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<document id="0962D36CF757D7CE3760E58CF2A9A5FD" ID-DOI="10.5852/ejt.2024.958.2665" ID-ISSN="2118-9773" ID-Zenodo-Dep="13837028" ID-ZooBank="425BCF68-144B-4861-BCED-CB570D8246D1" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1727252768769" checkinUser="plazi" docAuthor="Constant, Jérôme &amp; Pham, Hong Thai" docDate="2024" docId="03B887FAFFBEFFE92340FB5E5CEF130E" docLanguage="en" docName="EJT.2024.958.114-150.pdf" docOrigin="European Journal of Taxonomy 958" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2665/12349" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zanna bidoupana Constant &amp; Pham, 2024, sp. nov." docType="treatment" docUuid="587A685F-A6C1-465F-A601-1C841D650B7B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="11" masterDocId="FF81FF82FFBAFFE3213DFFAE5E6A1776" masterDocTitle="Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)" masterLastPageNumber="150" masterPageNumber="114" pageNumber="5" updateTime="1727378517280" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5AA34D4B208FDC90ABC3F5952F006639">Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<mods:namePart id="2AB95B7E555B6A84D0B76E01322EABC8">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="8949ECA84F79474D75CA8592D027CC0A">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier Street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
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<mods:namePart id="39097E2483D680698BE1861A6461309C">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="DC9A3364D762344D45FE77CCBC68DB0C">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.</mods:affiliation>
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<mods:date id="EE68554EF2E17C96A72E9A08DCD993B2">2024</mods:date>
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<treatment id="03B887FAFFBEFFE92340FB5E5CEF130E" ID-DOI="http://doi.org/10.5281/zenodo.13837038" ID-Zenodo-Dep="13837038" LSID="urn:lsid:zoobank.org:act:587A685F-A6C1-465F-A601-1C841D650B7B" httpUri="http://treatment.plazi.org/id/03B887FAFFBEFFE92340FB5E5CEF130E" lastPageId="10" lastPageNumber="11" pageId="4" pageNumber="5">
<subSubSection id="C30B6567FFBEFFE72340FB5E5DDD127C" box="[637,951,1263,1290]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BAE36ECFFBEFFE72340FB5E5DDD127C" blockId="4.[368,1219,1263,1361]" box="[637,951,1263,1290]" pageId="4" pageNumber="5">
<heading id="D0E68180FFBEFFE72340FB5E5DDD127C" box="[637,951,1263,1290]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="4C114D6FFFBEFFE72340FB5E5D3F127C" authorityName="Constant &amp; Pham" authorityYear="2024" box="[637,853,1263,1290]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE72340FB5E5D3F127C" bold="true" box="[637,853,1263,1290]" italics="true" pageId="4" pageNumber="5">Zanna bidoupana</emphasis>
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<taxonomicNameLabel id="A2565785FFBEFFE72261FB5E5DDD127C" box="[860,951,1264,1290]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BAE36ECFFBEFFE72345FA985DD61226" blockId="4.[368,1219,1263,1361]" box="[632,956,1334,1361]" pageId="4" pageNumber="5">
<figureCitation id="132A2A69FFBEFFE72345FA985C8A1227" box="[632,736,1334,1361]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="6.[189,230,1928,1954]" captionStartId-1="7.[190,230,1927,1953]" captionStartId-2="8.[189,230,1909,1935]" captionStartId-3="9.[188,229,1994,2020]" captionTargetBox-0="[217,1369,263,1889]" captionTargetBox-1="[216,1346,273,1889]" captionTargetBox-2="[257,1349,265,1872]" captionTargetBox-3="[215,1372,718,1954]" captionTargetId-0="figure-46@6.[212,1369,258,1890]" captionTargetId-1="figure-62@7.[212,1369,260,1892]" captionTargetId-2="figure-45@8.[257,1349,265,1872]" captionTargetId-3="figure-180@9.[215,1372,718,1954]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." captionText-1="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." captionText-2="Fig. 4. AB. Zanna bidoupana sp. nov. at the light trap, Vietnam, Bidoup-Nui Ba National Park, 23 Jul. 2014. A. Dorsal view. B. Lateral view. C. Habitat of Z.bidoupana sp. nov. around Hon Giao station, 23 Jul. 2014." captionText-3="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi-0="http://doi.org/10.5281/zenodo.13837032" figureDoi-1="http://doi.org/10.5281/zenodo.13837034" figureDoi-2="http://doi.org/10.5281/zenodo.13837036" figureDoi-3="http://doi.org/10.5281/zenodo.13837040" httpUri-0="https://zenodo.org/record/13837032/files/figure.png" httpUri-1="https://zenodo.org/record/13837034/files/figure.png" httpUri-2="https://zenodo.org/record/13837036/files/figure.png" httpUri-3="https://zenodo.org/record/13837040/files/figure.png" pageId="4" pageNumber="5">Figs 25</figureCitation>
,
<figureCitation id="132A2A69FFBEFFE723D1FA985D301227" box="[748,858,1334,1361]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="4" pageNumber="5">18A, E, I</figureCitation>
,
<figureCitation id="132A2A69FFBEFFE7225AFA985DD61226" box="[871,956,1334,1360]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="4" pageNumber="5">19AB</figureCitation>
</paragraph>
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<subSubSection id="C30B6567FFBEFFE72180FAD95D2A104E" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BAE36ECFFBEFFE72180FAD95F5C12E7" blockId="4.[189,1406,1399,1849]" box="[189,310,1399,1425]" pageId="4" pageNumber="5">
<heading id="D0E68180FFBEFFE72180FAD95F5C12E7" bold="true" box="[189,310,1399,1425]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B965EAFEFFBEFFE72180FAD95F5C12E7" bold="true" box="[189,310,1399,1425]" pageId="4" pageNumber="5">Diagnosis</emphasis>
