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<document id="545748AEBED06A3434142CEA131DD14A" ID-DOI="10.11646/zootaxa.5424.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10808050" ID-ZooBank="3DF7F8AA-BA4C-4C78-BB1E-450537B79CB6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1710236571554" checkinUser="plazi" docAuthor="Mendes, Samuel Lucas Da Silva Delgado, Paiva, Paulo Cesar De &amp; Rizzo, Alexandra E." docDate="2024" docId="03B887F73E096F7AF884B22E00D4FBFE" docLanguage="en" docName="zootaxa.5424.1.4.pdf" docOrigin="Zootaxa 5424 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oligobregma bakkeni Mendes &amp; Paiva &amp; Rizzo 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="94" masterDocId="FF81FF8F3E056F74F813B770034EFFCB" masterDocTitle="First record of Oligobregma Kudenov &amp; Blake, 1975 (Annelida: Polychaeta: Scalibregmatidae Malmgren, 1867) from Brazil with the description of three new species" masterLastPageNumber="98" masterPageNumber="80" pageNumber="92" updateTime="1710356895884" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="EB28D2DB40B8C2EB0E63791766FE9869">
<mods:title id="6E8AAC0D35F4B332CFF46079F00E495E">First record of Oligobregma Kudenov &amp; Blake, 1975 (Annelida: Polychaeta: Scalibregmatidae Malmgren, 1867) from Brazil with the description of three new species</mods:title>
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<mods:name id="E7F1BABFBDF565FD0DDE715D1A10F29A" type="personal">
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<mods:namePart id="694F92064A9BDAAA3A24DA9727000004">Mendes, Samuel Lucas Da Silva Delgado</mods:namePart>
<mods:nameIdentifier id="B1F37E9593A58F1A9A0206DAE3C5C4A6" type="ORCID">0000-0003-4894-3270</mods:nameIdentifier>
<mods:affiliation id="9B9E73A86A24CBA7EBED027750E9EEB7">slucasmendes 013 @ gmail. com</mods:affiliation>
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<mods:namePart id="782C61E69055BAA3F53B3648F2676128">Paiva, Paulo Cesar De</mods:namePart>
<mods:nameIdentifier id="3067585E510E369417DB8A87E008492C" type="ORCID">0000-0003-1061-6549</mods:nameIdentifier>
<mods:affiliation id="1FD351A4C34A9DF81675B283A93306B9">paulopaiva @ gmail. com</mods:affiliation>
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<mods:namePart id="254393B5C23D8697B130E402670B2FE0">Rizzo, Alexandra E.</mods:namePart>
<mods:nameIdentifier id="B89050385B3BFB918D3F5F55C93542EF" type="ORCID">0000-0001-5056-5358</mods:nameIdentifier>
<mods:affiliation id="29E7A07A47212CF0B5BF1A8EEF6507A7">Universidade Federal do Rio de Janeiro, Programa de Pós-Graduação em Zoologia, Museu Nacional.</mods:affiliation>
<mods:nameIdentifier id="FFFE17CB82518A05E21FD1927F47A68F" type="email">aerizzo@hotmail.com</mods:nameIdentifier>
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<mods:title id="C4557DFE888F21F8F31A4FD78AEFF6DB">Zootaxa</mods:title>
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<mods:date id="F7A94393D57BFB090282D5C72B04724D">2024</mods:date>
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<mods:number id="118114FDABE5D4D03CE2AB9EFEC798C3">2024-03-12</mods:number>
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<mods:number id="0F0F8F190F9C0A5B7353A3279765A75C">5424</mods:number>
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<treatment id="03B887F73E096F7AF884B22E00D4FBFE" LSID="urn:lsid:plazi:treatment:03B887F73E096F7AF884B22E00D4FBFE" httpUri="http://treatment.plazi.org/id/03B887F73E096F7AF884B22E00D4FBFE" lastPageId="14" lastPageNumber="94" pageId="12" pageNumber="92">
<subSubSection id="C30B656A3E096F78F884B22E07CCFA56" pageId="12" pageNumber="92" type="nomenclature">
<paragraph id="8BAE36E13E096F78F884B22E014CFAB2" blockId="12.[151,1154,1374,1473]" box="[151,514,1374,1401]" pageId="12" pageNumber="92">
<heading id="D0E6818D3E096F78F884B22E014CFAB2" bold="true" box="[151,514,1374,1401]" fontSize="11" level="1" pageId="12" pageNumber="92" reason="1">
<emphasis id="B965EAF33E096F78F884B22E014CFAB2" bold="true" box="[151,514,1374,1401]" pageId="12" pageNumber="92">
<taxonomicName id="4C114D623E096F78F884B22E02D3FAB2" authority="Mendes &amp; Paiva &amp; Rizzo, 2024" authorityName="Mendes &amp; Paiva &amp; Rizzo" authorityYear="2024" box="[151,413,1374,1401]" class="Polychaeta" family="Scalibregmatidae" genus="Oligobregma" kingdom="Animalia" pageId="12" pageNumber="92" phylum="Annelida" rank="species" species="bakkeni" status="sp. nov.">
<emphasis id="B965EAF33E096F78F884B22E02D3FAB2" bold="true" box="[151,413,1374,1401]" italics="true" pageId="12" pageNumber="92">Oligobregma bakkeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25657883E096F78F9B7B22F014CFAB2" box="[420,514,1375,1401]" pageId="12" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36E13E096F78F884B2F207CCFA56" blockId="12.[151,1154,1374,1473]" box="[151,1154,1410,1437]" pageId="12" pageNumber="92">
https://zoobank.org/NomenclaturalActs/
<uuid id="FFB70C343E096F78FA70B2F207CCFA56" box="[611,1154,1410,1437]" pageId="12" pageNumber="92">F5A5522E-AA2B-4B26-B238-F9FC339A3CAB</uuid>
</paragraph>
</subSubSection>
<subSubSection id="C30B656A3E096F78F884B2D7027CFA0A" box="[151,306,1447,1473]" pageId="12" pageNumber="92" type="description">
<paragraph id="8BAE36E13E096F78F884B2D7027CFA0A" blockId="12.[151,1154,1374,1473]" box="[151,306,1447,1473]" pageId="12" pageNumber="92">
<figureCitation id="132A2A643E096F78F884B2D7024EFA0A" box="[151,256,1447,1473]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="12" pageNumber="92">Figures 6</figureCitation>
&amp;
<figureCitation id="132A2A643E096F78F937B2D7027CFA0A" box="[292,306,1447,1473]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="12" pageNumber="92">7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B656A3E096F78F884B29F0393F99A" pageId="12" pageNumber="92" type="materials_examined">
<paragraph id="8BAE36E13E096F78F884B29F0393F99A" blockId="12.[151,1437,1518,2013]" pageId="12" pageNumber="92">
<emphasis id="B965EAF33E096F78F884B29F0206F9C3" bold="true" box="[151,328,1518,1545]" pageId="12" pageNumber="92">Type material.</emphasis>
<materialsCitation id="3B793CBC3E096F78F941B29E0264F9E6" collectingDate="2012-01-04" collectionCode="UERJ, R" pageId="12" pageNumber="92" specimenCode="UERJ9225, R1" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB79E9A3E096F78F941B29E0283F9C2" box="[338,461,1518,1545]" collectionCode="UERJ" pageId="12" pageNumber="92">UERJ9225</specimenCode>
(
<typeStatus id="54AA88433E096F78F9CCB29F0103F9C2" box="[479,589,1519,1545]" pageId="12" pageNumber="92" type="holotype">Holotype</typeStatus>
: Ambes5; A9
<specimenCode id="DBB79E9A3E096F78FAFDB29F0040F9C2" box="[750,782,1519,1545]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="92">R1</specimenCode>
; Lat: -21.161194; Long: -38.873622;
<date id="FFAF10213E096F78FCD7B29F0618F9C2" box="[1220,1366,1519,1545]" pageId="12" pageNumber="92" value="2012-01-04">
<collectingDate id="EFEBE9C93E096F78FCD7B29F0618F9C2" box="[1220,1366,1519,1545]" pageId="12" pageNumber="92" value="2012-01-04">04 Jan 2012</collectingDate>
</date>
; 2498 meters deep)
</materialsCitation>
.
<materialsCitation id="3B793CBC3E096F78F920B1630397F99A" collectingDate="2012-01-06" collectionCode="UERJ, R" pageId="12" pageNumber="92" specimenCode="UERJ9226, R1" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBB79E9A3E096F78F920B16302E1F9E6" box="[307,431,1555,1581]" collectionCode="UERJ" pageId="12" pageNumber="92">UERJ9226</specimenCode>
(
<typeStatus id="54AA88433E096F78F9AFB163016AF9E6" box="[444,548,1555,1581]" pageId="12" pageNumber="92" type="paratype">Paratype</typeStatus>
: Ambes5; D9
<specimenCode id="DBB79E9A3E096F78FAAEB1630193F9E6" box="[701,733,1555,1581]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="92">R1</specimenCode>
; Lat: -20.576756; Long: -38.681556;
<date id="FFAF10213E096F78FC93B1630644F9E6" box="[1152,1290,1555,1581]" pageId="12" pageNumber="92" value="2012-01-06">
<collectingDate id="EFEBE9C93E096F78FC93B1630644F9E6" box="[1152,1290,1555,1581]" pageId="12" pageNumber="92" value="2012-01-06">06 Jan 2012</collectingDate>
</date>
; 2460 meters deep)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B656A3E096F7AF8D4B12A0716FDD2" lastPageId="14" lastPageNumber="94" pageId="12" pageNumber="92" type="description">
<paragraph id="8BAE36E13E096F78F8D4B12A02A7F8E2" blockId="12.[151,1437,1518,2013]" pageId="12" pageNumber="92">
<emphasis id="B965EAF33E096F78F8D4B12A0217F9BF" bold="true" box="[199,345,1626,1652]" pageId="12" pageNumber="92">Description.</emphasis>
<typeStatus id="54AA88433E096F78F971B12B0284F9BE" box="[354,458,1627,1653]" pageId="12" pageNumber="92" type="holotype">Holotype</typeStatus>
adult but incomplete, lacking posterior end. Measuring
<quantity id="4CE99B043E096F78FC42B12B07DAF9BE" box="[1105,1172,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="92" unit="mm" value="8.0">8 mm</quantity>
long,
<quantity id="4CE99B043E096F78FCCCB12A0676F9BE" box="[1247,1336,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="92" unit="mm" value="2.5">2.5 mm</quantity>
wide on expanded region and
<quantity id="4CE99B043E096F78F990B10F028DF952" box="[387,451,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="92" unit="mm" value="1.0">1 mm</quantity>
wide on narrowest region, for 18 chaetigers. Moderate sized species,
<typeStatus id="54AA88433E096F78FCD3B10F066CF952" box="[1216,1314,1663,1689]" pageId="12" pageNumber="92" type="paratype">Paratype</typeStatus>
measuring
<quantity id="4CE99B043E096F78F884B1D303A8F976" box="[151,230,1699,1725]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="92" unit="mm" value="10.0">10 mm</quantity>
long,
<quantity id="4CE99B043E096F78F938B1D30225F976" box="[299,363,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="92" unit="mm" value="1.0">1 mm</quantity>
wide on expanded region and 0.5 wide on narrowest region, for 23 chaetigerous segments. Both
<typeStatus id="54AA88433E096F78F884B1B703B7F92A" box="[151,249,1735,1761]" pageId="12" pageNumber="92" type="holotype">holotype</typeStatus>
and
<typeStatus id="54AA88433E096F78F93DB1B702C0F92A" box="[302,398,1735,1761]" pageId="12" pageNumber="92" type="paratype">paratype</typeStatus>
incomplete, lacking posterior end. Body arenicoliform, weakly expanded over chaetiger 3–8. Colour in alcohol yellowish. Body surface covered by secondarily annulated rings. Secondary annuli smooth in both anterior and posterior regions.
</paragraph>
<paragraph id="8BAE36E13E096F79F8D4B04301BBFF1E" blockId="12.[151,1437,1518,2013]" lastBlockId="13.[151,1436,151,213]" lastPageId="13" lastPageNumber="93" pageId="12" pageNumber="92">
Triangular prostomium, with two long horns, both projected laterally (
<figureCitation id="132A2A643E096F78FC13B043072CF886" box="[1024,1122,1843,1869]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="12" pageNumber="92">Figs 6A</figureCitation>
;
<figureCitation id="132A2A643E096F78FC61B04307D6F886" box="[1138,1176,1843,1869]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="12" pageNumber="92">7A</figureCitation>
). Eyes absent. Nuchal organs observed, present as dorso-lateral saccular projections between prostomium and peristomium. Peristomium achaetous, uniannulated dorsally and ventrally. Proboscis smooth (
<figureCitation id="132A2A643E096F78FB96B00B0090F85E" box="[901,990,1915,1941]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="12" pageNumber="92">Fig. 7B</figureCitation>
). Ventral groove present from chaetiger 2, with quadrangular uni- to triannulated pads from chaetiger 2, forming a ventral mid-ridge up to end of body. Each pad paired to a single chaetiger. Posterior pads smaller and narrower than anterior ones. Dorsally and ventrally, chaetiger 1 presents two secondary annuli; from chaetiger 2, three annuli connect to parapodial lobe, plus one superior and inferior annulation delimitating segment.
</paragraph>
<caption id="DF6E66693E086F79F884B2000396F9D3" ID-DOI="http://doi.org/10.5281/zenodo.10808069" ID-Zenodo-Dep="10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93" startId="13.[151,250,1392,1416]" targetBox="[151,1436,238,1368]" targetPageId="13" targetType="figure">
<paragraph id="8BAE36E13E086F79F884B2000396F9D3" blockId="13.[151,1437,1392,1560]" pageId="13" pageNumber="93">
<emphasis id="B965EAF33E086F79F884B200025DFA43" bold="true" box="[151,275,1392,1416]" pageId="13" pageNumber="93">FIGURE 7.</emphasis>
<taxonomicName id="4C114D623E086F79F90AB20002BBFA43" authority="Mendes &amp; Paiva &amp; Rizzo, 2024" authorityName="Mendes &amp; Paiva &amp; Rizzo" authorityYear="2024" box="[281,501,1392,1416]" class="Polychaeta" family="Scalibregmatidae" genus="Oligobregma" kingdom="Animalia" pageId="13" pageNumber="93" phylum="Annelida" rank="species" species="bakkeni" status="sp. nov.">
<emphasis id="B965EAF33E086F79F90AB20002BBFA43" box="[281,501,1392,1416]" italics="true" pageId="13" pageNumber="93">Oligobregma bakkeni</emphasis>
</taxonomicName>
<emphasis id="B965EAF33E086F79F9E8B20001F8FA43" bold="true" box="[507,694,1392,1416]" pageId="13" pageNumber="93">
<taxonomicNameLabel id="A25657883E086F79F9E8B2000102FA43" box="[507,588,1392,1416]" pageId="13" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
Holotype
</emphasis>
in optical microscopy.
<emphasis id="B965EAF33E086F79FBB1B20000F5FA43" bold="true" box="[930,955,1392,1416]" pageId="13" pageNumber="93">A.</emphasis>
Anterior body in dorsal view.
<emphasis id="B965EAF33E086F79FCE2B200064CFA43" bold="true" box="[1265,1282,1392,1416]" pageId="13" pageNumber="93">B</emphasis>
. Anterior body in ventro-lateral view.
<emphasis id="B965EAF33E086F79F968B2E402C3FA67" bold="true" box="[379,397,1428,1452]" pageId="13" pageNumber="93">C</emphasis>
. Posterior body parapodium in anterior view.
<emphasis id="B965EAF33E086F79FB4DB2E40039FA67" bold="true" box="[862,887,1428,1452]" pageId="13" pageNumber="93">D.</emphasis>
Acicular spines from chaetiger 1–2.
<emphasis id="B965EAF33E086F79FCFFB2E4064DFA67" bold="true" box="[1260,1283,1428,1452]" pageId="13" pageNumber="93">E.</emphasis>
Acicular spine from chaetiger 4.
<emphasis id="B965EAF33E086F79F95BB2C80215FA1B" bold="true" box="[328,347,1464,1488]" pageId="13" pageNumber="93">F.</emphasis>
Lyrate chaetae. Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus.
</paragraph>
</caption>
<paragraph id="8BAE36E13E086F79F8D4B1360169F93B" blockId="13.[151,1437,1606,2029]" pageId="13" pageNumber="93">
Dorsal and ventral cirri present from chaetigers 13–14 (
<figureCitation id="132A2A643E086F79FB29B13600FAF9AB" box="[826,948,1606,1632]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="13" pageNumber="93">Figs 6E–F</figureCitation>
;
<figureCitation id="132A2A643E086F79FBD3B13600AAF9AB" box="[960,996,1606,1632]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93">7C</figureCitation>
). Interramal papillae present (
<figureCitation id="132A2A643E086F79FD26B13606C0F9AB" box="[1333,1422,1606,1632]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93">Fig. 7C</figureCitation>
); rounded when everted, observed from chaetiger 13, filled with granular content. Dorsal and ventral cirri present as a broad basis transitioning smoothly to a rounded tip (
<figureCitation id="132A2A643E086F79FAFFB1FE002BF963" box="[748,869,1678,1704]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="13" pageNumber="93">Figs 6E–F</figureCitation>
;
<figureCitation id="132A2A643E086F79FB7CB1FE00DDF963" box="[879,915,1678,1704]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93">7C</figureCitation>
). Strong and numerous blackish tubular glands may be present in all cirriferous chaetigers; sparse and less numerous initially, then densely entangled, filling all internal space posteriorly (
<figureCitation id="132A2A643E086F79F9D1B1A60154F93B" box="[450,538,1750,1776]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="8BAE36E13E086F79F8D4B18A0662F827" blockId="13.[151,1437,1606,2029]" pageId="13" pageNumber="93">
Parapodial lobes reduced in anterior chaetigers (
<figureCitation id="132A2A643E086F79FAC7B18A0067F8DF" box="[724,809,1786,1812]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93">Fig. 7B</figureCitation>
), becoming evident from midbody to posterior chaetigers (
<figureCitation id="132A2A643E086F79F88CB06E0256F8F3" box="[159,280,1822,1848]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="13" pageNumber="93">Figs 6E–F</figureCitation>
;
<figureCitation id="132A2A643E086F79F931B06E0208F8F3" box="[290,326,1822,1848]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93">7C</figureCitation>
). Notopodial and neuropodial lobes differing in size. Notopodial lobe always larger and longer than neuropodial lobe (
<figureCitation id="132A2A643E086F79F970B0320292F897" box="[355,476,1858,1884]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="13" pageNumber="93">Figs 6E–F</figureCitation>
;
<figureCitation id="132A2A643E086F79F9F4B0320145F897" box="[487,523,1858,1884]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93">7C</figureCitation>
). Both podial lobes of same shape on midbody to posterior chaetigers. Parapodial lobes initially present as inconspicuous asymmetrical elevations on anterior to midbody chaetigers; then become longer and wider, emerging as a broad elevation transitioning to an apically rounded tip, from first cirriferous chaetigers on midbody (
<figureCitation id="132A2A643E086F79F9B4B0DE02B0F803" box="[423,510,1966,1992]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="13" pageNumber="93">Fig. 6E</figureCitation>
). On posterior chaetigers, lobes becoming longer and thinner, emerging as a broad basis abruptly transitioning to a pointed tip, forming a “knife-like” asymmetrical structure (
<figureCitation id="132A2A643E086F79FC85B0A207A1F827" box="[1174,1263,2002,2028]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="13" pageNumber="93">Figs 6F</figureCitation>
;
<figureCitation id="132A2A643E086F79FCE8B0A20651F827" box="[1275,1311,2002,2028]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="13" pageNumber="93">7C</figureCitation>
).
</paragraph>
<paragraph id="8BAE36E13E0B6F7AF8D4B7E701E6FEAE" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="94">
Acicular spines present from chaetiger 1–4 on noto and neuropodia (
<figureCitation id="132A2A643E0B6F7AFBBCB7E70764FF7A" box="[943,1066,151,177]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="14" pageNumber="94">Figs 6B–C</figureCitation>
;
<figureCitation id="132A2A643E0B6F7AFC21B7E70737FF7A" box="[1074,1145,151,177]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="14" pageNumber="94">7D–E</figureCitation>
). On notopodia, chaetigers 1–2 present two anterior rows of strong acicular spines, with a terminal curved tip, plus a posterior row of capillary chaetae (
<figureCitation id="132A2A643E0B6F7AF8EFB7AF0215FF32" box="[252,347,223,249]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="14" pageNumber="94">Figs 6C</figureCitation>
;
<figureCitation id="132A2A643E0B6F7AF97AB7AF02C0FF32" box="[361,398,223,249]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="14" pageNumber="94">7D</figureCitation>
); numbering 9–10 spines on anterior and posterior rows. Chaetiger 3 presents on noto and neuropodia a single anterior row of 9–10 acicular spines, with the same shape of chaetigers 1–2, plus two posterior rows of capillaries. Chaetiger 4 presents one anterior row of nine straight and apically pointed acicular spines, plus two posterior rows of capillaries (
<figureCitation id="132A2A643E0B6F7AFA03B63B0122FEAE" box="[528,620,331,357]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="14" pageNumber="94">Figs 6B</figureCitation>
;
<figureCitation id="132A2A643E0B6F7AFA6AB63B01D5FEAE" box="[633,667,331,357]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="14" pageNumber="94">7E</figureCitation>
).
</paragraph>
<paragraph id="8BAE36E13E0B6F7AF8D4B61F0716FDD2" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="94">
Short spinous chaetae absent. Lyrate chaetae present from chaetiger 3 (
<figureCitation id="132A2A643E0B6F7AFC10B61F072BFE42" box="[1027,1125,367,393]" captionStart="FIGURE 6" captionStartId="11.[151,250,1900,1924]" captionTargetBox="[233,1357,199,1840]" captionTargetId="figure-26@11.[183,1405,181,1876]" captionTargetPageId="11" captionText="FIGURE 6. Oligobregma bakkeni sp. nov. Holotype line drawings. A. Anterior body in dorsal view. B. Acicular spine on chaetiger 4. C. Acicular spine on chaetigers 1–3. D. Lyrate chaeta. E. Midbody parapodium in anterior view. F. Posterior parapodium in anterior view. Abbreviations: dC = Dorsal cirrus; Hor = Prostomial horn; iP = Interramal papilla; NuO = Nuchal organ; Pe = Peristomium; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808067" httpUri="https://zenodo.org/record/10808067/files/figure.png" pageId="14" pageNumber="94">Figs 6D</figureCitation>
;
<figureCitation id="132A2A643E0B6F7AFC67B61F07DBFE42" box="[1140,1173,367,393]" captionStart="FIGURE 7" captionStartId="13.[151,250,1392,1416]" captionTargetBox="[151,1436,238,1368]" captionTargetId="figure-49@13.[151,1436,237,1369]" captionTargetPageId="13" captionText="FIGURE 7. Oligobregma bakkeni sp. nov. Holotype in optical microscopy. A. Anterior body in dorsal view. B. Anterior body in ventro-lateral view. C. Posterior body parapodium in anterior view. D. Acicular spines from chaetiger 1–2. E. Acicular spine from chaetiger 4. F. Lyrate chaetae.Abbreviations: As = Acicular spine; dC = Dorsal cirrus; iP = Interramal papillae; Ly = Lyrate chaeta; Nel = Neuropodial lobe; No = Nuchal organ; Nol = Notopodial Lobe; Pb = Proboscis; Pro = Prostomium; vC = Ventral cirrus." figureDoi="http://doi.org/10.5281/zenodo.10808069" httpUri="https://zenodo.org/record/10808069/files/figure.png" pageId="14" pageNumber="94">7F</figureCitation>
), numbering
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each fascicle on anterior chaetigers, reaching
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number on midbody chaetigers, then 4–5 on posterior chaetigers; always with subequal tynes (tyne’s ratio = 1.2) and in a single row anterior to capillaries. Anterior lyrate chaetae smaller, becoming longer on midbody to posterior chaetigers. Capillaries organized in one row on chaetigers 1–2, then two rows from chaetiger 3 up to posterior chaetigers. Pygidium nature unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30B656A3E0B6F7AF8D4B5530066FC6E" pageId="14" pageNumber="94" type="discussion">
<paragraph id="8BAE36E13E0B6F7AF8D4B55302DCFD06" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="94">
<emphasis id="B965EAF33E0B6F7AF8D4B5530275FDF6" bold="true" box="[199,315,547,573]" pageId="14" pageNumber="94">Remarks.</emphasis>
Although belonging to the subgroup of
<taxonomicName id="4C114D623E0B6F7AFB39B5530753FDF6" box="[810,1053,547,573]" pageId="14" pageNumber="94">
<emphasis id="B965EAF33E0B6F7AFB39B55300F0FDF6" box="[810,958,547,573]" italics="true" pageId="14" pageNumber="94">Oligobregma</emphasis>
species
</taxonomicName>
presenting acicular spines from chaetigers
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both rami,
<taxonomicName id="4C114D623E0B6F7AF9F9B5370129FDAA" authority="Mendes &amp; Paiva &amp; Rizzo, 2024" authorityName="Mendes &amp; Paiva &amp; Rizzo" authorityYear="2024" box="[490,615,583,609]" class="Polychaeta" family="Scalibregmatidae" genus="Oligobregma" kingdom="Animalia" pageId="14" pageNumber="94" phylum="Annelida" rank="species" species="bakkeni" status="sp. nov.">
<emphasis id="B965EAF33E0B6F7AF9F9B5370129FDAA" box="[490,615,583,609]" italics="true" pageId="14" pageNumber="94">O. bakkeni</emphasis>
</taxonomicName>
<emphasis id="B965EAF33E0B6F7AFA61B537019EFDAA" bold="true" box="[626,720,583,609]" pageId="14" pageNumber="94">
<taxonomicNameLabel id="A25657883E0B6F7AFA61B537019EFDAA" box="[626,720,583,609]" pageId="14" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from its congeners by the presence of acicular spines terminating with rounded curved tips on chaetigers 1–3, changing to transitional acicular spines on chaetiger 4. Only
<specimenCount id="9D17FD683E0B6F7AF884B5FF024CFD62" box="[151,258,655,681]" count="2" pageId="14" pageNumber="94" type="generic" typeStatus="types">two types</specimenCount>
of acicular spines are reported within
<taxonomicName id="4C114D623E0B6F7AFABAB5FF0073FD62" authorityName="Kudenov &amp; Blake" authorityYear="1975" box="[681,829,655,681]" class="Polychaeta" family="Scalibregmatidae" genus="Oligobregma" kingdom="Animalia" pageId="14" pageNumber="94" phylum="Annelida" rank="genus">
<emphasis id="B965EAF33E0B6F7AFABAB5FF0073FD62" box="[681,829,655,681]" italics="true" pageId="14" pageNumber="94">Oligobregma</emphasis>
</taxonomicName>
: curved or hirsute, both with or without terminal arista on their apical ending.
</paragraph>
<paragraph id="8BAE36E13E0B6F7AF8D4B5A70066FC6E" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="94">
Finally, the prostomium and its horns are also unique in this species, differing from all its congeners on the trapezoidal shape, “shield-like”, accompanied by long, and thin horns, these reaching the half of prostomium in length, always laterally directed. In
<taxonomicName id="4C114D623E0B6F7AFA27B46F01E1FCF2" authority="Mendes &amp; Paiva &amp; Rizzo, 2024" authorityName="Mendes &amp; Paiva &amp; Rizzo" authorityYear="2024" box="[564,687,799,825]" class="Polychaeta" family="Scalibregmatidae" genus="Oligobregma" kingdom="Animalia" pageId="14" pageNumber="94" phylum="Annelida" rank="species" species="bakkeni" status="sp. nov.">
<emphasis id="B965EAF33E0B6F7AFA27B46F01E1FCF2" box="[564,687,799,825]" italics="true" pageId="14" pageNumber="94">O. bakkeni</emphasis>
</taxonomicName>
<emphasis id="B965EAF33E0B6F7AFAABB46F005AFCF2" bold="true" box="[696,788,799,825]" pageId="14" pageNumber="94">
<taxonomicNameLabel id="A25657883E0B6F7AFAABB46F005AFCF2" box="[696,788,799,825]" pageId="14" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, lyrate chaetae co-occur with spinous chaetae, emerging from chaetiger 3. None of the species with acicular spines on chaetiger 1–4 has lyrate chaetae from chaetiger 3. However, species with acicular spines on chaetiger 1–2 possess lyrate chaetae emerging from chaetiger 3, but never co-occurring with acicular spines as in
<taxonomicName id="4C114D623E0B6F7AFA5DB4FB0189FC6E" authority="Mendes &amp; Paiva &amp; Rizzo, 2024" authorityName="Mendes &amp; Paiva &amp; Rizzo" authorityYear="2024" box="[590,711,907,933]" class="Polychaeta" family="Scalibregmatidae" genus="Oligobregma" kingdom="Animalia" pageId="14" pageNumber="94" phylum="Annelida" rank="species" species="bakkeni" status="sp. nov.">
