treatments-xml/data/03/B5/EB/03B5EB07870CFFF3FF63FD9467B4A6A3.xml

<document id="DAC746DD99EAA8D0F3DFD88670A83F8A" ID-DOI="10.11646/zootaxa.5369.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10151336" ID-ZooBank="B16AF2B9-ADB0-4FA3-BA52-838560C328F5" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1700255300510" checkinUser="plazi" docAuthor="Cipola, Nikolas Gioia &amp; Viana, Stéphanie Dos Santos" docDate="2023" docId="03B5EB07870CFFF3FF63FD9467B4A6A3" docLanguage="en" docName="zootaxa.5369.3.3.pdf" docOrigin="Zootaxa 5369 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5369.3.3/52248" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidocyrtus (Fractocyrtus) andensis Cipola &amp; Viana 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="387" masterDocId="FF8C937F8706FFE1FFF4FF896507A427" masterDocTitle="Two new species of Lepidocyrtus (Fractocyrtus) Cipola &amp; Bellini (Collembola Entomobryidae, Lepidocyrtinae), with new records and a new name for Lepidocyrtus (F.) americanus Cipola &amp; Bellini, 2018" masterLastPageNumber="399" masterPageNumber="369" pageNumber="379" updateTime="1700596547940" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="28133F94B4D6B49228758F8155A85CDE">Two new species of Lepidocyrtus (Fractocyrtus) Cipola &amp; Bellini (Collembola Entomobryidae, Lepidocyrtinae), with new records and a new name for Lepidocyrtus (F.) americanus Cipola &amp; Bellini, 2018</mods:title>
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<mods:namePart id="5A197CEEE6C1AA82D7ADA5B6E659F209">Cipola, Nikolas Gioia</mods:namePart>
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<mods:affiliation id="E3802747BD23326FAED915D16BAB00C3">Laboratório de Sistemática e Ecologia de Invertebrados do Solo, Instituto Nacional de Pesquisas da Amazônia - INPA, CPEN. Avenida André Araújo, 2936, Aleixo, 69067 - 375, Manaus, AM, Brazil. &amp; Corresponding Author: Nikolas Gioia Cipola: nikolasgc @ gmail. com; https: // orcid. org / 0000 - 0002 - 7594 - 9574</mods:affiliation>
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<mods:namePart id="5C7CB500C98B237E586B702C5347CD88">Viana, Stéphanie Dos Santos</mods:namePart>
<mods:affiliation id="471A35896559AC12BFEF5EFA5E4B655F">Laboratório de Sistemática e Ecologia de Invertebrados do Solo, Instituto Nacional de Pesquisas da Amazônia - INPA, CPEN. Avenida André Araújo, 2936, Aleixo, 69067 - 375, Manaus, AM, Brazil.</mods:affiliation>
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<paragraph id="8BA35A11870CFFEBFF63FD9467B8A61F" blockId="10.[151,703,541,604]" box="[151,703,541,568]" pageId="10" pageNumber="379">
<heading id="D0EBED7D870CFFEBFF63FD9467B8A61F" bold="true" box="[151,703,541,568]" fontSize="11" level="1" pageId="10" pageNumber="379" reason="1">
<emphasis id="B9688603870CFFEBFF63FD9467B8A61F" bold="true" box="[151,703,541,568]" pageId="10" pageNumber="379">
<taxonomicName id="4C1C2192870CFFEBFF63FD94675CA61F" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[151,603,541,568]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="10" pageNumber="379" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B9688603870CFFEBFF63FD946434A61F" bold="true" box="[151,307,541,568]" italics="true" pageId="10" pageNumber="379">Lepidocyrtus</emphasis>
(
<emphasis id="B9688603870CFFEBFEB1FD9764E7A61F" bold="true" box="[325,480,542,568]" italics="true" pageId="10" pageNumber="379">Fractocyrtus</emphasis>
) 
<emphasis id="B9688603870CFFEBFE05FD94675CA61F" bold="true" box="[497,603,541,568]" italics="true" pageId="10" pageNumber="379">andensis</emphasis>
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<taxonomicNameLabel id="A25B3B78870CFFEBFD96FD9767B8A61F" box="[610,703,542,568]" pageId="10" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C306099A870CFFEBFF63FDCB6495A67B" box="[151,402,578,604]" pageId="10" pageNumber="379" type="description">
