171 lines
18 KiB
XML
171 lines
18 KiB
XML
<document ID-DOI="10.37520/fi.2021.017" ID-ISSN="2533-4069" ID-Zenodo-Dep="7167657" approvalRequired="6" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="4" checkinTime="1665261192240" checkinUser="felipe" docAuthor="Golovneva, Lina, Bugdaeva, Eugenia, Volynets, Elena, Sun, Yuewu & Zolina, Anastasia" docDate="2021" docId="03CBF058E406FFAAFCF85DA5DA469C71" docLanguage="en" docName="FossilImprint.77.2.231-255.pdf" docOrigin="Fossil Imprint 77 (2)" docSource="http://dx.doi.org/10.37520/fi.2021.017" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.2:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="2" docTitle="Araliaephyllum vittenburgii GOLOVN. et VOLYNETS, Podgorodenka 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="247" masterDocId="FFF28820E409FFBAFFA6595CD8709976" masterDocTitle="Angiosperm Diversification In The Early Cretaceous Of Primorye, Far East Of Russia" masterLastPageNumber="255" masterPageNumber="231" pageNumber="246" updateTime="1665274038775" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Angiosperm Diversification In The Early Cretaceous Of Primorye, Far East Of Russia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Golovneva, Lina</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Bugdaeva, Eugenia</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Volynets, Elena</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Sun, Yuewu</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Zolina, Anastasia</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Fossil Imprint</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2021</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2021-12-24</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>77</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>231</mods:start>
|
||
<mods:end>255</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.37520/fi.2021.017</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.37520/fi.2021.017</mods:identifier>
|
||
<mods:identifier type="ISSN">2533-4069</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7167657</mods:identifier>
|
||
</mods:mods>
|
||
<treatment LSID="urn:lsid:plazi:treatment:03CBF058E406FFAAFCF85DA5DA469C71" httpUri="http://treatment.plazi.org/id/03CBF058E406FFAAFCF85DA5DA469C71" lastPageId="16" lastPageNumber="247" pageId="15" pageNumber="246">
|
||
<subSubSection pageId="15" pageNumber="246" type="nomenclature">
|
||
<paragraph blockId="15.[862,1418,1273,1330]" pageId="15" pageNumber="246">
|
||
<heading centered="true" fontSize="8" level="2" pageId="15" pageNumber="246" reason="2">
|
||
<emphasis bold="true" pageId="15" pageNumber="246">
|
||
<taxonomicName authority="GOLOVN. et VOLYNETS" authorityName="GOLOVN. et VOLYNETS, Podgorodenka" authorityYear="2021" box="[862,1410,1273,1298]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="15" pageNumber="246" phylum="Spermatophyta" rank="species" species="vittenburgii" status="sp. nov.">
|
||
<emphasis bold="true" box="[862,1167,1273,1297]" italics="true" pageId="15" pageNumber="246">Araliaephyllum vittenburgii</emphasis>
|
||
GOLOVN. et VOLYNETS
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[1097,1182,1306,1330]" pageId="15" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="16" lastPageNumber="247" pageId="15" pageNumber="246" type="description">
|
||
<paragraph blockId="15.[1051,1228,1339,1363]" box="[1051,1228,1339,1363]" pageId="15" pageNumber="246">
|
||
<figureCitation box="[1051,1190,1339,1363]" captionStart="Text-fig" captionStartId="8.[147,228,1657,1678]" captionTargetBox="[147,1462,151,1641]" captionTargetId="figure-96@8.[147,1462,151,1641]" captionTargetPageId="8" captionText="Text-fig. 10. a–d: Araliaephyllum luciferum (KRYSHT.) GOLOVN., Partizansk, Severosuchan Formation, Aptian, a – spec. TSNIGRM 3013/1, lectotype, b – spec TSNIGRM 3013/2, c – spec. TSNIGRM 3013/4, d – spec. TSNIGRM 3013/3; e, j: Pandanites ahnertii (KRYSHT.) GOLOVN., Konstantinovka, Lipovtsy Formation, Aptian, TSNIGRM 3013/5, e – fragment of leaf margin, j – general view, j1 – lectotype, j2 – leaf, figured in (e); f: Araliaephyllum ussuriense (KRASSILOV) GOLOVN., Andreev Inlet, Frentsevka Formation, early-middle Albian, spec. IBBS 28/103; g: Araliaephyllum vittenburgii GOLOVN. et VOLYNETS, Podgorodenka coalfield, Galenki Formation, early-middle Albian, spec. IBBS 41/1, g1 – spec. IBBS 41/1a, holotype, g2 – spec. IBBS 41/1b; h: Dicotylophyllum sp. 1, Podgorodenka coalfield, Lipovtsy Formation, Aptian, spec. IBBS 28L-4; i: Sapindopsis sp., 3rd Kamenka, Frentsevka Formation, early-middle Albian, spec. IBSS 325/8. Scale bar 5 mm in a–h and 1 cm in i, j." figureDoi="http://doi.org/10.5281/zenodo.7167677" httpUri="https://zenodo.org/record/7167677/files/figure.png" pageId="15" pageNumber="246">
|
||
Text-fig.
