pageNumber="596">Wing</emphasis>
(
<figureCitation id="1CE82A4DCB79FF9AFE89F9D2FE6B1977" box="[270,393,1584,1608]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="1" pageNumber="596">Figs. 2J, K</figureCitation>
). Medium brown, somewhat greyish. non-marginal veins with narrow scales. Without pattern of pigmentation. Apex of R
<subScript id="1857348DCB79FF9AFE26F986FE4B194D" attach="left" box="[417,425,1636,1650]" fontSize="6" pageId="1" pageNumber="596">2</subScript>
slightly basal to apex of M
<subScript id="1857348DCB79FF9AFD56F986FD3B194D" attach="left" box="[721,729,1636,1650]" fontSize="6" pageId="1" pageNumber="596">1</subScript>
. Ratio of lengths of R
<subScript id="1857348DCB79FF9AFC4DF986FC30194D" attach="left" box="[970,978,1636,1650]" fontSize="6" pageId="1" pageNumber="596">3</subScript>
/R
<subScript id="1857348DCB79FF9AFC6CF986FC11194D" attach="left" box="[1003,1011,1636,1650]" fontSize="6" pageId="1" pageNumber="596">1</subScript>
: 0.64; R
<subScript id="1857348DCB79FF9AFBD6F986FB8B194D" attach="left" box="[1105,1129,1636,1650]" fontSize="6" pageId="1" pageNumber="596">2+3</subScript>
/R
<subScript id="1857348DCB79FF9AFB05F986FB68194D" attach="left" box="[1154,1162,1636,1650]" fontSize="6" pageId="1" pageNumber="596">2</subScript>
: 0.37. Pigment of haltere same as that of scutellum.
</paragraph>
<paragraph id="846C36C8CB79FF9AFF40F97FFE1B181F" blockId="1.[151,1437,1224,2040]" pageId="1" pageNumber="596">
<emphasis id="B6A7EADACB79FF9AFF40F97FFF1B198B" box="[199,249,1693,1716]" italics="true" pageId="1" pageNumber="596">Legs</emphasis>
(
<figureCitation id="1CE82A4DCB79FF9AFE8EF97EFEB9198B" box="[265,347,1692,1716]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="1" pageNumber="596">Fig. 2F</figureCitation>
). Medium brown, with trochanters and bases of hind and midfemora pale. Tibiae and femora without scales. Midleg tarsus apparently lacking distinctively thick subapical setae. Ratio of lengths of Ta1/Ta2: 3.00; Ta3/Ta4: 0.52. Tarsal claws of foreleg unequal and longer than hind and midleg claws. Empodium (
<figureCitation id="1CE82A4DCB79FF9AFBDBF906FB4719C3" box="[1116,1189,1764,1788]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="1" pageNumber="596">Fig. 2I</figureCitation>
) of intermediate thickness and length, with 4 branches.
</paragraph>
<paragraph id="846C36C8CB79FF9AFF40F8CEFAA2187B" blockId="1.[151,1437,1224,2040]" box="[199,1344,1836,1860]" pageId="1" pageNumber="596">
<emphasis id="B6A7EADACB79FF9AFF40F8CEFECE187B" box="[199,300,1836,1860]" italics="true" pageId="1" pageNumber="596">Abdomen</emphasis>
(
<figureCitation id="1CE82A4DCB79FF9AFEBCF8CEFE6C187B" box="[315,398,1836,1860]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="1" pageNumber="596">Fig. 2L</figureCitation>
). Medium brown, with segments 79 and gonocoxite darker. Segment 8 pale posteriorly.
</paragraph>
<paragraph id="846C36C8CB79FF9AFF40F8B2FC7118C7" blockId="1.[151,1437,1224,2040]" pageId="1" pageNumber="596">
<emphasis id="B6A7EADACB79FF9AFF40F8B2FECE1857" box="[199,300,1872,1896]" italics="true" pageId="1" pageNumber="596">Genitalia</emphasis>
(
<figureCitation id="1CE82A4DCB79FF9AFEBEF8B2FE5E1857" box="[313,444,1872,1896]" captionStart="FIGURE 2" captionStartId="2.[151,250,1943,1967]" captionTargetBox="[192,1405,187,1910]" captionTargetId="figure-24@2.[179,1408,181,1920]" captionTargetPageId="2" captionText="FIGURE 2. Corethrella stenostyla sp. n., adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia." figureDoi="http://doi.org/10.5281/zenodo.5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="1" pageNumber="596">Figs. 2M, N</figureCitation>
). Gonocoxite slender, without distinctively elongate spicules on anteromedial area; dorsal row well-defined with 6 stout elongate setae restricted to dorsal region; 1 dorsomedial stout seta, tapering from base, located at 0.17 from base of gonocoxite. Gonostylus slender, with nearly constant width; sub-basal seta simple and elongate, situated anteriorly, at 0.28 from base; gonostylus with blunt, peg-like bristle inserted at apex and 5 small subapical setae. Aedeagus strongly tapering from midlength, margins fused subapically.