<paragraph id="8BA35A11870CFFEBFF63FDCB6495A67B" blockId="10.[151,703,541,604]" box="[151,402,578,604]" pageId="10" pageNumber="379">
<figureCitation id="13274694870CFFEBFF63FDCB640EA67B" box="[151,265,578,604]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStart-5="FIGURE 11" captionStart-6="FIGURE 12" captionStartId-0="11.[151,250,1873,1897]" captionStartId-1="12.[151,250,1943,1967]" captionStartId-2="13.[151,250,1943,1967]" captionStartId-3="14.[151,250,1943,1967]" captionStartId-4="15.[151,250,1943,1967]" captionStartId-5="16.[151,250,1451,1475]" captionStartId-6="17.[151,250,1908,1932]" captionTargetBox-0="[152,1434,346,1850]" captionTargetBox-1="[224,1363,181,1919]" captionTargetBox-2="[238,1356,181,1919]" captionTargetBox-3="[227,1360,181,1919]" captionTargetBox-4="[194,1394,191,1919]" captionTargetBox-5="[156,1436,181,1418]" captionTargetBox-6="[317,1272,187,1885]" captionTargetId-0="figure-117@11.[152,1435,345,1850]" captionTargetId-1="figure-22@12.[224,1363,181,1919]" captionTargetId-2="figure-17@13.[232,1356,181,1919]" captionTargetId-3="figure-22@14.[227,1360,181,1919]" captionTargetId-4="figure-17@15.[194,1394,181,1919]" captionTargetId-5="figure-22@16.[151,1436,181,1428]" captionTargetId-6="figure-17@17.[316,1272,181,1885]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="15" captionTargetPageId-5="16" captionTargetPageId-6="17" captionText-0="FIGURE 6A–C. Lepidocyrtus (Fractocyrtus) andensis sp. nov., habitus of specimens fixed in ethanol (lateral view); A, specimen with weak lateral spot on Abd IV (frequent color pattern); B, specimen with strong lateral spot on Th II (frequent color pattern); C, specimen with both spots on Th II and Abd IV, including a small spot between the Ant I base and eyepatches, and coxae I–III pigmented (atypical color pattern). Scale bar: 1 mm." captionText-1="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." captionText-2="FIGURE 8A–D. Lepidocyrtus (Fractocyrtus) andensis sp. nov., dorsal chaetotaxy (left side); A, Th II; B, Th III; C, Abd I; D, Abd II." captionText-3="FIGURE 9A–C. Lepidocyrtus (Fractocyrtus) andensis sp. nov., dorsal chaetotaxy (left side); A, Abd III; B, Abd IV; C, Abd V." captionText-4="FIGURE 10A–B. Lepidocyrtus (F.) andensis sp. nov., body psp distribution; A, dorsal side; B, ventral side." captionText-5="FIGURE 11A–E. Lepidocyrtus (F.) andensis sp. nov., legs; A–C, chaetotaxy of subcoxa I–III, respectively; D, trochanteral organ (posterior view); E, distal end of tibiotarsus and empodial complex I (anterior view)." captionText-6="FIGURE 12A–D. Lepidocyrtus (F.) andensis sp. nov., abdominal appendages; A, collophore (lateral view); B, distal end of manubrium (anterior view), arrow shows elongated scale; C, distal end of manubrium and proximal end of dens (lateral view), arrows on manubrial plate identify chaetae of different lengths, arrows on dens indicate dental spines and pseudopore present or absent; D, distal end of dens and mucro (outer view)." figureDoi-0="http://doi.org/10.5281/zenodo.10151352" figureDoi-1="http://doi.org/10.5281/zenodo.10151354" figureDoi-2="http://doi.org/10.5281/zenodo.10151356" figureDoi-3="http://doi.org/10.5281/zenodo.10151358" figureDoi-4="http://doi.org/10.5281/zenodo.10151360" figureDoi-5="http://doi.org/10.5281/zenodo.10151362" figureDoi-6="http://doi.org/10.5281/zenodo.10151364" httpUri-0="https://zenodo.org/record/10151352/files/figure.png" httpUri-1="https://zenodo.org/record/10151354/files/figure.png" httpUri-2="https://zenodo.org/record/10151356/files/figure.png" httpUri-3="https://zenodo.org/record/10151358/files/figure.png" httpUri-4="https://zenodo.org/record/10151360/files/figure.png" httpUri-5="https://zenodo.org/record/10151362/files/figure.png" httpUri-6="https://zenodo.org/record/10151364/files/figure.png" pageId="10" pageNumber="379">Figs 6–12</figureCitation>
, 
<figureCitation id="13274694870CFFEBFEE0FDCB6434A67B" box="[276,307,578,604]" captionStart="FIGURE 19" captionStartId="26.[151,250,885,909]" captionTargetBox="[163,1425,181,861]" captionTargetId="figure-22@26.[163,1425,181,861]" captionTargetPageId="26" captionText="FIGURE 19. Records map of Lepidocyrtus (Fractocyrtus) species; stars represent type localities and circles additional localities." figureDoi="http://doi.org/10.5281/zenodo.10151380" httpUri="https://zenodo.org/record/10151380/files/figure.png" pageId="10" pageNumber="379">19</figureCitation>
; 
<tableCitation id="C69E6FAA870CFFEBFECBFDCB6495A67B" box="[319,402,578,604]" captionStart="TABLE" captionStartId="7.[152,244,872,896]" captionTargetBox="[163,1398,918,1952]" captionTargetPageId="7" captionText="TABLE. 1. Comparison among Lepidocyrtus (Fractocyrtus) species." httpUri="http://table.plazi.org/id/DF630A998701FFE6FF6CFCE1667AA7A7" pageId="10" pageNumber="379" tableUuid="DF630A998701FFE6FF6CFCE1667AA7A7">Table 1</tableCitation>
</paragraph>
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<subSubSection id="C306099A870CFFEBFF63FD006467A7CF" pageId="10" pageNumber="379" type="diagnosis">
<paragraph id="8BA35A11870CFFEBFF63FD006467A7CF" blockId="10.[151,1437,649,2009]" pageId="10" pageNumber="379">
<emphasis id="B9688603870CFFEBFF63FD006415A684" bold="true" box="[151,274,649,675]" pageId="10" pageNumber="379">Diagnosis.</emphasis>
Body depigmented, if present, dark blue pigment on Th II laterally (rarely on Th III), a weak lateral spot present or absent on Abd IV and rarely, on coxae I–III pigmented (
<figureCitation id="13274694870CFFEBFC46FD2766F1A6EF" box="[946,1014,686,712]" captionStart="FIGURE 6" captionStartId="11.[151,250,1873,1897]" captionTargetBox="[152,1434,346,1850]" captionTargetId="figure-117@11.[152,1435,345,1850]" captionTargetPageId="11" captionText="FIGURE 6A–C. Lepidocyrtus (Fractocyrtus) andensis sp. nov., habitus of specimens fixed in ethanol (lateral view); A, specimen with weak lateral spot on Abd IV (frequent color pattern); B, specimen with strong lateral spot on Th II (frequent color pattern); C, specimen with both spots on Th II and Abd IV, including a small spot between the Ant I base and eyepatches, and coxae I–III pigmented (atypical color pattern). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10151352" httpUri="https://zenodo.org/record/10151352/files/figure.png" pageId="10" pageNumber="379">Fig. 6</figureCitation>
); Ant III sense organ club-shape (
<figureCitation id="13274694870CFFEBFA9BFD2765BAA6CB" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7A</figureCitation>
); head 
<emphasis id="B9688603870CFFEBFEF9FD586436A6CC" bold="true" box="[269,305,721,747]" pageId="10" pageNumber="379">An</emphasis>
series with 14–15 mac; series 
<emphasis id="B9688603870CFFEBFD7DFD58679AA6CC" bold="true" box="[649,669,721,747]" pageId="10" pageNumber="379">A</emphasis>
with 2 supplementary mac between 
<emphasis id="B9688603870CFFEBFBCDFD58615CA6CC" bold="true" box="[1081,1115,721,747]" pageId="10" pageNumber="379">A0</emphasis>
and 
<emphasis id="B9688603870CFFEBFB65FD5861B3A6CC" bold="true" box="[1169,1204,721,747]" pageId="10" pageNumber="379">A2</emphasis>
; 5 interocular ciliate chaetae (
<figureCitation id="13274694870CFFEBFF0FFD7F6455A737" box="[251,338,758,784]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7B</figureCitation>
); clypeal formula with 2 
<emphasis id="B9688603870CFFEBFD93FD7F6768A737" bold="true" box="[615,623,758,784]" pageId="10" pageNumber="379">l</emphasis>
, 9 
<emphasis id="B9688603870CFFEBFD7FFD7C6793A728" bold="true" box="[651,660,757,783]" pageId="10" pageNumber="379">f</emphasis>
and 3 
<emphasis id="B9688603870CFFEBFD2DFD7C67F5A728" bold="true" box="[729,754,757,783]" pageId="10" pageNumber="379">pf</emphasis>
chaetae (
<figureCitation id="13274694870CFFEBFCA8FD7F66B2A737" box="[860,949,758,784]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7C</figureCitation>
); labial papilla E 
<emphasis id="B9688603870CFFEBFB8FFD7F61A7A737" bold="true" box="[1147,1184,758,784]" pageId="10" pageNumber="379">l.p.</emphasis>
pointed and short, not reaching base of distal guard appendages papilla (
<figureCitation id="13274694870CFFEBFD20FC936636A713" box="[724,817,794,820]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7D</figureCitation>
); postlabial chaetotaxy with 3 spines, cephalic groove with 7 chaetae and 
<emphasis id="B9688603870CFFEBFE84FCB76494A77F" bold="true" box="[368,403,830,856]" pageId="10" pageNumber="379">X2</emphasis>
chaeta present (
<figureCitation id="13274694870CFFEBFDBDFCB767A7A77F" box="[585,672,830,856]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7E</figureCitation>
); Abd III with 5 lateral mac and 27 chaetae on lateral tuft (
<figureCitation id="13274694870CFFEBFAC0FCB76089A77F" box="[1332,1422,830,856]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[227,1360,181,1919]" captionTargetId="figure-22@14.[227,1360,181,1919]" captionTargetPageId="14" captionText="FIGURE 9A–C. Lepidocyrtus (Fractocyrtus) andensis sp. nov., dorsal chaetotaxy (left side); A, Abd III; B, Abd IV; C, Abd V." figureDoi="http://doi.org/10.5281/zenodo.10151358" httpUri="https://zenodo.org/record/10151358/files/figure.png" pageId="10" pageNumber="379">Fig. 9A</figureCitation>
); Abd IV with 12 lateral mac, 13 median sens and 29 posterior mes (
<figureCitation id="13274694870CFFEBFC8AFCEB66D2A75B" box="[894,981,866,892]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[227,1360,181,1919]" captionTargetId="figure-22@14.[227,1360,181,1919]" captionTargetPageId="14" captionText="FIGURE 9A–C. Lepidocyrtus (Fractocyrtus) andensis sp. nov., dorsal chaetotaxy (left side); A, Abd III; B, Abd IV; C, Abd V." figureDoi="http://doi.org/10.5281/zenodo.10151358" httpUri="https://zenodo.org/record/10151358/files/figure.png" pageId="10" pageNumber="379">Fig. 9B</figureCitation>
); BP4 region with 25–29 psp (
<figureCitation id="13274694870CFFEBFADDFCEB6089A75B" box="[1321,1422,866,892]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-17@15.[194,1394,181,1919]" captionTargetPageId="15" captionText="FIGURE 10A–B. Lepidocyrtus (F.) andensis sp. nov., body psp distribution; A, dorsal side; B, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10151360" httpUri="https://zenodo.org/record/10151360/files/figure.png" pageId="10" pageNumber="379">Fig. 10B</figureCitation>
); trochanteral organ with about 34 spine-like chaetae (
<figureCitation id="13274694870CFFEBFD1EFC0F6656A787" box="[746,849,902,928]" captionStart="FIGURE 11" captionStartId="16.[151,250,1451,1475]" captionTargetBox="[156,1436,181,1418]" captionTargetId="figure-22@16.[151,1436,181,1428]" captionTargetPageId="16" captionText="FIGURE 11A–E. Lepidocyrtus (F.) andensis sp. nov., legs; A–C, chaetotaxy of subcoxa I–III, respectively; D, trochanteral organ (posterior view); E, distal end of tibiotarsus and empodial complex I (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.10151362" httpUri="https://zenodo.org/record/10151362/files/figure.png" pageId="10" pageNumber="379">Fig. 11D</figureCitation>
); collophore anteriorly with 34 and posteriorly with 36 ciliate chaetae; lateral flap with 18 ciliate and 8 smooth chaetae (
<figureCitation id="13274694870CFFEBFC62FC2366F8A7E3" box="[918,1023,938,964]" captionStart="FIGURE 12" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[317,1272,187,1885]" captionTargetId="figure-17@17.[316,1272,181,1885]" captionTargetPageId="17" captionText="FIGURE 12A–D. Lepidocyrtus (F.) andensis sp. nov., abdominal appendages; A, collophore (lateral view); B, distal end of manubrium (anterior view), arrow shows elongated scale; C, distal end of manubrium and proximal end of dens (lateral view), arrows on manubrial plate identify chaetae of different lengths, arrows on dens indicate dental spines and pseudopore present or absent; D, distal end of dens and mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10151364" httpUri="https://zenodo.org/record/10151364/files/figure.png" pageId="10" pageNumber="379">Fig. 12A</figureCitation>
); dens dorsally with 10–12 proximal spines (
<figureCitation id="13274694870CFFEBFF18FC476453A7CF" box="[236,340,974,1000]" captionStart="FIGURE 12" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[317,1272,187,1885]" captionTargetId="figure-17@17.[316,1272,181,1885]" captionTargetPageId="17" captionText="FIGURE 12A–D. Lepidocyrtus (F.) andensis sp. nov., abdominal appendages; A, collophore (lateral view); B, distal end of manubrium (anterior view), arrow shows elongated scale; C, distal end of manubrium and proximal end of dens (lateral view), arrows on manubrial plate identify chaetae of different lengths, arrows on dens indicate dental spines and pseudopore present or absent; D, distal end of dens and mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10151364" httpUri="https://zenodo.org/record/10151364/files/figure.png" pageId="10" pageNumber="379">Fig. 12C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306099A870CFFEBFF33FC7B615CA0E7" pageId="10" pageNumber="379" type="materials_examined">
<paragraph id="8BA35A11870CFFEBFF33FC7B6175A05F" blockId="10.[151,1437,649,2009]" pageId="10" pageNumber="379">
<emphasis id="B9688603870CFFEBFF33FC7B6471A02C" bold="true" box="[199,374,1009,1036]" pageId="10" pageNumber="379">Type material.</emphasis>
<materialsCitation id="3B74504C870CFFEBFE89FC7B6455A073" collectingDate="2020-07" collectingMethod="malaise trap" collectionCode="CC, UFRN" collectorName="L Mazariegos" country="Colombia" elevation="3" latitude="5.495833" location="Mesenia-Paramillo" longLatPrecision="21" longitude="-75.88917" municipality="Jardin" pageId="10" pageNumber="379" specimenCode="INPA-CLL 0000303-305" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Antioquia Department" typeStatus="holotype">
<typeStatus id="54A7E4B3870CFFEBFE89FC7B64E2A02B" box="[381,485,1010,1036]" pageId="10" pageNumber="379" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1A9198870CFFEBFE19FC7B673EA02B" box="[493,569,1010,1036]" pageId="10" pageNumber="379" type="female">female</specimenCount>
on slide (UNIANDES): 
<collectingCountry id="F30B1A81870CFFEBFCA5FC7B66C5A02B" box="[849,962,1010,1036]" name="Colombia" pageId="10" pageNumber="379">Colombia</collectingCountry>
, 
<collectingRegion id="49D894F3870CFFEBFC39FC7B61CCA02B" box="[973,1227,1010,1036]" country="Colombia" name="Antioquia" pageId="10" pageNumber="379">Antioquia Department</collectingRegion>
, 
<collectingMunicipality id="6BC7C06B870CFFEBFB23FC7B601CA02B" box="[1239,1307,1010,1036]" pageId="10" pageNumber="379">Jardín</collectingMunicipality>
municipality, “ 
<location id="8EC30CCA870CFFEBFF3FFB9F64A2A017" LSID="urn:lsid:plazi:treatment:03B5EB07870CFFF3FF63FD9467B4A6A3:8EC30CCA870CFFEBFF3FFB9F64A2A017" box="[203,421,1046,1072]" country="Colombia" latitude="5.495833" longLatPrecision="21" longitude="-75.88917" municipality="Jardin" name="Mesenia-Paramillo" pageId="10" pageNumber="379" stateProvince="Antioquia Department">Mesenia-Paramillo</location>
” 
<location id="8EC30CCA870CFFEBFE43FB9F6768A017" LSID="urn:lsid:plazi:treatment:03B5EB07870CFFF3FF63FD9467B4A6A3:8EC30CCA870CFFEBFE43FB9F6768A017" box="[439,623,1046,1072]" country="Colombia" latitude="5.495833" longLatPrecision="21" longitude="-75.88917" municipality="Jardin" name="Natural Reserve" pageId="10" pageNumber="379" stateProvince="Antioquia Department">Natural Reserve</location>
, 
<geoCoordinate id="EE283CD6870CFFEBFD8EFB9F6601A016" box="[634,774,1046,1073]" degrees="05" direction="north" minutes="29" orientation="latitude" pageId="10" pageNumber="379" precision="15" seconds="45" value="5.495833">05°29’45”N</geoCoordinate>
, 
<geoCoordinate id="EE283CD6870CFFEBFCE6FB9F66A6A016" box="[786,929,1046,1073]" degrees="75" direction="west" minutes="53" orientation="longitude" pageId="10" pageNumber="379" precision="15" seconds="21" value="-75.88917">75°53’21”W</geoCoordinate>
(
<figureCitation id="13274694870CFFEBFC45FB9F6104A017" box="[945,1027,1046,1072]" captionStart="FIGURE 19" captionStartId="26.[151,250,885,909]" captionTargetBox="[163,1425,181,861]" captionTargetId="figure-22@26.[163,1425,181,861]" captionTargetPageId="26" captionText="FIGURE 19. Records map of Lepidocyrtus (Fractocyrtus) species; stars represent type localities and circles additional localities." figureDoi="http://doi.org/10.5281/zenodo.10151380" httpUri="https://zenodo.org/record/10151380/files/figure.png" pageId="10" pageNumber="379">Fig. 19</figureCitation>
), 
<quantity id="4CE4F7F4870CFFEBFBEDFB9F617DA017" box="[1049,1146,1046,1072]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="379" unit="m" value="3.0">
<elevation id="0031BD22870CFFEBFBEDFB9F617DA017" box="[1049,1146,1046,1072]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="379" unit="m" value="3.0">3.000 m</elevation>
</quantity>
., 
<date id="FFA27CD1870CFFEBFB7EFB9F61EDA017" box="[1162,1258,1046,1072]" pageId="10" pageNumber="379" value="2020-07">
<collectingDate id="EFE68539870CFFEBFB7EFB9F61EDA017" box="[1162,1258,1046,1072]" pageId="10" pageNumber="379" value="2020-07">vii.2020</collectingDate>
</date>
, 
<collectingMethod id="525D2206870CFFEBFB01FB9F6087A017" box="[1269,1408,1046,1072]" pageId="10" pageNumber="379">malaise trap</collectingMethod>
, 
<collectorName id="26E93FC7870CFFEBFA78FB9F641DA073" pageId="10" pageNumber="379">L Mazariegos</collectorName>
coll.
</materialsCitation>
<materialsCitation id="3B74504C870CFFEBFEACFBB36175A05F" collectingDate="2020-07" collectingMethod="malaise trap" collectionCode="CC, UFRN" collectorName="L Mazariegos" country="Colombia" elevation="3" latitude="5.495833" location="Mesenia-Paramillo" longLatPrecision="21" longitude="-75.88917" municipality="Jardin" pageId="10" pageNumber="379" specimenCode="INPA-CLL 0000303-305" specimenCount="18" specimenCount-female="11" specimenCount-male="1" stateProvince="Antioquia" typeStatus="paratype">
18 
<typeStatus id="54A7E4B3870CFFEBFE8EFBB364EDA073" box="[378,490,1082,1108]" pageId="10" pageNumber="379">paratypes</typeStatus>
: 
<specimenCount id="9D1A9198870CFFEBFE00FBB36739A073" box="[500,574,1082,1108]" count="1" pageId="10" pageNumber="379" type="male">1 male</specimenCount>
and 
<specimenCount id="9D1A9198870CFFEBFD80FBB367D8A073" box="[628,735,1082,1108]" count="6" pageId="10" pageNumber="379" type="female">6 females</specimenCount>
on slides and 
<specimenCount id="9D1A9198870CFFEBFC89FBB36101A073" box="[893,1030,1082,1108]" count="6" pageId="10" pageNumber="379" type="generic">6 specimens</specimenCount>
in alcohol (UNIANDES), 
<specimenCount id="9D1A9198870CFFEBFAC5FBB3609BA073" box="[1329,1436,1082,1108]" count="3" pageId="10" pageNumber="379" type="female">3 females</specimenCount>
(
<specimenCode id="DBBAF26A870CFFEBFF6AFBD764B8A05E" box="[158,447,1118,1145]" collectionCode="INPA-CLL" pageId="10" pageNumber="379">INPA-CLL 0000303–305</specimenCode>
) and 
<specimenCount id="9D1A9198870CFFEBFE09FBD7676EA05F" box="[509,617,1118,1144]" count="2" pageId="10" pageNumber="379" type="female">2 females</specimenCount>
on slides (
<collectionCode id="ED0DC2D4870CFFEBFD16FBD7660EA05F" box="[738,777,1118,1144]" country="Australia" name="CSIRO Canberra Rhizobium Collection" pageId="10" pageNumber="379">CC</collectionCode>
/ 
<collectionCode id="ED0DC2D4870CFFEBFCE5FBD76666A05F" box="[785,865,1118,1144]" country="Brazil" lsid="urn:lsid:biocol.org:col:15771" name="Universidade Federal do Rio Grande do Norte" pageId="10" pageNumber="379" type="Herbarium">UFRN</collectionCode>
): same date as holotype.
</materialsCitation>
</paragraph>
<paragraph id="8BA35A11870CFFEBFF33FB08615CA0E7" blockId="10.[151,1437,649,2009]" pageId="10" pageNumber="379">
<emphasis id="B9688603870CFFEBFF33FB0864F8A0BC" bold="true" box="[199,511,1153,1179]" pageId="10" pageNumber="379">Other examined material.</emphasis>
<materialsCitation id="3B74504C870CFFEBFDFCFB0B615CA0E7" collectingDate="2022-09-29" collectionCode="INPA" collectorName="U Rendon" elevation="2" pageId="10" pageNumber="379" specimenCount="1" typeStatus="holotype">
<specimenCount id="9D1A9198870CFFEBFDFCFB0B678FA0BB" box="[520,648,1154,1180]" count="1" pageId="10" pageNumber="379" type="generic">1 specimen</specimenCount>
preserved in absolute alcohol (
<collectionCode id="ED0DC2D4870CFFEBFC1AFB0B6134A0BB" box="[1006,1075,1154,1180]" collectionName="Brazil, Amazonas, Manaus, Instituto Nacional de Pesquisas da Amazoonia, Colecao Sistematica da Entomologia" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="10" pageNumber="379">INPA</collectionCode>
), same date as holotype, except 
<quantity id="4CE4F7F4870CFFEBFF63FB2F65FFA0E7" box="[151,248,1190,1216]" metricMagnitude="0" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="379" unit="m" value="2.4">
<elevation id="0031BD22870CFFEBFF63FB2F65FFA0E7" box="[151,248,1190,1216]" metricMagnitude="0" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="379" unit="m" value="2.4">2.400 m</elevation>
</quantity>
., 
<date id="FFA27CD1870CFFEBFEFCFB2F6485A0E7" box="[264,386,1190,1216]" pageId="10" pageNumber="379" value="2022-09-29">
<collectingDate id="EFE68539870CFFEBFEFCFB2F6485A0E7" box="[264,386,1190,1216]" pageId="10" pageNumber="379" value="2022-09-29">29.ix.2022</collectingDate>
</date>
, manual collect in decomposing trunk in the soil, 
<collectorName id="26E93FC7870CFFEBFC45FB2F6124A0E7" box="[945,1059,1190,1216]" pageId="10" pageNumber="379">U Rendon</collectorName>
coll.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C306099A870CFFF1FF33FB4066D7A3BF" lastPageId="16" lastPageNumber="385" pageId="10" pageNumber="379" type="description">
<paragraph id="8BA35A11870CFFEBFF33FB4061FCA177" blockId="10.[151,1437,649,2009]" pageId="10" pageNumber="379">
<emphasis id="B9688603870CFFEBFF33FB40645EA0C4" bold="true" box="[199,345,1225,1251]" pageId="10" pageNumber="379">Description.</emphasis>
Total length (head+ trunk) of specimens 
<quantity id="4CE4F7F4870CFFEBFCCDFB4366D8A0C3" box="[825,991,1226,1253]" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.57" metricValueMin="2.18" pageId="10" pageNumber="379" unit="mm" value="2.375" valueMax="2.57" valueMin="2.18">2.18–2.57 mm</quantity>
(n =4), 
<typeStatus id="54A7E4B3870CFFEBFBCBFB4361A6A0C3" box="[1087,1185,1226,1252]" pageId="10" pageNumber="379" type="holotype">holotype</typeStatus>
<quantity id="4CE4F7F4870CFFEBFB5FFB436012A0C3" box="[1195,1301,1226,1252]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="10" pageNumber="379" unit="mm" value="2.18">2.18 mm</quantity>
. Specimens whitish with violet pigments on distal half or completely of Ant IV and Ant III–II distally (present or absent); dark blue pigment rarely present on Ant I base and eyepatches, Th II–III and Abd IV laterally with or without one weak lateral spot, and rarely one spot present on coxae I–III; eyepatches black (present or absent) (