|
||
<quantity box="[1148,1190,1339,1363]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="15" pageNumber="246" unit="g" value="10.0">10g</quantity>
|
||
</figureCitation>
|
||
<figureCitation box="[1189,1228,1339,1363]" captionStart-0="Text-fig" captionStart-1="Text-fig" captionStartId-0="0.[843,924,2048,2069]" captionStartId-1="1.[125,206,2048,2069]" captionTargetBox-0="[843,1478,1612,2032]" captionTargetBox-1="[125,1440,1292,2032]" captionTargetId-0="figure-654@0.[843,1478,1612,2032]" captionTargetId-1="figure-452@1.[125,1440,1292,2032]" captionTargetPageId-0="0" captionTargetPageId-1="1" captionText-0="Text-fig. 1. Map of Primorye region, Russia. a – Razdolnaya coal basin; b – Partizansk coal basin." captionText-1="Text-fig. 2. Map of Partizansk coal basin with early angiosperm localities. a: Severosuchan Formation, Aptian; b: Frentsevka Formation, early-middle Albian. 1 – Novoveselaya village; 2 – 3rd Kamenka River; 3 – Bolshoy Kuvshin; 4 – Andreev Inlet." figureDoi-0="http://doi.org/10.5281/zenodo.7167659" figureDoi-1="http://doi.org/10.5281/zenodo.7167661" httpUri-0="https://zenodo.org/record/7167659/files/figure.png" httpUri-1="https://zenodo.org/record/7167661/files/figure.png" pageId="15" pageNumber="246">1–2</figureCitation>
|
||
</paragraph>
|
||
<paragraph blockId="15.[858,1363,1386,1411]" box="[858,1363,1386,1411]" pageId="15" pageNumber="246">
|
||
H o l o t y p e. No. IBSS 41/1a (
|
||
<figureCitation box="[1195,1341,1386,1410]" captionStart="Text-fig" captionStartId="8.[147,228,1657,1678]" captionTargetBox="[147,1462,151,1641]" captionTargetId="figure-96@8.[147,1462,151,1641]" captionTargetPageId="8" captionText="Text-fig. 10. a–d: Araliaephyllum luciferum (KRYSHT.) GOLOVN., Partizansk, Severosuchan Formation, Aptian, a – spec. TSNIGRM 3013/1, lectotype, b – spec TSNIGRM 3013/2, c – spec. TSNIGRM 3013/4, d – spec. TSNIGRM 3013/3; e, j: Pandanites ahnertii (KRYSHT.) GOLOVN., Konstantinovka, Lipovtsy Formation, Aptian, TSNIGRM 3013/5, e – fragment of leaf margin, j – general view, j1 – lectotype, j2 – leaf, figured in (e); f: Araliaephyllum ussuriense (KRASSILOV) GOLOVN., Andreev Inlet, Frentsevka Formation, early-middle Albian, spec. IBBS 28/103; g: Araliaephyllum vittenburgii GOLOVN. et VOLYNETS, Podgorodenka coalfield, Galenki Formation, early-middle Albian, spec. IBBS 41/1, g1 – spec. IBBS 41/1a, holotype, g2 – spec. IBBS 41/1b; h: Dicotylophyllum sp. 1, Podgorodenka coalfield, Lipovtsy Formation, Aptian, spec. IBBS 28L-4; i: Sapindopsis sp., 3rd Kamenka, Frentsevka Formation, early-middle Albian, spec. IBSS 325/8. Scale bar 5 mm in a–h and 1 cm in i, j." figureDoi="http://doi.org/10.5281/zenodo.7167677" httpUri="https://zenodo.org/record/7167677/files/figure.png" pageId="15" pageNumber="246">
|
||
Text-fig.