</paragraph>
<caption id="D0AC6640CB7AFF99FF10F875FAC018C7" ID-DOI="http://doi.org/10.5281/zenodo.5638068" ID-Zenodo-Dep="5638068" httpUri="https://zenodo.org/record/5638068/files/figure.png" pageId="2" pageNumber="597" startId="2.[151,250,1943,1967]" targetBox="[192,1405,187,1910]" targetPageId="2">
<paragraph id="846C36C8CB7AFF99FF10F875FAC018C7" blockId="2.[151,1436,1943,2040]" pageId="2" pageNumber="597">
<emphasis id="B6A7EADACB7AFF99FF10F875FEF11890" bold="true" box="[151,275,1943,1967]" pageId="2" pageNumber="597">FIGURE 2.</emphasis>
<taxonomicName id="43D34D4BCB7AFF99FE9DF87AFE19188F" authority="Amaral &amp; Bello-González &amp; Pinho, 2021" authorityName="Amaral &amp; Bello-González &amp; Pinho" authorityYear="2021" box="[282,507,1944,1968]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="stenostyla" status="sp. nov.">
<emphasis id="B6A7EADACB7AFF99FE9DF87AFE19188F" box="[282,507,1944,1968]" italics="true" pageId="2" pageNumber="597">Corethrella stenostyla</emphasis>
</taxonomicName>
<emphasis id="B6A7EADACB7AFF99FD85F87AFDD9188F" bold="true" box="[514,571,1944,1968]" pageId="2" pageNumber="597">
<taxonomicNameLabel id="AD9457A1CB7AFF99FD85F87AFDD9188F" box="[514,571,1944,1968]" pageId="2" pageNumber="597" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult male. A, head in dorsal view. B, cranial setae, anterior and posterior views. C, palpus. D, antenna. E, flagellomeres. F, thorax and legs, lateral view. G, detail of thorax, lateral view. H, thoracic setae. I, empodium. J, wing. K, wing venation. L, abdomen. M, abdominal segments 89 and genitalia, dorsal view. N, genitalia.
</paragraph>
</caption>
<paragraph id="846C36C8CB7BFF98FF40FF75FDD81F90" blockId="3.[151,1437,151,859]" box="[199,570,151,175]" pageId="3" pageNumber="598">
<emphasis id="B6A7EADACB7BFF98FF40FF75FE5E1F90" box="[199,444,151,175]" italics="true" pageId="3" pageNumber="598">Female and immatures</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="CCC96543CB7BFF98FF40FF59FF361C64" pageId="3" pageNumber="598" type="distribution">
<paragraph id="846C36C8CB7BFF98FF40FF59FAEB1FC7" blockId="3.[151,1437,151,859]" pageId="3" pageNumber="598">
<emphasis id="B6A7EADACB7BFF98FF40FF59FE061FEC" box="[199,484,187,212]" inLineHeading="true" pageId="3" pageNumber="598" reason="1">
<emphasis id="B6A7EADACB7BFF98FF40FF59FE3F1FEC" bold="true" box="[199,477,187,212]" pageId="3" pageNumber="598">Distribution and biology</emphasis>
.
</emphasis>
This species is known solely from the male
<typeStatus id="5B68886ACB7BFF98FC3EFF59FBFB1FEC" box="[953,1049,187,211]" pageId="3" pageNumber="598" type="holotype">holotype</typeStatus>
, collected in a Malaise trap set in La Bayamesa National Park,
<collectingCountry id="FCC47658CB7BFF98FE2BFF3DFE051FC8" box="[428,487,223,247]" name="Cuba" pageId="3" pageNumber="598">Cuba</collectingCountry>
, in an area of montane rainforest, at approximately
<quantity id="432B9B2DCB7BFF98FB89FF02FBB51FC7" box="[1038,1111,224,248]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="598" unit="m" value="1400.0">1400m</quantity>
a.s.l. (
<figureCitation id="1CE82A4DCB7BFF98FB19FF3DFB1B1FC7" box="[1182,1273,223,248]" captionStart="FIGURE 1" captionStartId="1.[151,250,1152,1176]" captionTargetBox="[151,1436,271,1127]" captionTargetId="figure-66@1.[151,1436,271,1127]" captionTargetPageId="1" captionText="FIGURE 1. Collection site in La Bayamesa National Park, Cuba." figureDoi="http://doi.org/10.5281/zenodo.5638066" httpUri="https://zenodo.org/record/5638066/files/figure.png" pageId="3" pageNumber="598">Figure 1</figureCitation>
).
</paragraph>
<paragraph id="846C36C8CB7BFF98FF40FEE1FF1F1DD0" blockId="3.[151,1437,151,859]" pageId="3" pageNumber="598">
<emphasis id="B6A7EADACB7BFF98FF40FEE1FE251E24" box="[199,455,259,283]" inLineHeading="true" pageId="3" pageNumber="598" reason="1">
<emphasis id="B6A7EADACB7BFF98FF40FEE1FE221E24" bold="true" box="[199,448,259,283]" pageId="3" pageNumber="598">Taxonomic discussion</emphasis>
.
</emphasis>
<taxonomicName id="43D34D4BCB7BFF98FE49FEE1FD591E24" authority="Amaral &amp; Bello-González &amp; Pinho, 2021" authorityName="Amaral &amp; Bello-González &amp; Pinho" authorityYear="2021" box="[462,699,259,283]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="stenostyla" status="sp. nov.">
<emphasis id="B6A7EADACB7BFF98FE49FEE1FD591E24" box="[462,699,259,283]" italics="true" pageId="3" pageNumber="598">Corethrella stenostyla</emphasis>
</taxonomicName>
<emphasis id="B6A7EADACB7BFF98FD44FEE6FD1D1E23" bold="true" box="[707,767,260,284]" pageId="3" pageNumber="598">
<taxonomicNameLabel id="AD9457A1CB7BFF98FD44FEE6FD1D1E23" box="[707,767,260,284]" pageId="3" pageNumber="598" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is morphologically peculiar. According to the phylogeny proposed by
<bibRefCitation id="E0424B39CB7BFF98FF7BFEC5FE401E7F" author="Borkent, A." box="[252,418,295,320]" pageId="3" pageNumber="598" pagination="1 - 456" refId="ref1985" refString="Borkent, A. (2008) The frog-biting midges of the world (Corethrellidae: Diptera). Zootaxa, 1804, 1 - 456. https: // doi. org / 10.11646 / zootaxa. 1804.1.1" type="journal article" year="2008">Borkent (2008)</bibRefCitation>
, it presents the derived conditions of characters 13, 510, 1318, 21, 2526, 2830, 33, 41, 43, 6365, 6869, 82, but is plesiomorphic to 4244, 7081, 8489. Among the derived traits are: a short third anterior tarsomere, the presence of a slender, anteriorly positioned sub-basal seta on the gonostylus and the gonocoxite with the dorsomedial seta more basally located. Conflicting plesiomorphic characters are the posterior anepisternum without a thick, discrete anterodorsal margin and apex of gonostylus with an apical peg-like bristle instead of a slender seta. Three species,
<taxonomicName id="43D34D4BCB7BFF98FF10FE39FE9C1ECC" authorityName="Borkent" authorityYear="2008" box="[151,382,475,499]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="evenhuisi">
<emphasis id="B6A7EADACB7BFF98FF10FE39FE9C1ECC" box="[151,382,475,499]" italics="true" pageId="3" pageNumber="598">Corethrella evenhuisi</emphasis>
</taxonomicName>
,
<taxonomicName id="43D34D4BCB7BFF98FE0BFE3EFDF71ECC" authorityName="Borkent" authorityYear="2008" box="[396,533,476,499]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="canningsi">
<emphasis id="B6A7EADACB7BFF98FE0BFE3EFDF71ECC" box="[396,533,476,499]" italics="true" pageId="3" pageNumber="598">C. canningsi</emphasis>
</taxonomicName>
and
<taxonomicName id="43D34D4BCB7BFF98FDCDFE3EFD321ECC" authorityName="Borkent" authorityYear="2008" box="[586,720,476,499]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="pauciseta">
<emphasis id="B6A7EADACB7BFF98FDCDFE3EFD321ECC" box="[586,720,476,499]" italics="true" pageId="3" pageNumber="598">C. pauciseta</emphasis>
</taxonomicName>
, belonging to Oceanian region (the latter also present in Orienatal region), present a similar combination of apomorphic and plesiomorphic character states (