<emphasis id="B965EAF33E0B6F7AFA5DB4FB0189FC6E" box="[590,711,907,933]" italics="true" pageId="14" pageNumber="94">O. bakkeni</emphasis>
</taxonomicName>
<emphasis id="B965EAF33E0B6F7AFADDB4FB0066FC6E" bold="true" box="[718,808,907,933]" pageId="14" pageNumber="94">
<taxonomicNameLabel id="A25657883E0B6F7AFADDB4FB0066FC6E" box="[718,808,907,933]" pageId="14" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30B656A3E0B6F7AF8D4B4DE009DFC26" pageId="14" pageNumber="94" type="distribution">
<paragraph id="8BAE36E13E0B6F7AF8D4B4DE009DFC26" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="94">
<emphasis id="B965EAF33E0B6F7AF8D4B4DE022FFC03" bold="true" box="[199,353,942,968]" pageId="14" pageNumber="94">Distribution.</emphasis>
<taxonomicName id="4C114D623E0B6F7AF975B4DF0293FC02" authority="Mendes &amp; Paiva &amp; Rizzo, 2024" authorityName="Mendes &amp; Paiva &amp; Rizzo" authorityYear="2024" box="[358,477,943,969]" class="Polychaeta" family="Scalibregmatidae" genus="Oligobregma" kingdom="Animalia" pageId="14" pageNumber="94" phylum="Annelida" rank="species" species="bakkeni" status="sp. nov.">
<emphasis id="B965EAF33E0B6F7AF975B4DF0293FC02" box="[358,477,943,969]" italics="true" pageId="14" pageNumber="94">O. bakkeni</emphasis>
</taxonomicName>
<emphasis id="B965EAF33E0B6F7AF9F1B4DF0174FC02" bold="true" box="[482,570,943,969]" pageId="14" pageNumber="94">
<taxonomicNameLabel id="A25657883E0B6F7AF9F1B4DF0174FC02" box="[482,570,943,969]" pageId="14" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found living on
<collectingRegion id="49D5F8033E0B6F7AFB0DB4DF00F5FC02" box="[798,955,943,969]" country="Brazil" name="Espirito Santo" pageId="14" pageNumber="94">Espírito Santo</collectingRegion>
basin continental slope, from a bathymetric range of 2460–2498 meters deep, predominantly among muddy sediment.
</paragraph>
</subSubSection>
<subSubSection id="C30B656A3E0B6F7AF8D4B48700D4FBFE" pageId="14" pageNumber="94" type="etymology">
<paragraph id="8BAE36E13E0B6F7AF8D4B48700D4FBFE" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="94">
<emphasis id="B965EAF33E0B6F7AF8D4B4870200FBDA" bold="true" box="[199,334,1015,1041]" pageId="14" pageNumber="94">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C114D623E0B6F7AFA5AB48701FDFBDA" authorityName="Mendes &amp; Paiva &amp; Rizzo" authorityYear="2024" box="[585,691,1015,1041]" class="Polychaeta" family="Scalibregmatidae" genus="Oligobregma" kingdom="Animalia" pageId="14" pageNumber="94" phylum="Annelida" rank="species" species="bakkeni">
“
<emphasis id="B965EAF33E0B6F7AFA45B48701FDFBDA" box="[598,691,1015,1041]" italics="true" pageId="14" pageNumber="94">bakkeni</emphasis>
</taxonomicName>
<emphasis id="B965EAF33E0B6F7AFAA1B48701F2FBDA" box="[690,700,1015,1041]" italics="true" pageId="14" pageNumber="94">”</emphasis>
was chosen to honor to Dr. Torkild Bakken, for his important contributions to
<taxonomicName id="4C114D623E0B6F7AF95CB36B015FFBFE" authorityName="Malmgren" authorityYear="1867" box="[335,529,1051,1077]" class="Polychaeta" family="Scalibregmatidae" kingdom="Animalia" pageId="14" pageNumber="94" phylum="Annelida" rank="family">Scalibregmatidae</taxonomicName>
taxonomy and annelids in general.
</paragraph>
</subSubSection>
</treatment>
</document>
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