|
||
<quantity box="[1294,1341,1386,1410]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="15" pageNumber="246" unit="g" value="10.0">10g</quantity>
|
||
</figureCitation>
|
||
<figureCitation box="[1340,1350,1386,1410]" captionStart="Text-fig" captionStartId="0.[843,924,2048,2069]" captionTargetBox="[843,1478,1612,2032]" captionTargetId="figure-654@0.[843,1478,1612,2032]" captionTargetPageId="0" captionText="Text-fig. 1. Map of Primorye region, Russia. a – Razdolnaya coal basin; b – Partizansk coal basin." figureDoi="http://doi.org/10.5281/zenodo.7167659" httpUri="https://zenodo.org/record/7167659/files/figure.png" pageId="15" pageNumber="246">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="15.[820,1455,1433,1490]" box="[858,1455,1433,1458]" pageId="15" pageNumber="246">P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r:</paragraph>
|
||
<paragraph blockId="15.[820,1455,1433,1490]" box="[820,1147,1466,1490]" pageId="15" pageNumber="246">PFN002696 (for new species).</paragraph>
|
||
<paragraph blockId="15.[820,1459,1513,1634]" pageId="15" pageNumber="246">
|
||
R e p o s i t o r y. Federal Scientific Center of the
|
||
<collectingCountry name="China" pageId="15" pageNumber="246">East Asia</collectingCountry>
|
||
Terrestrial Biodiversity (previously Institute of Botany and Soil Science), Far Eastern Branch, Russian Academy of Sciences, Vladivostok,
|
||
<collectingCountry box="[1070,1145,1610,1634]" name="Russia" pageId="15" pageNumber="246">Russia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="15.[858,1376,1658,1682]" box="[858,1376,1658,1682]" pageId="15" pageNumber="246">E t y m o l o g y. After geologist P. V. Vittenburg.</paragraph>
|
||
<paragraph blockId="15.[820,1459,1705,1794]" pageId="15" pageNumber="246">
|
||
T y p e l o c a l i t y a n d a g e. Dachny Creek, Podgorodenka coalfield, Razdolnaya coal basin, Southern Primorye,
|
||
<collectingCountry box="[933,1008,1770,1794]" name="Russia" pageId="15" pageNumber="246">Russia</collectingCountry>
|
||
; Galenki Formation, early-middle Albian.
|
||
</paragraph>
|
||
<paragraph blockId="15.[820,1459,1817,2100]" pageId="15" pageNumber="246">D i a g n o s i s. Leaves very small, 5–7-lobed, rounded or widely ovate in outline, with cordate base; margin entire; lobes linear-elliptic, with wide rounded apices; central lobe equal in length to upper lateral lobes or slightly shorter; lateral lobes gradually decrease in size towards leaf base; lobe length equal to 1/2 of blade length; sinuses between lobes narrow with rounded apices; basal veins end before lobe apex; lowest basiscopic branches of lateral basal veins strong and innervate 1–2 additional pairs of lateral lobes.</paragraph>
|
||
<paragraph blockId="16.[129,767,151,208]" pageId="16" pageNumber="247">A d d i t i o n a l m a t e r i a l. Specimens IBSS 17/1a and IBSS 17/1b.</paragraph>
|
||
<paragraph blockId="16.[129,768,231,804]" pageId="16" pageNumber="247">
|
||
D e s c r i p t i o n. Leaves are simple, 5–7-lobed, rounded or widely ovate in outline, with cordate base, 1.4–2.0 cm long, 2.0–
|
||
<quantity box="[240,316,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="16" pageNumber="247" unit="cm" value="3.6">3.6 cm</quantity>
|
||
wide. Margin is entire. Lobes are wide, linear-elliptic, with wide rounded apices. The central lobe is equal in length to the upper lateral lobes, or slightly shorter.
|
||
</paragraph>
|
||
<paragraph blockId="16.[129,768,231,804]" pageId="16" pageNumber="247">
|
||
The upper lateral lobes are usually the longest and widest. Lateral lobes gradually decrease in size towards the leaf base. The lobe length is equal to 1/2 of the blade length. Sinuses between lobes are narrow, triangular, with rounded apices. The primary venation is palmate, the secondary venation is brochidodromous. Lateral basal veins diverge from the midvein at an angle of 65–75°, strong, slightly curved, ending before the lobe apex. They bear several curved acroscopic and basiscopic branches, which
|
||
<taxonomicName form="loops" pageId="16" pageNumber="247" rank="form">form loops</taxonomicName>
|
||
near the margin. The lowest basiscopic branches are stronger, and innervate 1–2 additional pairs of lateral lobes. Tertiary venation is very thin, usually irregular. Higher-order venation is not well preserved.
|
||
</paragraph>
|
||
<paragraph blockId="16.[129,768,828,1207]" pageId="16" pageNumber="247">
|
||
C o m p a r i s o n.