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<paragraph id="8BAE36ECFFBEFFE72180FA355D2A104E" blockId="4.[189,1406,1399,1849]" pageId="4" pageNumber="5">
The species is closest to
<taxonomicName id="4C114D6FFFBEFFE720EFFA325C9312C3" authority="Constant" authorityName="Constant" authorityYear="2024" box="[466,761,1435,1461]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE720EFFA325CE012C3" box="[466,650,1435,1461]" italics="true" pageId="4" pageNumber="5">Zanna chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE723C3FA355D3312C3" box="[766,857,1435,1461]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C114D6FFFBEFFE7225CFA325D8F12C3" authorityName="Constant &amp; Pham" authorityYear="2024" box="[865,997,1435,1461]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE7225CFA325D8F12C3" box="[865,997,1435,1461]" italics="true" pageId="4" pageNumber="5">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE722D1FA355A2F12C3" box="[1004,1093,1435,1461]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C114D6FFFBEFFE7254AFA325B1412C3" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1143,1406,1435,1461]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE7254AFA325B6312C3" box="[1143,1289,1435,1461]" italics="true" pageId="4" pageNumber="5">Z.limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE72180FA115F7312AF" box="[189,281,1471,1497]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can be separated by the following characters: basal portion of head (
<figureCitation id="132A2A69FFBEFFE725EAFA105B0512AE" box="[1239,1391,1470,1496]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="4" pageNumber="5">Fig. 2B, EF</figureCitation>
) covered in dense, irregular, rather large, coarse, sometimes coalescent black pitting with few smaller pits (moderately large, rather well-spaced, black pitting in
<taxonomicName id="4C114D6FFFBEFFE7224EF9AB5D931168" authorityName="Constant &amp; Pham" authorityYear="2024" box="[883,1017,1540,1566]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFBEFFE7224EF9AB5D931168" box="[883,1017,1540,1566]" italics="true" pageId="4" pageNumber="5">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE72525F9AA5AA91169" box="[1048,1219,1540,1567]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="4" pageNumber="5">Fig. 12B, EF</figureCitation>
; same but with background of dense, very small black pitting in
<taxonomicName id="4C114D6FFFBEFFE72226F9865DC91134" authorityName="Constant" authorityYear="2024" box="[795,931,1576,1602]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chartieri">
<emphasis id="B965EAFEFFBEFFE72226F9865DC91134" box="[795,931,1576,1602]" italics="true" pageId="4" pageNumber="5">Z. chartieri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFBEFFE722E2F9865A191134" authorityName="Constant" authorityYear="2024" box="[991,1139,1576,1602]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFBEFFE722E2F9865A191134" box="[991,1139,1576,1602]" italics="true" pageId="4" pageNumber="5">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE725ACF9865B5B1134" box="[1169,1329,1575,1602]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="4" pageNumber="5">Figs 6B, EF</figureCitation>
,
<figureCitation id="132A2A69FFBEFFE72403F9895E991113" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="4" pageNumber="5">15B, EF</figureCitation>
); anal tube of male rather high in lateral view (
<figureCitation id="132A2A69FFBEFFE7220CF9E55DFB1113" box="[817,913,1611,1637]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="4" pageNumber="5">Fig. 3A</figureCitation>
), about 1.60 times as long as high, with ventral angle in basal half (rather flattened dorsoventrally, 1.952.14 times as long as high in lateral view, with ventral angle at midlength in
<taxonomicName id="4C114D6FFFBEFFE723A6F93C5D4B11DD" authorityName="Constant" authorityYear="2024" box="[667,801,1681,1707]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chartieri">
<emphasis id="B965EAFEFFBEFFE723A6F93C5D4B11DD" box="[667,801,1681,1707]" italics="true" pageId="4" pageNumber="5">Z. chartieri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFBEFFE72261F93C5D8811DD" authorityName="Constant &amp; Pham" authorityYear="2024" box="[860,994,1681,1707]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFBEFFE72261F93C5D8811DD" box="[860,994,1681,1707]" italics="true" pageId="4" pageNumber="5">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE7253DF93F5A0B11DD" box="[1024,1121,1681,1707]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="4" pageNumber="5">Figs 7A</figureCitation>
,
<figureCitation id="132A2A69FFBEFFE72553F93F5ACE11DD" box="[1134,1188,1681,1707]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="4" pageNumber="5">13A</figureCitation>
); pygofer of male in lateral view (
<figureCitation id="132A2A69FFBEFFE72047F91A5FBD11B9" box="[378,471,1716,1743]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="4" pageNumber="5">Fig. 3A</figureCitation>
) with posterior margin sinuate, forming large, broadly rounded posterior lobe in dorsal half (posterior margin projecting into large posterior lobe angularly rounded in midheight in
<taxonomicName id="4C114D6FFFBEFFE72182F9525F291063" authorityName="Constant &amp; Pham" authorityYear="2024" box="[191,323,1787,1813]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFBEFFE72182F9525F291063" box="[191,323,1787,1813]" italics="true" pageId="4" pageNumber="5">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE72061F9555FAD1063" box="[348,455,1787,1813]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="4" pageNumber="5">Fig. 13A</figureCitation>
; posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle in
<taxonomicName id="4C114D6FFFBEFFE7232AF8B15CC3104E" authorityName="Constant" authorityYear="2024" box="[535,681,1822,1848]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFBEFFE7232AF8B15CC3104E" box="[535,681,1822,1848]" italics="true" pageId="4" pageNumber="5">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE723FBF8B05D58104E" box="[710,818,1822,1848]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="4" pageNumber="5">Fig. 16A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBEFFE72180F8CE5FC010BD" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BAE36ECFFBEFFE72180F8CE5F2E100C" blockId="4.[189,324,1888,1914]" box="[189,324,1888,1914]" pageId="4" pageNumber="5">
<heading id="D0E68180FFBEFFE72180F8CE5F2E100C" bold="true" box="[189,324,1888,1914]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B965EAFEFFBEFFE72180F8CE5F2E100C" bold="true" box="[189,324,1888,1914]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180F8205FC010BD" blockId="4.[189,1398,1934,1995]" pageId="4" pageNumber="5">
The species epithet refers to the
<typeStatus id="54AA884EFFBEFFE72378F8215C1210DF" box="[581,632,1935,1961]" pageId="4" pageNumber="5">type</typeStatus>
location, Bidoup-Nui Ba National Park in
<collectingRegion id="49D5F80EFFBEFFE72540F8215B1810DE" box="[1149,1394,1934,1961]" country="Vietnam" name="Lam Dong" pageId="4" pageNumber="5">Lam Dong Province</collectingRegion>
, Highlands
<collectingCountry id="F306767CFFBEFFE72003F81F5FCF10BD" box="[318,421,1969,1995]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBFFFE62180FEA75F741460" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BAE36ECFFBFFFE62180FEA75F061654" blockId="5.[189,364,264,291]" box="[189,364,264,291]" pageId="5" pageNumber="6">
<heading id="D0E68180FFBFFFE62180FEA75F061654" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B965EAFEFFBFFFE62180FEA75F061654" bold="true" box="[189,364,264,291]" pageId="5" pageNumber="6">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFBFFFE621D7FE995A3916A8" blockId="5.[189,1399,311,478]" pageId="5" pageNumber="6">
<materialsCitation id="3B793CB1FFBFFFE621D7FE995A3916A8" collectingDate="2014-07-21" collectingDateMax="2014-07-25" collectingDateMin="2014-07-21" collectingMethod="Light trap" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="12.186611" location="Hon Giao station" longLatPrecision="1" longitude="108.714806" municipality="Bidoup-Nui Ba National Park" pageId="5" pageNumber="6" specimenCode="32.779" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong Province" typeStatus="holotype">
<emphasis id="B965EAFEFFBFFFE621D7FE995F361627" bold="true" box="[234,348,311,337]" pageId="5" pageNumber="6">
<typeStatus id="54AA884EFFBFFFE621D7FE995F361627" box="[234,348,311,337]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F306767CFFBFFFE62180FEF45F2E1602" box="[189,324,346,372]" name="Vietnam" pageId="5" pageNumber="6">VIETNAM</collectingCountry>
<specimenCount id="9D17FD65FFBFFFE62059FEF45FFC1603" box="[356,406,346,373]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D5F80EFFBFFFE6209BFEF55CF71602" box="[422,669,346,373]" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="6">Lam Dong Province</collectingRegion>
,
<collectingMunicipality id="6BCAAC96FFBFFFE62391FEF45A7D1602" box="[684,1047,346,373]" pageId="5" pageNumber="6">Bidoup-Nui Ba National Park</collectingMunicipality>
,
<location id="8ECE6037FFBFFFE62518FEF55A921602" LSID="urn:lsid:plazi:treatment:03B887FAFFBEFFE92340FB5E5CEF130E:8ECE6037FFBFFFE62518FEF55A921602" box="[1061,1272,346,373]" country="Vietnam" latitude="12.186611" longLatPrecision="1" longitude="108.714806" municipality="Bidoup-Nui Ba National Park" name="Hon Giao station" pageId="5" pageNumber="6" stateProvince="Lam Dong Province">Hon Giao station</location>
;
<geoCoordinate id="EE25502BFFBFFFE6243AFEF75E9F16EE" degrees="12" direction="north" minutes="11" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="11. 8" value="12.186611">12°11ʹ11. 8ʺ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFBFFFE6203DFED25FDE16EE" box="[256,436,380,408]" degrees="108" direction="east" minutes="42" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="53.3" value="108.714806">108°42ʹ53.3ʺ E</geoCoordinate>
;
<date id="FFAF102CFFBFFFE620FDFED35C1716EE" box="[448,637,381,408]" pageId="5" pageNumber="6" value="2014-07-21" valueMax="2014-07-25" valueMin="2014-07-21">
<collectingDate id="EFEBE9C4FFBFFFE620FDFED35C1716EE" box="[448,637,381,408]" pageId="5" pageNumber="6" value="2014-07-21" valueMax="2014-07-25" valueMin="2014-07-21">2125 Jul. 2014</collectingDate>
</date>
;
<collectorName id="26E4533AFFBFFFE623B5FED05D6016EE" box="[648,778,382,408]" pageId="5" pageNumber="6">J. Constant</collectorName>
and
<collectorName id="26E4533AFFBFFFE6227CFED05DD716E1" box="[833,957,381,408]" pageId="5" pageNumber="6">J. Bresseel</collectorName>
leg.; GTI project; I.G.:
<specimenCode id="DBB79E97FFBFFFE625EDFED05B4E16EE" box="[1232,1316,382,408]" pageId="5" pageNumber="6">32.779</specimenCode>
; “Coll. I.R.Sc.N.B., Vietnam, Lam Dong prov., Bidoup-Nui Ba N.P., 12°26ʹ N, 108°30ʹ E, 21-25.VII.2014,
<collectingMethod id="52504EFBFFBFFFE62180FE6A5F5F16A8" box="[189,309,452,478]" pageId="5" pageNumber="6">Light trap</collectingMethod>
, Leg. J. Constant &amp; J. Bresseel, GTI Project, I.G.: 32.779”;
<collectionCode id="ED00AE29FFBFFFE622C8FE6A5A2516A8" box="[1013,1103,452,478]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="5" pageNumber="6" type="Museum">RBINS</collectionCode>
.
</materialsCitation>
</paragraph>
<materialsCitation id="3B793CB1FFBFFFE621D7FDAB5A7D1535" collectingDate="2014-07-21" collectingDateMax="2014-07-25" collectingDateMin="2014-07-21" collectingMethod="Light trap" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="12.186611" location="VIETNAM" longLatPrecision="1" longitude="108.714806" municipality="Bidoup-Nui Ba National Park" pageId="5" pageNumber="6" specimenCount="10" specimenCount-female="2" specimenCount-male="8" stateProvince="Lam Dong" typeStatus="paratype">
<paragraph id="8BAE36ECFFBFFFE621D7FDAB5C4E1569" blockId="5.[189,1399,517,790]" box="[234,548,517,544]" pageId="5" pageNumber="6">
<emphasis id="B965EAFEFFBFFFE621D7FDAB5F0D1569" bold="true" box="[234,359,517,543]" pageId="5" pageNumber="6">
<typeStatus id="54AA884EFFBFFFE621D7FDAB5F0D1569" box="[234,359,517,543]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
</emphasis>
(12 ♂♂, 2 ♀♀)
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FD875F741460" blockId="5.[189,1399,517,790]" pageId="5" pageNumber="6">
<collectingCountry id="F306767CFFBFFFE62180FD875F2E1535" box="[189,324,553,579]" name="Vietnam" pageId="5" pageNumber="6">VIETNAM</collectingCountry>
<specimenCount id="9D17FD65FFBFFFE62058FD875FC01535" box="[357,426,553,579]" count="8" pageId="5" pageNumber="6" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="9D17FD65FFBFFFE6208AFD875F901535" box="[439,506,553,579]" count="2" pageId="5" pageNumber="6" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED00AE29FFBFFFE622FCFD875A7D1535" box="[961,1047,553,579]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="5" pageNumber="6" type="Museum">RBINS</collectionCode>
<materialsCitation id="3B793CB1FFBFFFE6250CFD875C5A15FF" collectingDate="2013-06-04" collectingMethod="light trap" collectionCode="VNMN" collectorName="H. T. Pham" country="Vietnam" location="Bidoup-Nui Ba National Park" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCount id="9D17FD65FFBFFFE6250CFD875A0A1535" box="[1073,1120,553,579]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D5F80EFFBFFFE62550FD875A811535" box="[1133,1259,553,579]" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="6">Lam Dong</collectingRegion>
[Province],
<location id="8ECE6037FFBFFFE62180FDE25C491510" LSID="urn:lsid:plazi:treatment:03B887FAFFBEFFE92340FB5E5CEF130E:8ECE6037FFBFFFE62180FDE25C491510" box="[189,547,588,614]" country="Vietnam" name="Bidoup-Nui Ba National Park" pageId="5" pageNumber="6" stateProvince="Lam Dong">Bidoup-Nui Ba National Park</location>
; 4 Jun. 2013; [
<collectingMethod id="52504EFBFFBFFFE623EEFDE25D221510" box="[723,840,588,614]" pageId="5" pageNumber="6">light trap</collectingMethod>
]; [
<collectorName id="26E4533AFFBFFFE62259FDE25D8C1510" box="[868,998,588,614]" pageId="5" pageNumber="6">H.T. Pham</collectorName>
leg.]; “VQG Bidup-Nui Ba, Lam Dong,
<date id="FFAF102CFFBFFFE6202CFDC15F1615FF" box="[273,380,623,649]" pageId="5" pageNumber="6" value="2013-06-04">
<collectingDate id="EFEBE9C4FFBFFFE6202CFDC15F1615FF" box="[273,380,623,649]" pageId="5" pageNumber="6" value="2013-06-04">4/6/2013</collectingDate>
</date>
, VD”;
<collectionCode id="ED00AE29FFBFFFE620EBFDC15C5A15FF" box="[470,560,623,649]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
</materialsCitation>
<materialsCitation id="3B793CB1FFBFFFE6236CFDC15C1D15DA" collectingDate="2016-10-13" collectionCode="VNMN" collectorName="T. T. Men" location="Bidoup National Park" pageId="5" pageNumber="6" specimenCode="VNMN_E 000009331" stateProvince="Lam Dong Province" typeStatus="paratype">1 ♂; [Lam Dong Province], Bidoup National Park; 13 Oct. 2016; T.T. Men leg.; VNMN_E 000009331</materialsCitation>
; “Bidoup NP, LD, 13/10/2016, light, T.T. Men leg.”, “VNMN_E 000 009 331”; VNMN •
<materialsCitation id="3B793CB1FFBFFFE620D0FD1B5B1D15A6" box="[493,1399,693,720]" collectingDate="2016-10-13" collectionCode="VNMN" collectorName="T. T. Men" country="Vietnam" location="Bidoup National Park" pageId="5" pageNumber="6" specimenCode="VNMN_E 000009330" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCount id="9D17FD65FFBFFFE620D0FD1B5C7515A6" box="[493,543,693,720]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<specimenCode id="DBB79E97FFBFFFE6253FFD185B6515B9" box="[1026,1295,693,720]" pageId="5" pageNumber="6">VNMN_E 000009330</specimenCode>
;
<collectionCode id="ED00AE29FFBFFFE62420FD185B1D15A6" box="[1309,1399,694,720]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
</materialsCitation>
<materialsCitation id="3B793CB1FFBFFFE621F3FD775F741460" collectingDate="2016-10-13" collectingMethod="by net" collectionCode="VNMN" collectorName="T. T. Men &amp; N. V. Dat" country="Vietnam" location="Bidoup National Park" pageId="5" pageNumber="6" specimenCode="VNMN_E 000009332" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCount id="9D17FD65FFBFFFE621F3FD775E971585" box="[206,253,729,755]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectingMethod id="52504EFBFFBFFFE623F4FD765D7E1585" box="[713,788,728,755]" pageId="5" pageNumber="6">by net</collectingMethod>
;
<collectorName id="26E4533AFFBFFFE62222FD775DE01585" box="[799,906,729,755]" pageId="5" pageNumber="6">T.T. Men</collectorName>
and
<collectorName id="26E4533AFFBFFFE622FFFD775A421585" box="[962,1064,729,755]" pageId="5" pageNumber="6">N.V. Dat</collectorName>
leg.;
<specimenCode id="DBB79E97FFBFFFE62554FD775B181585" box="[1129,1394,729,755]" pageId="5" pageNumber="6">VNMN_E 000009332</specimenCode>
;
<collectionCode id="ED00AE29FFBFFFE62180FD525F711460" box="[189,283,764,790]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
.
</materialsCitation>
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C30B6567FFBFFFE92180FC935B191402" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6" type="description">
<paragraph id="8BAE36ECFFBFFFE62180FC935F241421" blockId="5.[189,334,829,855]" box="[189,334,829,855]" pageId="5" pageNumber="6">
<heading id="D0E68180FFBFFFE62180FC935F241421" bold="true" box="[189,334,829,855]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B965EAFEFFBFFFE62180FC935F241421" bold="true" box="[189,334,829,855]" pageId="5" pageNumber="6">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FCC25CAB14BB" blockId="5.[189,1399,876,973]" pageId="5" pageNumber="6">
MEASUREMENTS AND RATIOS. LT:
<specimenCount id="9D17FD65FFBFFFE62369FCC25C0014F0" box="[596,618,876,902]" count="1" pageId="5" pageNumber="6" type="male"></specimenCount>
(n =10): 41.0 mm (38.243.1),
<specimenCount id="9D17FD65FFBFFFE622C7FCC25A7A14F0" box="[1018,1040,876,902]" count="1" pageId="5" pageNumber="6" type="female"></specimenCount>
(n=2): 41.942.0 mm. LTg/ BTg =3.48; LH/BH=4.0; LF/BF =5.3; BH/BPrH = 2.2; LH/LT =0.35; wingspan (extrapolated):
<specimenCount id="9D17FD65FFBFFFE62180FC1D5EBD14BB" box="[189,215,947,973]" count="1" pageId="5" pageNumber="6" type="male"></specimenCount>
:
<quantity id="4CE99B09FFBFFFE621DFFC1C5F2114BB" box="[226,331,946,973]" metricMagnitude="-2" metricUnit="m" metricValue="5.279999999999999" pageId="5" pageNumber="6" unit="mm" value="52.8">52.8 mm</quantity>
(49.255.5),
<specimenCount id="9D17FD65FFBFFFE620D7FC1D5C6814BB" box="[490,514,947,973]" count="1" pageId="5" pageNumber="6" type="female"></specimenCount>
: 54.0
<quantity id="4CE99B09FFBFFFE6236EFC1C5CD714BB" box="[595,701,946,973]" metricMagnitude="-2" metricUnit="m" metricValue="5.41" pageId="5" pageNumber="6" unit="mm" value="54.1">54.1 mm</quantity>
.
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FC5A5F2D125E" blockId="5.[189,1399,1011,1320]" pageId="5" pageNumber="6">
HEAD (
<figureCitation id="132A2A69FFBFFFE62032FC5D5FC71378" box="[271,429,1011,1038]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="5" pageNumber="6">Fig. 2B, EF</figureCitation>
). Strongly elongate, representing 35% of total length, gently and evenly narrowing towards apex; apex obliquely cut in lateral view and with incomplete, rim-shaped orange carina (missing in ventral portion); rather pale pinkish brown entirely covered in dense, irregular black pitting with smaller pits towards distal portion and majority of rather large, sometimes coalescent pits in basal portion. Lateral carinae of frons weakly zigzagged, more prominent towards apex, upcurved near apex and merging into a short dorsal carina reaching apex; lateral carinae of vertex merging with upcurved part of lateral carinae of frons; weakly marked, weakly zigzagged carina in middle portion of vertex, and some cross-carinae between latter and lateral carinae of vertex. Labium pale brown, reaching mesocoxae.
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FAE15FB912D9" blockId="5.[189,1398,1358,1455]" pageId="5" pageNumber="6">
THORAX (
<figureCitation id="132A2A69FFBFFFE6200FFAE05FB3121F" box="[306,473,1358,1385]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="5" pageNumber="6">Fig. 2B, EF</figureCitation>
). Pro- and mesonotum rather pale pinkish brown entirely covered in dense, irregular, strong black pitting with pits often larger than space between them and sometimes coalescent; median carina obsolete.
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FA785C5511D7" blockId="5.[189,1399,1494,1697]" pageId="5" pageNumber="6">
TEGMINA (
<figureCitation id="132A2A69FFBFFFE62006FA785F821286" box="[315,488,1494,1520]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="5" pageNumber="6">Fig. 2A, CD</figureCitation>
). Strongly elongate and reticulate, pale pinkish brown, progressively paler, subhyaline towards apex, basal portion densely covered with minute pink spots; all surface irregularly covered in small to rather large black spots, mostly on veins, larger ones slightly protruding, large middle portion with very dense small spots; costal and postclaval margins subparallel, weakly diverging from base towards apex and curved at level of apex of clavus; apical margin rounded. Veins orange in large basal portion. Clavus open.
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180F9665B221073" blockId="5.[189,1398,1734,1798]" pageId="5" pageNumber="6">VENATION. ScP+R forked in basal ⅓, MP forked more basally in basal 1/4; CuA forked near apex of clavus; PCu and A1 fused near apex of clavus; PCu+A1 fused with postclaval margin at nodal line.</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180F8835D11101C" blockId="5.[189,1399,1836,1898]" pageId="5" pageNumber="6">
HIND WINGS (
<figureCitation id="132A2A69FFBFFFE6205DF8825F881031" box="[352,482,1836,1863]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="5" pageNumber="6">Fig. 2A, C</figureCitation>
). Subhyaline, milky white with veins slightly darker, vein PCu dark brown in basal half; postclaval margin weakly emarginate at A1 vein.