<bibRefCitation id="E0424B39CB7BFF98FBD5FE1DFB121D27" author="Borkent, A." box="[1106,1264,511,536]" pageId="3" pageNumber="598" pagination="1 - 456" refId="ref1985" refString="Borkent, A. (2008) The frog-biting midges of the world (Corethrellidae: Diptera). Zootaxa, 1804, 1 - 456. https: // doi. org / 10.11646 / zootaxa. 1804.1.1" type="journal article" year="2008">Borkent, 2008</bibRefCitation>
). Unfortunately, those species are only known as females, thereby hampering further comparisons, given that many of the relevant characters in
<taxonomicName id="43D34D4BCB7BFF98FF67FDAAFE881D60" authorityName="Amaral &amp; Bello-González &amp; Pinho" authorityYear="2021" box="[224,362,583,607]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="stenostyla">
<emphasis id="B6A7EADACB7BFF98FF67FDAAFE881D60" box="[224,362,583,607]" italics="true" pageId="3" pageNumber="598">C. stenostyla</emphasis>
</taxonomicName>
are related to male genitalia. Still,
<taxonomicName id="43D34D4BCB7BFF98FD58FDA5FC291D60" authorityName="Amaral &amp; Bello-González &amp; Pinho" authorityYear="2021" box="[735,971,583,607]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="stenostyla">
<emphasis id="B6A7EADACB7BFF98FD58FDA5FC291D60" box="[735,971,583,607]" italics="true" pageId="3" pageNumber="598">Corethrella stenostyla</emphasis>
</taxonomicName>
differs from those three species by its plain wings with narrow veins and mid and hind femora pale at base. It is nevertheless remarkable that the species with greatest similarity in morphology are found in such distant biogeographical regions, though only a new phylogenetic analysis of the family, including adults and immatures, molecular and other sources of character data will satisfactorily resolve their affinities.
</paragraph>
<paragraph id="846C36C8CB7BFF98FF40FD19FF361C64" blockId="3.[151,1437,151,859]" pageId="3" pageNumber="598">
The other species recorded for
<collectingCountry id="FCC47658CB7BFF98FD9EFD19FDB61C2C" box="[537,596,763,787]" name="Cuba" pageId="3" pageNumber="598">Cuba</collectingCountry>
,
<taxonomicName id="43D34D4BCB7BFF98FDE7FD1EFC981C2B" authority="(Lane, 1939)" baseAuthorityName="Lane" baseAuthorityYear="1939" box="[608,890,763,788]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="B6A7EADACB7BFF98FDE7FD1EFD061C2C" box="[608,740,763,787]" italics="true" pageId="3" pageNumber="598">C. maculata</emphasis>
(
<bibRefCitation id="E0424B39CB7BFF98FD74FD1EFC901C2B" author="Lane, J." box="[755,882,764,788]" pageId="3" pageNumber="598" pagination="99 - 113" refId="ref2100" refString="Lane, J. (1939) Notes on non hematophagous Culicidae. Boletim Biologico, Clube Zoologico do Brasil, 4, 99 - 113." type="journal article" year="1939">Lane, 1939</bibRefCitation>
)
</taxonomicName>
, can be promptly differentiated from
<taxonomicName id="43D34D4BCB7BFF98FA96FD1EFA7F1C2C" authority="Amaral &amp; Bello-González &amp; Pinho, 2021" authorityName="Amaral &amp; Bello-González &amp; Pinho" authorityYear="2021" box="[1297,1437,763,787]" class="Insecta" family="Corethrellidae" genus="Corethrella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="stenostyla" status="sp. nov.">
<emphasis id="B6A7EADACB7BFF98FA96FD1EFA7F1C2C" box="[1297,1437,763,787]" italics="true" pageId="3" pageNumber="598">C. stenostyla</emphasis>
</taxonomicName>
<emphasis id="B6A7EADACB7BFF98FF10FCC2FF301C07" bold="true" box="[151,210,800,824]" pageId="3" pageNumber="598">
<taxonomicNameLabel id="AD9457A1CB7BFF98FF10FCC2FF301C07" box="[151,210,800,824]" pageId="3" pageNumber="598" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by its banded wings, pale halteres, tergites with whitish apical bands and gonocoxite with a dorsal row of five stout setae.
</paragraph>
</subSubSection>
</treatment>
</document>