|
||
<taxonomicName box="[371,654,828,852]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="luciferum">
|
||
<emphasis box="[371,654,828,852]" italics="true" pageId="16" pageNumber="247">Araliaephyllum luciferum</emphasis>
|
||
</taxonomicName>
|
||
from the Partizansk differs from
|
||
<taxonomicName authorityName="GOLOVN. et VOLYNETS, Podgorodenka" authorityYear="2021" box="[390,546,860,884]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="vittenburgii">
|
||
<emphasis box="[390,546,860,884]" italics="true" pageId="16" pageNumber="247">A. vittenburgii</emphasis>
|
||
</taxonomicName>
|
||
in having a dentate margin and three lobes.
|
||
<taxonomicName box="[390,533,893,916]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="ussuriense">
|
||
<emphasis box="[390,533,893,916]" italics="true" pageId="16" pageNumber="247">A. ussuriense</emphasis>
|
||
</taxonomicName>
|
||
from the Palets Cape also has only three narrow lobes, with the central lobe the largest. Specimens of
|
||
<taxonomicName authorityName="GOLOVN. et VOLYNETS, Podgorodenka" authorityYear="2021" box="[376,533,957,981]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="vittenburgii">
|
||
<emphasis box="[376,533,957,981]" italics="true" pageId="16" pageNumber="247">A. vittenburgii</emphasis>
|
||
</taxonomicName>
|
||
are the most similar to leaves of
|
||
<taxonomicName authority="(G. G. FILIPPOVA) G. G. FILIPPOVA" authorityName="G. G. FILIPPOVA" baseAuthorityName="G. G. FILIPPOVA" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="arenarium">
|
||
<emphasis box="[289,584,989,1013]" italics="true" pageId="16" pageNumber="247">Araliaephyllum arenarium</emphasis>
|
||
(G.G.FILIPPOVA) G.G.FILIPPOVA
|
||
</taxonomicName>
|
||
from the Cenomanian Krivorechenskaya Formation of the Anadyr River (
|
||
<bibRefCitation author="Filippova, G. G. & Abramova, L. N." pageId="16" pageNumber="247" refId="ref13811" refString="Filippova, G. G., Abramova, L. N. (1993): Pozdnemelovaya flora Severo-Vostoka Rossii [The Late Cretaceous Flora of the North-East of Russia]. - Nedra, Moskva [Moscow], 348 pp. (in Russian)" type="book" year="1993">Filippova and Abramova, 1993</bibRefCitation>
|
||
) in number and arrangement of the lobes, but the lobes of
|
||
<taxonomicName box="[155,295,1119,1142]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="arenarium">
|
||
<emphasis box="[155,295,1119,1142]" italics="true" pageId="16" pageNumber="247">A. arenarium</emphasis>
|
||
</taxonomicName>
|
||
are narrower and longer. In addition, the base of
|
||
<taxonomicName box="[158,299,1151,1175]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="arenarium">
|
||
<emphasis box="[158,299,1151,1175]" italics="true" pageId="16" pageNumber="247">A. arenarium</emphasis>
|
||
</taxonomicName>
|
||
is cuneate, while the base of
|
||
<taxonomicName authorityName="GOLOVN. et VOLYNETS, Podgorodenka" authorityYear="2021" box="[614,767,1151,1175]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="vittenburgii">
|
||
<emphasis box="[614,767,1151,1175]" italics="true" pageId="16" pageNumber="247">A. vittenburgii</emphasis>
|
||
</taxonomicName>
|
||
is cordate. The leaves of
|
||
<taxonomicName authorityName="GOLOVN. et VOLYNETS, Podgorodenka" authorityYear="2021" box="[395,547,1183,1207]" class="Magnoliopsida" family="Araliaceae" genus="Araliaephyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="247" phylum="Spermatophyta" rank="species" species="vittenburgii">
|
||
<emphasis box="[395,547,1183,1207]" italics="true" pageId="16" pageNumber="247">A. vittenburgii</emphasis>
|
||
</taxonomicName>
|
||
are much smaller.
|
||
</paragraph>
|
||
<paragraph blockId="16.[129,765,1231,1287]" box="[166,765,1231,1255]" pageId="16" pageNumber="247">S t r a t i g r a p h i c h o r i z o n a n d o c c u r r e n c e.</paragraph>
|
||
<paragraph blockId="16.[129,765,1231,1287]" box="[129,566,1263,1287]" pageId="16" pageNumber="247">Galenki Formation, early-middle Albian.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |