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<mods:title id="3EF9DEAA7BC82F79EB54DCC92C5B1E17">Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<mods:namePart id="2883064185AEF0DF5F40F4B9B146D5AD">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="A307D676FA16547C4DACD4464F530BB0">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.</mods:affiliation>
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Genus
<taxonomicName id="4C114D6FFFB9FFE02382FB555DAD1263" ID-CoL="6496H" ID-ENA="548218" authority="Kirkaldy, 1902" authorityName="Kirkaldy" authorityYear="1902" box="[703,967,1275,1301]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB9FFE02382FB555D651263" bold="true" box="[703,783,1275,1301]" italics="true" pageId="3" pageNumber="4">Zanna</emphasis>
<bibRefCitation id="EF804B1DFFB9FFE0222AFB555DAD1263" author="Kirkaldy G. W." box="[791,967,1275,1301]" pageId="3" pageNumber="4" pagination="46 - 58" refId="ref13368" refString="Kirkaldy G. W. 1902. Memoirs on Oriental Rhynchota. Journal of the Bombay Natural History Society 14: 46 - 58." type="journal article" year="1902">Kirkaldy, 1902</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C30B6567FFB9FFE02180FA935B631174" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="8BAE36ECFFB9FFE02180FA935A551221" blockId="3.[189,1087,1340,1367]" box="[189,1087,1340,1367]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="AB0111C2FFB9FFE02180FA935A551221" box="[189,1087,1340,1367]" pageId="3" pageNumber="4">
<treatmentCitation id="0AB010FDFFB9FFE02180FA935F851221" author="Kirkaldy G. W." box="[189,495,1340,1367]" page="47" pageId="3" pageNumber="4" year="1902">
<taxonomicName id="4C114D6FFFB9FFE02180FA935F851221" ID-CoL="6496H" authority="Kirkaldy, 1902: 47" authorityName="Kirkaldy" authorityPageNumber="47" authorityYear="1902" box="[189,495,1340,1367]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB9FFE02180FA935F621220" box="[189,264,1341,1366]" italics="true" pageId="3" pageNumber="4">Zanna</emphasis>
<bibRefCitation id="EF804B1DFFB9FFE02032FA925F851221" author="Kirkaldy G. W." box="[271,495,1340,1367]" pageId="3" pageNumber="4" pagination="46 - 58" refId="ref13368" refString="Kirkaldy G. W. 1902. Memoirs on Oriental Rhynchota. Journal of the Bombay Natural History Society 14: 46 - 58." type="journal article" year="1902">Kirkaldy, 1902: 47</bibRefCitation>
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.
<typeStatus id="54AA884EFFB9FFE020C7FA935C5E1221" box="[506,564,1341,1367]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<emphasis id="B965EAFEFFB9FFE0239DFA925D101220" box="[672,890,1340,1366]" italics="true" pageId="3" pageNumber="4">Fulgoratenebrosa</emphasis>
Fabricius, 1775.
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</paragraph>
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<treatmentCitationGroup id="AB0111C2FFB9FFE02180FAD15B1D12EE" box="[189,1399,1406,1433]" pageId="3" pageNumber="4">
<taxonomicName id="4C114D6FFFB9FFE02180FAD15F6212EE" ID-CoL="6496H" authorityName="Kirkaldy" authorityYear="1902" box="[189,264,1407,1432]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB9FFE02180FAD15F6212EE" box="[189,264,1407,1432]" italics="true" pageId="3" pageNumber="4">Zanna</emphasis>
</taxonomicName>
<treatmentCitation id="0AB010FDFFB9FFE02016FAD05C6612EE" author="Melichar L." box="[299,524,1406,1432]" page="13" pageId="3" pageNumber="4" year="1903">
<bibRefCitation id="EF804B1DFFB9FFE02016FAD05C6612EE" author="Melichar L." box="[299,524,1406,1432]" pageId="3" pageNumber="4" refId="ref13563" refString="Melichar L. 1903. Homopteren-Fauna von Ceylon. Verlag von Felix L. Dames, Berlin." type="book" year="1903">Melichar 1903: 13</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AB010FDFFB9FFE0237FFAD05D4E12EE" author="Metcalf Z. P." box="[578,804,1406,1432]" page="246" pageId="3" pageNumber="4" year="1947">
<bibRefCitation id="EF804B1DFFB9FFE0237FFAD05D4E12EE" author="Metcalf Z. P." box="[578,804,1406,1432]" pageId="3" pageNumber="4" refId="ref13618" refString="Metcalf Z. P. 1947. General Catalogue of the Homoptera. Fascicle IV Fulgoroidea. Part 9 Fulgoridae. North Carolina State College, Raleigh (USA)." type="book" year="1947">Metcalf 1947: 246</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AB010FDFFB9FFE02264FAD05A3712EE" author="Lallemand V." box="[857,1117,1406,1433]" page="101" pageId="3" pageNumber="4" year="1959">
<bibRefCitation id="EF804B1DFFB9FFE02264FAD05A3712EE" author="Lallemand V." box="[857,1117,1406,1433]" pageId="3" pageNumber="4" pagination="37 - 123" refId="ref13393" refString="Lallemand V. 1959. Revision des especes africaines de la famille Fulgoridae (super-famille Fulgoroides - sous-ordre des Homopteres). Publicacoes Culturais, Companhia de Diamantes de Angola (Diamang), Lisboa 41: 37 - 123." type="book chapter" year="1959">Lallemand 1959: 101</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB010FDFFB9FFE02551FAD05AB012EE" author="Lallemand V." box="[1132,1242,1406,1432]" page="90" pageId="3" pageNumber="4" year="1963">
<bibRefCitation id="EF804B1DFFB9FFE02551FAD05AB012EE" author="Lallemand V." box="[1132,1242,1406,1432]" pageId="3" pageNumber="4" pagination="1 - 99" refId="ref13434" refString="Lallemand V. 1963. Revision des Fulgoridae (Homoptera). Deuxieme partie. Faunes asiatique et australienne. Memoires de l'Institut royal des Sciences naturelles de Belgique (2 e serie) 75: 1 - 99." type="book chapter" year="1963">1963: 90</bibRefCitation>
</treatmentCitation>
. —
<bibRefCitation id="EF804B1DFFB9FFE02432FAD05B1D12EE" author="Nagai S. &amp; Porion T." box="[1295,1399,1406,1432]" pageId="3" pageNumber="4" refId="ref13649" refString="Nagai S. &amp; Porion T. 1996. Fulgoridae 2. Illustrated Catalogue of the Asiatic and Australian Fauna. Sciences Nat., Compiegne." type="book" year="1996">Nagai &amp;</bibRefCitation>
</treatmentCitationGroup>
Porion 1996: 27. —
<bibRefCitation id="EF804B1DFFB9FFE020E8FA0F5CA612CD" author="Urban J. M. &amp; Cryan J. R." box="[469,716,1441,1467]" pageId="3" pageNumber="4" pagination="471 - 484" refId="ref13902" refString="Urban J. M. &amp; Cryan J. R. 2009. Entomologically famous, evolutionarily unexplored: The first phylogeny of the lanternfly family Fulgoridae (Insecta: Hemiptera: Fulgoroidea). Molecular Phylogenetics and Evolution 50: 471 - 484. https: // doi. org / 10.1016 / j. ympev. 2008.12.004" type="journal article" year="2009">Urban &amp; Cryan 2009</bibRefCitation>
. Replacement name for
<taxonomicName id="4C114D6FFFB9FFE022DCFA0C5B1C12CD" ID-CoL="765X" authority="Amyot &amp; Audinet-Serville, 1843" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[993,1398,1441,1467]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB9FFE022DCFA0C5A5912CD" box="[993,1075,1442,1467]" italics="true" pageId="3" pageNumber="4">Pyrops</emphasis>
<bibRefCitation id="EF804B1DFFB9FFE02504FA0F5B1C12CD" author="Amyot C. &amp; Audinet-Serville J." box="[1081,1398,1441,1467]" pageId="3" pageNumber="4" refId="ref12739" refString="Amyot C. &amp; Audinet-Serville J. 1843. Deuxieme partie. Homopteres. Homoptera Latr. Histoire naturelle des Insectes. Hemipteres. Librairie encyclopedique de Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 8471" type="book" year="1843">Amyot &amp; Audinet-Serville,</bibRefCitation>
</taxonomicName>
1843 nec
<taxonomicName id="4C114D6FFFB9FFE02061FA6B5C3612A8" authority="Spinola, 1839" authorityName="Spinola" authorityYear="1839" box="[348,604,1476,1502]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB9FFE02061FA6B5FC412A8" box="[348,430,1477,1502]" italics="true" pageId="3" pageNumber="4">Pyrops</emphasis>
Spinola, 1839
</taxonomicName>
.
</paragraph>
<paragraph id="8BAE36ECFFB9FFE02180FA465B631174" blockId="3.[189,1399,1406,1538]" box="[189,1289,1511,1538]" pageId="3" pageNumber="4">
<taxonomicName id="4C114D6FFFB9FFE02180FA465F651177" authorityName="Spinola" authorityYear="1839" box="[189,271,1512,1537]" class="Insecta" family="Fulgoridae" genus="Pyrops" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB9FFE02180FA465F651177" box="[189,271,1512,1537]" italics="true" pageId="3" pageNumber="4">Pyrops</emphasis>
</taxonomicName>
<bibRefCitation id="EF804B1DFFB9FFE02016FA465C8F1174" author="Amyot C. &amp; Audinet-Serville J." box="[299,741,1511,1538]" pageId="3" pageNumber="4" refId="ref12739" refString="Amyot C. &amp; Audinet-Serville J. 1843. Deuxieme partie. Homopteres. Homoptera Latr. Histoire naturelle des Insectes. Hemipteres. Librairie encyclopedique de Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 8471" type="book" year="1843">Amyot &amp; Audinet-Serville 1843: 491</bibRefCitation>
. —
<bibRefCitation id="EF804B1DFFB9FFE0222EFA495D951174" author="Atkinson E. T." box="[787,1023,1511,1538]" pageId="3" pageNumber="4" pagination="127 - 158" refId="ref12784" refString="Atkinson E. T. 1885. Notes on Indian Rhynchota. No. 4. Journal and Proceedings of the Asiatic Society of Bengal. Calcutta 54: 127 - 158." type="journal article" year="1885">Atkinson 1885: 139</bibRefCitation>
. —
<bibRefCitation id="EF804B1DFFB9FFE02512FA495B6E1174" author="Distant W. L." box="[1071,1284,1511,1538]" pageId="3" pageNumber="4" refId="ref13146" refString="Distant W. L. 1906. Rhynchota Vol. 3. Heteroptera-Homoptera. In: Bingham C. T. (ed.) The Fauna of British India, including Ceylon and Burma. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 48423" type="book" year="1906">Distant 1906: 179</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFB9FFE72180F9875A191581" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BAE36ECFFB9FFE02180F9875F5C1135" blockId="3.[189,310,1577,1603]" box="[189,310,1577,1603]" pageId="3" pageNumber="4">
<heading id="D0E68180FFB9FFE02180F9875F5C1135" bold="true" box="[189,310,1577,1603]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B965EAFEFFB9FFE02180F9875F5C1135" bold="true" box="[189,310,1577,1603]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB9FFE02180F9F95F8B11E3" blockId="3.[189,1398,1623,1685]" pageId="3" pageNumber="4">
Species of the genus
<taxonomicName id="4C114D6FFFB9FFE0208EF9F65F941107" authorityName="Kirkaldy" authorityYear="1902" box="[435,510,1624,1649]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB9FFE0208EF9F65F941107" box="[435,510,1624,1649]" italics="true" pageId="3" pageNumber="4">Zanna</emphasis>
</taxonomicName>
can be separated from the other genera of
<taxonomicName id="4C114D6FFFB9FFE022D2F9F95A041107" authorityName="Latreille" authorityYear="1807" box="[1007,1134,1623,1649]" class="Insecta" family="Fulgoridae" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Fulgoridae</taxonomicName>
by the combination of the following characters:
</paragraph>
<paragraph id="8BAE36ECFFB9FFE02180F9125A2F118F" blockId="3.[189,1398,1724,1785]" pageId="3" pageNumber="4">(1) General shape of body distinctly narrow, slender and elongate, usually pale brownish ochraceous, without colorful markings, usually densely and prominently spotted in black;</paragraph>
<paragraph id="8BAE36ECFFB9FFE02180F88F5D991028" blockId="3.[189,1398,1825,1886]" pageId="3" pageNumber="4">(2) Head distinctly produced into an elongate, often very long cephalic process, which can vary from as long as, to more than twice as long as pro- and mesonotum combined;</paragraph>
<paragraph id="8BAE36ECFFB9FFE02180F8285CC21090" blockId="3.[189,1398,1925,2022]" pageId="3" pageNumber="4">(3) Cephalic process relatively robust, projecting straight anteriorly, more or less hexagonal in crosssection, narrowing from base to apex, somewhat upturned at apex in lateral view (some Afrotropical species with apex clavate in lateral view);</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FEA75FCD1630" blockId="4.[189,1398,265,326]" pageId="4" pageNumber="5">(4) Vertex very elongate, lateral margins weakly ridged with carinae being slightly zigzagged, often with faint median carina;</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FEC35C1A16B8" blockId="4.[189,1406,365,463]" pageId="4" pageNumber="5">(5) Frons distinctly elongate, with lateral margins weakly ridged with carinae being slightly zigzagged, with slightly zigzagged submedian carinae, carinae sometimes obsolete; median carina either absent or incomplete and very weakly marked;</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FE5B5F0E1545" blockId="4.[189,1398,501,563]" pageId="4" pageNumber="5">(6) Antennae with pedicel relatively short, subbulbose, fully covered with sensory plaque organs over entire surface;</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FDF45B361503" blockId="4.[189,1372,602,629]" box="[189,1372,602,629]" pageId="4" pageNumber="5">(7) Tegmina narrow and elongate, almost entirely reticulate, inner marginal areas overlapping at rest;</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FD325CE815C3" blockId="4.[189,642,667,694]" box="[189,642,667,694]" pageId="4" pageNumber="5">(8) Hind tibiae with 57 lateral spines;</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FD735A191581" blockId="4.[189,1139,733,759]" box="[189,1139,733,759]" pageId="4" pageNumber="5">(9) Gonoplacs of adult females with basal inner area strongly and densely pilose.</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBEFFE72180FCB05F1114D8" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BAE36ECFFBEFFE72180FCB05E92144E" blockId="4.[189,248,798,824]" box="[189,248,798,824]" pageId="4" pageNumber="5">
<heading id="D0E68180FFBEFFE72180FCB05E92144E" bold="true" box="[189,248,798,824]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B965EAFEFFBEFFE72180FCB05E92144E" bold="true" box="[189,248,798,824]" pageId="4" pageNumber="5">Note</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FCE35F1114D8" blockId="4.[189,1401,845,942]" pageId="4" pageNumber="5">
The above diagnosis was modified from
<bibRefCitation id="EF804B1DFFBEFFE7239BFCE35D281411" author="Liang A. - P." box="[678,834,845,871]" pageId="4" pageNumber="5" pagination="361 - 373" refId="ref13506" refString="Liang A. - P. 2017. Zanna robusticephalica sp. nov. (Hemiptera: Fulgoromorpha: Fulgoridae) from China, with comments on some ultrastructural characters of the new species. Zootaxa 4338 (2): 361 - 373. https: // doi. org / 10.11646 / zootaxa. 4338.2.10" type="journal article" year="2017">Liang (2017)</bibRefCitation>
as it was necessary to expand the definition of the genus in order to encompass many Afrotropical taxa, including the
<typeStatus id="54AA884EFFBEFFE72528FCDF5A2214FD" box="[1045,1096,881,907]" pageId="4" pageNumber="5">type</typeStatus>
species
<taxonomicName id="4C114D6FFFBEFFE7258FFCDF5F1D14D8" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tenebrosa">
<emphasis id="B965EAFEFFBEFFE7258FFCDF5B1314FC" box="[1202,1401,880,906]" italics="true" pageId="4" pageNumber="5">Zanna tenebrosa</emphasis>
Fabricius, 1775
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBEFFE72180FC7A5FB213F1" pageId="4" pageNumber="5" type="taxon_list">
<paragraph id="8BAE36ECFFBEFFE72180FC7A5DA11498" blockId="4.[189,971,980,1007]" box="[189,971,980,1007]" pageId="4" pageNumber="5">
<emphasis id="B965EAFEFFBEFFE72180FC7A5DA11498" bold="true" box="[189,971,980,1007]" pageId="4" pageNumber="5">
Checklist of the species of
<taxonomicName id="4C114D6FFFBEFFE72338FC7B5C3F1499" authorityName="Kirkaldy" authorityYear="1902" box="[517,597,981,1007]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFBEFFE72338FC7B5C3F1499" bold="true" box="[517,597,981,1007]" italics="true" pageId="4" pageNumber="5">Zanna</emphasis>
</taxonomicName>
from
<collectingCountry id="F306767CFFBEFFE7239CFC7A5D491498" box="[673,803,980,1006]" name="Cambodia" pageId="4" pageNumber="5">Cambodia</collectingCountry>
and
<collectingCountry id="F306767CFFBEFFE7225CFC7A5DA11498" box="[865,971,980,1006]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FBAA5F851368" blockId="4.[189,596,1027,1195]" box="[189,495,1027,1054]" pageId="4" pageNumber="5">
<taxonomicName id="4C114D6FFFBEFFE72180FBAA5FE6136B" authorityName="Constant &amp; Pham" authorityYear="2024" box="[189,396,1027,1053]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE72180FBAA5FE6136B" box="[189,396,1027,1053]" italics="true" pageId="4" pageNumber="5">Zanna bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE720A9FBAA5F851368" box="[404,495,1028,1054]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FB895C221337" blockId="4.[189,596,1027,1195]" box="[189,584,1062,1089]" pageId="4" pageNumber="5">
<taxonomicName id="4C114D6FFFBEFFE72180FB895F8F1337" authority="Constant" authorityName="Constant" authorityYear="2024" box="[189,485,1062,1089]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE72180FB895F1C1336" box="[189,374,1062,1088]" italics="true" pageId="4" pageNumber="5">Zanna chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE720D0FB895C221337" box="[493,584,1063,1089]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FBE45C5F1312" blockId="4.[189,596,1027,1195]" box="[189,565,1098,1124]" pageId="4" pageNumber="5">
<taxonomicName id="4C114D6FFFBEFFE72180FBE45C5F1312" authority="(Distant, 1893)" baseAuthorityName="Distant" baseAuthorityYear="1893" box="[189,565,1098,1124]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B965EAFEFFBEFFE72180FBE45F161312" box="[189,380,1098,1124]" italics="true" pageId="4" pageNumber="5">Zanna chinensis</emphasis>
(
<bibRefCitation id="EF804B1DFFBEFFE720B1FBE45C471312" author="Distant W. L." box="[396,557,1098,1124]" pageId="4" pageNumber="5" pagination="443 - 449" refId="ref13113" refString="Distant W. L. 1893. On the Homopterous genus Pyrops, with descriptions of two new species. Transactions of the Entomological Society of London 1893: 443 - 449." type="journal article" year="1893">Distant, 1893</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FBC05FB213F1" blockId="4.[189,596,1027,1195]" box="[189,472,1133,1159]" pageId="4" pageNumber="5">
<taxonomicName id="4C114D6FFFBEFFE72180FBC05F1C13F1" authorityName="Constant &amp; Pham" authorityYear="2024" box="[189,374,1133,1159]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE72180FBC05F1C13F1" box="[189,374,1133,1159]" italics="true" pageId="4" pageNumber="5">Zanna kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE72040FBC35FB213F1" box="[381,472,1133,1159]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBEFFE72180FB3E5C3E13DD" box="[189,596,1168,1195]" pageId="4" pageNumber="5" type="type_taxon">
<paragraph id="8BAE36ECFFBEFFE72180FB3E5C3E13DD" blockId="4.[189,596,1027,1195]" box="[189,596,1168,1195]" pageId="4" pageNumber="5">
<taxonomicName id="4C114D6FFFBEFFE72180FB3E5F9B13DC" authority="Constant" authorityName="Constant" authorityYear="2024" box="[189,497,1168,1194]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE72180FB3E5FE813DC" box="[189,386,1168,1194]" italics="true" pageId="4" pageNumber="5">Zannalimbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE720C4FB3F5C3E13DD" box="[505,596,1169,1195]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA5FFFC2180FA1A5A0812A7" box="[189,1122,1460,1489]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BAE36ECFFA5FFFC2180FA1A5A0812A7" blockId="31.[189,1122,1460,1489]" box="[189,1122,1460,1489]" pageId="31" pageNumber="32">
<emphasis id="B965EAFEFFA5FFFC2180FA1A5A0812A7" bold="true" box="[189,1122,1460,1489]" italics="true" pageId="31" pageNumber="32">
Identification key to the species of
<taxonomicName id="4C114D6FFFA5FFFC23B9FA1B5CB112A7" ID-CoL="6496H" ID-ENA="548218" authority="Kirkaldy, 1902" authorityName="Kirkaldy" authorityYear="1902" box="[644,731,1461,1489]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Zanna</taxonomicName>
from
<collectingCountry id="F306767CFFA5FFFC2216FA1A5DD812A7" box="[811,946,1460,1489]" name="Cambodia" pageId="31" pageNumber="32">Cambodia</collectingCountry>
and
<collectingCountry id="F306767CFFA5FFFC22C9FA1B5A0812A7" box="[1012,1122,1461,1489]" name="Vietnam" pageId="31" pageNumber="32">Vietnam</collectingCountry>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA5FFC32180FA475B1D1479" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="key">
<key id="FF8061DDFFA5FFC32180FA475B1D1479" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<keyStep id="30E52E49FFA5FFFC2180FA475B1C111B" pageId="31" pageNumber="32">
<paragraph id="8BAE36ECFFA5FFFC2180FA475B1D1150" blockId="31.[189,1399,1513,1645]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180FA475B1D1150" pageId="31" pageNumber="32">
1. Smaller, wingspan up to
<quantity id="4CE99B09FFA5FFFC232DFA475C0D1172" box="[528,615,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="32" unit="mm" value="55.0">55 mm</quantity>
, usually less than
<quantity id="4CE99B09FFA5FFFC2203FA475DFE1172" box="[830,916,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="mm" value="50.0">50 mm</quantity>
; cephalic process exceeding 1.3 times as long as abdomen ................................................................................................................................ 2
</keyLead>
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180F99E5B1C111B" blockId="31.[189,1399,1513,1645]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180F99E5B1C111B" pageId="31" pageNumber="32">
Larger, wingspan
<quantity id="4CE99B09FFA5FFFC20FDF9815C29113C" box="[448,579,1583,1610]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="31" pageNumber="32" unit="mm" value="62.5" valueMax="65.0" valueMin="60.0">6065 mm</quantity>
; cephalic process less than 1.3 times as long as abdomen (
<figureCitation id="132A2A69FFA5FFFC25C0F9815E94111B" captionStart="Fig" captionStartId="32.[188,229,1947,1973]" captionTargetBox="[189,1398,864,1901]" captionTargetId="figure-207@32.[186,1401,864,1901]" captionTargetPageId="32" captionText="Fig. 20. Zanna chinensis (Distant, 1893), syntype, ♀ (BMNH). A. Dorsal view. B. Lateral view. C. Labels (photograph by D. Rendon-Mera, © The Trustees of the Natural History Museum, London)." figureDoi="http://doi.org/10.5281/zenodo.13837084" httpUri="https://zenodo.org/record/13837084/files/figure.png" pageId="31" pageNumber="32">Fig. 20A B</figureCitation>
) ............................................................................................................
<taxonomicName id="4C114D6FFFA5FFFC2516F9FD5B1C111B" ID-CoL="5CW46" authority="(Distant, 1893)" baseAuthorityName="Distant" baseAuthorityYear="1893" box="[1067,1398,1618,1645]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B965EAFEFFA5FFFC2516F9FD5AD7111B" bold="true" box="[1067,1213,1618,1645]" italics="true" pageId="31" pageNumber="32">Z. chinensis</emphasis>
(
<bibRefCitation id="EF804B1DFFA5FFFC25F0F9FD5B04111B" author="Distant W. L." box="[1229,1390,1619,1645]" pageId="31" pageNumber="32" pagination="443 - 449" refId="ref13113" refString="Distant W. L. 1893. On the Homopterous genus Pyrops, with descriptions of two new species. Transactions of the Entomological Society of London 1893: 443 - 449." type="journal article" year="1893">Distant, 1893</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E49FFA5FFFC2180F93A5B1D106E" pageId="31" pageNumber="32">
<paragraph id="8BAE36ECFFA5FFFC2180F93A5B1D11A7" blockId="31.[189,1399,1684,1816]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180F93A5B1D11A7" pageId="31" pageNumber="32">
2. Anal tube of male rather flattened dorsoventrally, 1.952.14 times as long as high in lateral view; ventral angle of anal tube at midlength (
<figureCitation id="132A2A69FFA5FFFC2382F9195D2311A7" box="[703,841,1719,1745]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="31" pageNumber="32">Fig. 19C, E</figureCitation>
) ....................................................................... 3
</keyLead>
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180F9755B1D106E" blockId="31.[189,1399,1684,1816]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180F9755B1D106E" pageId="31" pageNumber="32">
Anal tube of male rather high, 1.601.65 times as long as high in lateral view; ventral angle of anal tube before midlength (
<figureCitation id="132A2A69FFA5FFFC20C0F9505CE1106E" box="[509,651,1790,1816]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="31" pageNumber="32">Fig. 19A, G</figureCitation>
) ................................................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E49FFA5FFC32180F8915B1D16FB" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<paragraph id="8BAE36ECFFA5FFFC2180F8915B1D1091" blockId="31.[189,1399,1855,2023]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180F8915B1D1091" pageId="31" pageNumber="32">
3. Pygofer with posterior margin projecting into a large rounded posterior lobe in dorsal half (
<figureCitation id="132A2A69FFA5FFFC243BF8915B04102F" box="[1286,1390,1855,1881]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="31" pageNumber="32">Fig. 19C</figureCitation>
) and with sides rather strongly excavate in dorsal portion (
<figureCitation id="132A2A69FFA5FFFC22A9F8CC5A68100B" box="[916,1026,1890,1917]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="31" pageNumber="32">Fig. 19D</figureCitation>
); basal portion of head covered in dense, irregular black pitting with smaller pits and some dispersed, moderately large, not or very rarely coalescent pits (
<figureCitation id="132A2A69FFA5FFFC20CFF8075C1610B5" box="[498,636,1961,1987]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="31" pageNumber="32">Fig. 18B, F</figureCitation>
); tegmina more elongate, 3.80 times as long as wide (
<figureCitation id="132A2A69FFA5FFFC25CEF8075B3E10B5" box="[1267,1364,1961,1987]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="31" pageNumber="32">Fig. 18J</figureCitation>
) .. ..............................................................................................................
<taxonomicName id="4C114D6FFFA5FFFC2521F8625B7F1090" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1052,1301,1995,2022]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFA5FFFC2521F8625ACF1090" bold="true" box="[1052,1189,1995,2022]" italics="true" pageId="31" pageNumber="32">Z. chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA5FFFC2421F8635B1D1091" box="[1308,1399,1997,2023]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAE36ECFF9AFFC32180FEA75B1D16FB" blockId="32.[189,1399,265,397]" pageId="32" pageNumber="33">
<keyLead id="30E095D9FF9AFFC32180FEA75B1D16FB" pageId="32" pageNumber="33">
Pygofer with posterior margin projecting into a large posterior lobe angularly rounded in midheight (
<figureCitation id="132A2A69FF9AFFC321CEFE825F311630" box="[243,347,300,326]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19E</figureCitation>
) and with sides weakly excavate in dorsal portion (
<figureCitation id="132A2A69FF9AFFC322FDFE825A431630" box="[960,1065,300,326]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19F</figureCitation>
); basal portion of head with irregular, moderately large, rather well-spaced, black pitting but without background of smaller pits (
<figureCitation id="132A2A69FF9AFFC321CEFEDC5FEA16FB" box="[243,384,370,397]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="32" pageNumber="33">Fig. 18C, G</figureCitation>
); tegmina less elongate, 3.25 times as long as wide (
<figureCitation id="132A2A69FF9AFFC322DCFEDC5A2016FB" box="[993,1098,370,397]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="32" pageNumber="33">Fig. 18K</figureCitation>
).......
<taxonomicName id="4C114D6FFF9AFFC325B6FEDD5B7F16FB" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1163,1301,370,397]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFF9AFFC325B6FEDD5B7F16FB" bold="true" box="[1163,1301,370,397]" italics="true" pageId="32" pageNumber="33">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FF9AFFC32420FEDD5B1D16FB" box="[1309,1399,371,397]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E49FF9AFFC32180FE1A5B1D1479" pageId="32" pageNumber="33">
<paragraph id="8BAE36ECFF9AFFC32180FE1A5B1D152D" blockId="32.[189,1399,436,783]" pageId="32" pageNumber="33">
<keyLead id="30E095D9FF9AFFC32180FE1A5B1D152D" pageId="32" pageNumber="33">
4. Pygofer in lateral view with posterior margin sinuate, forming a large, broadly rounded posterior lobe in dorsal half (
<figureCitation id="132A2A69FF9AFFC320F5FE795C591687" box="[456,563,471,497]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19A</figureCitation>
); anal tube in dorsal view 1.2 times as long as wide and with sides nearly straight, weakly diverging towards apex (
<figureCitation id="132A2A69FF9AFFC323FAFE545D5B1563" box="[711,817,506,533]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19B</figureCitation>
); basal portion of head covered in dense, irregular, rather large, coarse, sometimes coalescent black pitting with few smaller pits (
<figureCitation id="132A2A69FF9AFFC325BDFDB05B66154E" box="[1152,1292,542,568]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="32" pageNumber="33">Fig. 18A, E</figureCitation>
) ............ ..........................................................................................................................
<taxonomicName id="4C114D6FFF9AFFC32549FDEF5B7F152D" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1140,1301,576,603]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFF9AFFC32549FDEF5B7F152D" bold="true" box="[1140,1301,576,603]" italics="true" pageId="32" pageNumber="33">Z. bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FF9AFFC32421FDEF5B1D152D" box="[1308,1399,577,603]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAE36ECFF9AFFC32180FDCB5B1D1479" blockId="32.[189,1399,436,783]" pageId="32" pageNumber="33">
<keyLead id="30E095D9FF9AFFC32180FDCB5B1D1479" pageId="32" pageNumber="33">
Pygofer in lateral view with posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle (
<figureCitation id="132A2A69FF9AFFC323DFFD295D3815D7" box="[738,850,647,673]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19G</figureCitation>
); anal tube in dorsal view 1.08 times as long as wide, with sides roundly diverging towards apex (
<figureCitation id="132A2A69FF9AFFC32240FD045D9A15B3" box="[893,1008,682,709]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19H</figureCitation>
); basal portion of head densely covered in irregular, very small black pitting, with some moderately larger pits, rarely coalescent (
<figureCitation id="132A2A69FF9AFFC321CEFD5A5FE81478" box="[243,386,756,782]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="32" pageNumber="33">Fig. 18D, H</figureCitation>
).......................................................................................
<taxonomicName id="4C114D6FFF9AFFC32533FD5A5B7F1478" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1038,1301,755,782]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFF9AFFC32533FD5A5ACF1478" bold="true" box="[1038,1189,755,782]" italics="true" pageId="32" pageNumber="33">Z. limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FF9AFFC32421FD5B5B1D1479" box="[1308,1399,757,783]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>