</paragraph>
<caption id="DF6E6664FFBCFFE52180F8265FCC109E" ID-DOI="http://doi.org/10.5281/zenodo.13837032" ID-Zenodo-Dep="13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,230,1928,1954]" targetBox="[217,1369,263,1889]" targetPageId="6">
<paragraph id="8BAE36ECFFBCFFE52180F8265FCC109E" blockId="6.[189,1398,1928,2024]" pageId="6" pageNumber="7">
<emphasis id="B965EAFEFFBCFFE52180F8265F6610D4" bold="true" box="[189,268,1928,1954]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
<taxonomicName id="4C114D6FFFBCFFE52029F8275F8F10D4" authorityName="Constant &amp; Pham" authorityYear="2024" box="[276,485,1928,1954]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFBCFFE52029F8275F8F10D4" box="[276,485,1928,1954]" italics="true" pageId="6" pageNumber="7">Zanna bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBCFFE520D1F8265C2010D4" box="[492,586,1928,1954]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS).
<emphasis id="B965EAFEFFBCFFE52259F8265D1310D4" bold="true" box="[868,889,1928,1954]" pageId="6" pageNumber="7">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B965EAFEFFBCFFE525B4F8265AF710D4" bold="true" box="[1161,1181,1928,1954]" pageId="6" pageNumber="7">B</emphasis>
. Head and thorax, dorsal view.
<emphasis id="B965EAFEFFBCFFE52064F8055F0410B3" bold="true" box="[345,366,1963,1989]" pageId="6" pageNumber="7">C</emphasis>
. Habitus, ventral view.
<emphasis id="B965EAFEFFBCFFE523AEF8055CC210B3" bold="true" box="[659,680,1963,1989]" pageId="6" pageNumber="7">D</emphasis>
. Habitus, lateral view.
<emphasis id="B965EAFEFFBCFFE522F9F8055DB210B3" bold="true" box="[964,984,1963,1989]" pageId="6" pageNumber="7">E</emphasis>
. Head, ventral view.
<emphasis id="B965EAFEFFBCFFE525E2F8055A9B10B3" bold="true" box="[1247,1265,1963,1989]" pageId="6" pageNumber="7">F</emphasis>
. Head and thorax, lateral view.
</paragraph>
</caption>
<caption id="DF6E6664FFBDFFE42183F8295B7E109E" ID-DOI="http://doi.org/10.5281/zenodo.13837034" ID-Zenodo-Dep="13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="7" pageNumber="8" startId="7.[190,230,1927,1953]" targetBox="[216,1346,273,1889]" targetPageId="7">
<paragraph id="8BAE36ECFFBDFFE42183F8295B7E109E" blockId="7.[188,1400,1927,2024]" pageId="7" pageNumber="8">
<emphasis id="B965EAFEFFBDFFE42183F8295F6D10D7" bold="true" box="[190,263,1927,1953]" pageId="7" pageNumber="8">Fig. 3.</emphasis>
<emphasis id="B965EAFEFFBDFFE42030F8295FBD10D7" box="[269,471,1927,1953]" italics="true" pageId="7" pageNumber="8">Zannabidoupana</emphasis>
<taxonomicNameLabel id="A2565785FFBDFFE420E1F8265C5F10D4" box="[476,565,1928,1954]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS), terminalia.
<emphasis id="B965EAFEFFBDFFE42282F8295D9210D7" bold="true" box="[959,1016,1927,1953]" pageId="7" pageNumber="8">AD</emphasis>
. Pygofer, anal tube and gonostyli.
<emphasis id="B965EAFEFFBDFFE42181F8045EBB10B2" bold="true" box="[188,209,1962,1988]" pageId="7" pageNumber="8">A</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFBDFFE42098F8055FD310B3" bold="true" box="[421,441,1963,1989]" pageId="7" pageNumber="8">B</emphasis>
. Caudal view.
<emphasis id="B965EAFEFFBDFFE42362F8045C1E10B2" bold="true" box="[607,628,1962,1988]" pageId="7" pageNumber="8">C</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFBDFFE4222AF8055D4610B3" bold="true" box="[791,812,1963,1989]" pageId="7" pageNumber="8">D</emphasis>
. Ventral view.
<emphasis id="B965EAFEFFBDFFE422EEF8055A6610B3" bold="true" box="[979,1036,1963,1989]" pageId="7" pageNumber="8">EH</emphasis>
. Aedeagus.
<emphasis id="B965EAFEFFBDFFE425AEF8055ACD10B3" bold="true" box="[1171,1191,1963,1989]" pageId="7" pageNumber="8">E</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFBDFFE42180F8605EA5109E" bold="true" box="[189,207,1998,2024]" pageId="7" pageNumber="8">F</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFBDFFE42048F8635FE61091" bold="true" box="[373,396,1997,2023]" pageId="7" pageNumber="8">G</emphasis>
. Laterodorsal view.
<emphasis id="B965EAFEFFBDFFE42348F8605CE6109E" bold="true" box="[629,652,1998,2024]" pageId="7" pageNumber="8">H</emphasis>
. Ventral view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF6E6664FFB2FFEB2180F8DB5FDE10A0" ID-DOI="http://doi.org/10.5281/zenodo.13837036" ID-Zenodo-Dep="13837036" httpUri="https://zenodo.org/record/13837036/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,230,1909,1935]" targetBox="[257,1349,265,1872]" targetPageId="8">
<paragraph id="8BAE36ECFFB2FFEB2180F8DB5FDE10A0" blockId="8.[189,1398,1909,2006]" pageId="8" pageNumber="9">
<emphasis id="B965EAFEFFB2FFEB2180F8DB5F3F10F9" bold="true" box="[189,341,1909,1936]" pageId="8" pageNumber="9">Fig. 4. AB</emphasis>
.
<taxonomicName id="4C114D6FFFB2FFEB2055F8D95C5710E6" authorityName="Constant &amp; Pham" authorityYear="2024" box="[360,573,1910,1936]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFB2FFEB2055F8D95C5710E6" box="[360,573,1910,1936]" italics="true" pageId="8" pageNumber="9">Zanna bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB2FFEB2377F8D85CC010E6" box="[586,682,1910,1936]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
at the light trap, Vietnam, Bidoup-Nui Ba National Park, 23 Jul. 2014.
<emphasis id="B965EAFEFFB2FFEB205FF8375F1D10C5" bold="true" box="[354,375,1945,1971]" pageId="8" pageNumber="9">A</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFB2FFEB2315F8375C5610C5" bold="true" box="[552,572,1945,1971]" pageId="8" pageNumber="9">B</emphasis>
. Lateral view.
<emphasis id="B965EAFEFFB2FFEB23CCF8375D6C10C5" bold="true" box="[753,774,1945,1971]" pageId="8" pageNumber="9">C</emphasis>
. Habitat of
<taxonomicName id="4C114D6FFFB2FFEB22A6F8345A5D10C5" authorityName="Constant &amp; Pham" authorityYear="2024" box="[923,1079,1945,1971]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFB2FFEB22A6F8345A5D10C5" box="[923,1079,1945,1971]" italics="true" pageId="8" pageNumber="9">Z.bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB2FFEB257CF8375AF710C5" box="[1089,1181,1945,1971]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
around Hon Giao station, 23 Jul. 2014.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFB3FFEA2180FEA75B3F161F" blockId="9.[187,1399,265,362]" pageId="9" pageNumber="10">
VENATION. ScP+R and MP forked at midlength; CuA forked slightly more basally then MP fork; PCu forked at basal ⅓; A1 forked close to base, A1
<subScript id="179534A9FFB3FFEA2344FE915CE8163B" attach="left" box="[633,642,319,333]" fontSize="6" pageId="9" pageNumber="10">2</subScript>
fused with A
<quantity id="4CE99B09FFB3FFEA2226FE825D2C1630" box="[795,838,300,326]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="10" unit="in" value="2.0">2 in</quantity>
distal portion and A1
<subScript id="179534A9FFB3FFEA2501FE915A2E163B" attach="none" box="[1084,1092,319,333]" fontSize="6" pageId="9" pageNumber="10">2</subScript>
+A2 forked slightly before reaching postclaval margin. Numerous crossveins in large portion along apical and postclaval margins.
</paragraph>
<paragraph id="8BAE36ECFFB3FFEA2180FE3F5B6F1508" blockId="9.[189,1399,401,639]" pageId="9" pageNumber="10">
LEGS (
<figureCitation id="132A2A69FFB3FFEA203AFE3F5FC316DD" box="[263,425,401,427]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="9" pageNumber="10">Fig. 2A, CD</figureCitation>
). Pale pinkish brown, pilose, rather short and robust. Pro- and mesocoxae with some black markings; all femora with some small black spots and conspicuous ring in distal ⅓, formed from large, more or less coalescent, black spots; pro- and mesotibiae with some small black spots and with apex black; pro- and mesotarsi blackish brown; metatibiae brown basally with all spines apically black, 56 lateral spines and 8 apical spines; first metatarsomere with 11 apical spines ventrally; second metatarsomere with 9 apical spines ventrally; first and second metatarsomere with a dense pad of microsetae ventrally; third metatarsomere elongate with a blackish brown ring. Metatibiotarsal formula: (56) 8/11/9.
</paragraph>
<caption id="DF6E6664FFB3FFEA2181F8645A6D1093" ID-DOI="http://doi.org/10.5281/zenodo.13837040" ID-Zenodo-Dep="13837040" box="[188,1031,1994,2021]" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="9" pageNumber="10" startId="9.[188,229,1994,2020]" targetBox="[215,1372,718,1954]" targetPageId="9">
<paragraph id="8BAE36ECFFB3FFEA2181F8645A6D1093" blockId="9.[188,1031,1994,2021]" box="[188,1031,1994,2021]" pageId="9" pageNumber="10">
<emphasis id="B965EAFEFFB3FFEA2181F8645F601093" bold="true" box="[188,266,1994,2021]" pageId="9" pageNumber="10">Fig. 5.</emphasis>
<taxonomicName id="4C114D6FFFB3FFEA202FF8625F371093" authorityName="Kirkaldy" authorityYear="1902" box="[274,349,1996,2021]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB3FFEA202FF8625F371093" box="[274,349,1996,2021]" italics="true" pageId="9" pageNumber="10">Zanna</emphasis>
</taxonomicName>
, new spp. from Cambodia and Vietnam, distribution map.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFB0FFE92180FEA75C9116FA" blockId="10.[189,1398,265,396]" pageId="10" pageNumber="11">
ABDOMEN (
<figureCitation id="132A2A69FFB0FFE9207AFEA75FBB1655" box="[327,465,265,291]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="10" pageNumber="11">Fig. 2A, C</figureCitation>
). Tergites smooth and shiny, blackish brown bordered with yellowish along posterior margin; sternites pale pinkish brown with transverse irregular row of strong black dots and rounded, brown marking along middle portion of posterior margin, except pregenital sternite with large subtriangular blackish brown marking in middle.
</paragraph>
<paragraph id="8BAE36ECFFB0FFE92180FE1A5B191402" blockId="10.[189,1399,436,885]" pageId="10" pageNumber="11">
MALE TERMINALIA (
<figureCitation id="132A2A69FFB0FFE92099FE1A5F8616B8" box="[420,492,436,462]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
). Pygofer (
<emphasis id="B965EAFEFFB0FFE9234EFE1B5CF816B8" box="[627,658,437,462]" italics="true" pageId="10" pageNumber="11">Py</emphasis>
<figureCitation id="132A2A69FFB0FFE9238CFE1A5D5B16B8" box="[689,817,436,462]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3AD</figureCitation>
) about 1.77 times as high as long in lateral view, robust with posterior margin sinuate, forming large, broadly rounded posterior lobe in dorsal half; sides excavate in dorsal portion (visible in dorsal aspect,
<figureCitation id="132A2A69FFB0FFE9225FFE545DD41563" box="[866,958,506,533]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3C</figureCitation>
). Anal tube (
<emphasis id="B965EAFEFFB0FFE92565FE555A131563" box="[1112,1145,507,533]" italics="true" pageId="10" pageNumber="11">An</emphasis>
<figureCitation id="132A2A69FFB0FFE925AAFE545B7F1563" box="[1175,1301,506,533]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3AC</figureCitation>
) robust, 1.60 times as long as high and with ventral margin oblique in large posterior portion and angularly projecting in lateral view; 1.2 times as long as wide in dorsal view and with sides nearly straight, weakly diverging towards apex; more or less round in caudal view; epiproct and paraproct slightly supassing anal tube, rounded posteriorly in dorsal aspect. Gonostyli (
<emphasis id="B965EAFEFFB0FFE92253FD265DE915D4" box="[878,899,648,674]" italics="true" pageId="10" pageNumber="11">G</emphasis>
<figureCitation id="132A2A69FFB0FFE9229DFD295A4A15D7" box="[928,1056,647,673]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3AD</figureCitation>
) elongate, 2.46 times as long as high; in lateral view, ventral margin broadly rounded and dorsal margin rather broadly rounded after lateral hook (
<emphasis id="B965EAFEFFB0FFE92067FD605F15159E" box="[346,383,718,744]" italics="true" pageId="10" pageNumber="11">lhg</emphasis>
), apical margin rounded; lateral hook with basal portion projecting posterolaterad and apex strongly curved ventrad; in ventral view, lateral protrusion at about midlength, rather strongly and angularly marked. Aedeagus (
<figureCitation id="132A2A69FFB0FFE92318FCBA5CCF1458" box="[549,677,788,814]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3EH</figureCitation>
) rather robust with lateral processes (
<emphasis id="B965EAFEFFB0FFE9254FFCBA5AFD1458" box="[1138,1175,788,814]" italics="true" pageId="10" pageNumber="11">lpa</emphasis>
) forming multiple lobes; dorsal process (
<emphasis id="B965EAFEFFB0FFE920F7FC995F9C1427" box="[458,502,823,849]" italics="true" pageId="10" pageNumber="11">dpa</emphasis>
) moderately protruding dorsally in lateral view; dorsal endosomal process (
<emphasis id="B965EAFEFFB0FFE921FAFCF45E9B1402" box="[199,241,858,884]" italics="true" pageId="10" pageNumber="11">dep</emphasis>
) in dorsal view subtriangular with lateral margins weakly curved and median slit in distal portion.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFB0FFE92180FC325B20137E" pageId="10" pageNumber="11" type="biology_ecology">
<paragraph id="8BAE36ECFFB0FFE92180FC325F7614C0" blockId="10.[189,284,924,950]" box="[189,284,924,950]" pageId="10" pageNumber="11">
<heading id="D0E68180FFB0FFE92180FC325F7614C0" bold="true" box="[189,284,924,950]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B965EAFEFFB0FFE92180FC325F7614C0" bold="true" box="[189,284,924,950]" pageId="10" pageNumber="11">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB0FFE92180FC645B20137E" blockId="10.[189,1399,970,1032]" pageId="10" pageNumber="11">
The specimens were collected at a mercury vapour light trap at Hon Giao station in July (
<figureCitation id="132A2A69FFB0FFE925DAFC645B001492" box="[1255,1386,970,996]" captionStart="Fig" captionStartId="8.[189,230,1909,1935]" captionTargetBox="[257,1349,265,1872]" captionTargetId="figure-45@8.[257,1349,265,1872]" captionTargetPageId="8" captionText="Fig. 4. AB. Zanna bidoupana sp. nov. at the light trap, Vietnam, Bidoup-Nui Ba National Park, 23 Jul. 2014. A. Dorsal view. B. Lateral view. C. Habitat of Z.bidoupana sp. nov. around Hon Giao station, 23 Jul. 2014." figureDoi="http://doi.org/10.5281/zenodo.13837036" httpUri="https://zenodo.org/record/13837036/files/figure.png" pageId="10" pageNumber="11">Fig. 4AB</figureCitation>
). The habitat around was a clearing covered in grass, herbaceaous weeds and small bushes (
<figureCitation id="132A2A69FFB0FFE925DDFC435B57137E" box="[1248,1341,1005,1032]" captionStart="Fig" captionStartId="8.[189,230,1909,1935]" captionTargetBox="[257,1349,265,1872]" captionTargetId="figure-45@8.[257,1349,265,1872]" captionTargetPageId="8" captionText="Fig. 4. AB. Zanna bidoupana sp. nov. at the light trap, Vietnam, Bidoup-Nui Ba National Park, 23 Jul. 2014. A. Dorsal view. B. Lateral view. C. Habitat of Z.bidoupana sp. nov. around Hon Giao station, 23 Jul. 2014." figureDoi="http://doi.org/10.5281/zenodo.13837036" httpUri="https://zenodo.org/record/13837036/files/figure.png" pageId="10" pageNumber="11">Fig. 4C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFB0FFE92180FB815CEF130E" box="[189,645,1071,1144]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BAE36ECFFB0FFE92180FB815F3C133F" blockId="10.[189,342,1071,1097]" box="[189,342,1071,1097]" pageId="10" pageNumber="11">
<heading id="D0E68180FFB0FFE92180FB815F3C133F" bold="true" box="[189,342,1071,1097]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B965EAFEFFB0FFE92180FB815F3C133F" bold="true" box="[189,342,1071,1097]" pageId="10" pageNumber="11">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB0FFE92180FBF35CEF130E" blockId="10.[189,645,1117,1144]" box="[189,645,1117,1144]" pageId="10" pageNumber="11">
<collectingCountry id="F306767CFFB0FFE92180FBF35F4F1301" box="[189,293,1117,1143]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
:
<collectingRegion id="49D5F80EFFB0FFE9200DFBF05C771301" box="[304,541,1117,1144]" country="Vietnam" name="Lam Dong" pageId="10" pageNumber="11">Lam Dong Province</collectingRegion>
(
<figureCitation id="132A2A69FFB0FFE92313FBF35C1C130E" box="[558,630,1117,1144]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>