<figureCitation id="13274694870CFFEBFB5CFABF61EBA177" box="[1192,1260,1334,1360]" captionStart="FIGURE 6" captionStartId="11.[151,250,1873,1897]" captionTargetBox="[152,1434,346,1850]" captionTargetId="figure-117@11.[152,1435,345,1850]" captionTargetPageId="11" captionText="FIGURE 6A–C. Lepidocyrtus (Fractocyrtus) andensis sp. nov., habitus of specimens fixed in ethanol (lateral view); A, specimen with weak lateral spot on Abd IV (frequent color pattern); B, specimen with strong lateral spot on Th II (frequent color pattern); C, specimen with both spots on Th II and Abd IV, including a small spot between the Ant I base and eyepatches, and coxae I–III pigmented (atypical color pattern). Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10151352" httpUri="https://zenodo.org/record/10151352/files/figure.png" pageId="10" pageNumber="379">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BA35A11870CFFEBFF33FAD36037A29F" blockId="10.[151,1437,649,2009]" pageId="10" pageNumber="379">
Head. Ratio antennae: trunk = 1: 2.53–2.81 (n=4), 
<typeStatus id="54A7E4B3870CFFEBFD01FAD36650A153" box="[757,855,1370,1396]" pageId="10" pageNumber="379" type="holotype">holotype</typeStatus>
1: 2.72; antennal ratio as I: II: III: IV = 1: 1.82–2.07: 1.71–2.05: 2.78–3.28 (n=4), 
<typeStatus id="54A7E4B3870CFFEBFE2AFAF76747A1BF" box="[478,576,1406,1432]" pageId="10" pageNumber="379" type="holotype">holotype</typeStatus>
1: 1.82: 1.71: 2.78. Ant III distally with 2 club-shape sens (sense organ), 3 guard sens and at least 6 finger-shape sens (
<figureCitation id="13274694870CFFEBFD8DFA2B67D5A19B" box="[633,722,1442,1468]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7A</figureCitation>
). Eye B larger than others, G and H smallest, interocular chaetae (
<emphasis id="B9688603870CFFEBFF55FA4F65B6A1C7" bold="true" box="[161,177,1478,1504]" pageId="10" pageNumber="379">p</emphasis>
) as mac, others as mes (
<emphasis id="B9688603870CFFEBFE37FA4F64D4A1C7" bold="true" box="[451,467,1478,1504]" pageId="10" pageNumber="379">q</emphasis>
, 
<emphasis id="B9688603870CFFEBFE15FA4F64E8A1C7" bold="true" box="[481,495,1478,1504]" pageId="10" pageNumber="379">v</emphasis>
, 
<emphasis id="B9688603870CFFEBFE09FA4F670EA1C7" bold="true" box="[509,521,1478,1504]" pageId="10" pageNumber="379">r</emphasis>
, 
<emphasis id="B9688603870CFFEBFDE3FA4F6727A1C7" bold="true" box="[535,544,1478,1504]" pageId="10" pageNumber="379">t</emphasis>
); head dorsal chaetotaxy (
<figureCitation id="13274694870CFFEBFCBDFA4F6699A1C7" box="[841,926,1478,1504]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7B</figureCitation>
) with 14–15 ‘An’ (
<emphasis id="B9688603870CFFEBFB83FA4C61D4A1C7" bold="true" box="[1143,1235,1477,1504]" pageId="10" pageNumber="379">An1a–3</emphasis>
), 4–5 ‘A’ (
<emphasis id="B9688603870CFFEBFABAFA4C608BA1C7" bold="true" box="[1358,1420,1477,1504]" pageId="10" pageNumber="379">A0–5</emphasis>
), 6 ‘M’ (
<emphasis id="B9688603870CFFEBFF11FA63642EA223" bold="true" box="[229,297,1514,1540]" pageId="10" pageNumber="379">M0–4</emphasis>
), 6 ‘S’ (
<emphasis id="B9688603870CFFEBFE71FA6064A5A224" bold="true" box="[389,418,1513,1539]" pageId="10" pageNumber="379">S0</emphasis>
, 
<emphasis id="B9688603870CFFEBFE44FA6064EEA223" bold="true" box="[432,489,1513,1540]" pageId="10" pageNumber="379">S2–4</emphasis>
, 
<emphasis id="B9688603870CFFEBFE03FA606737A223" bold="true" box="[503,560,1513,1540]" pageId="10" pageNumber="379">S6–7</emphasis>
), 3 ‘Ps’ (
<emphasis id="B9688603870CFFEBFD62FA6367DBA223" bold="true" box="[662,732,1514,1540]" pageId="10" pageNumber="379">Ps2–3</emphasis>
, 
<emphasis id="B9688603870CFFEBFD1EFA636613A223" bold="true" box="[746,788,1514,1540]" pageId="10" pageNumber="379">Ps5</emphasis>
), 4 ‘Pa’ (
<emphasis id="B9688603870CFFEBFC88FA6366C2A223" bold="true" box="[892,965,1514,1540]" pageId="10" pageNumber="379">Pa2–5</emphasis>
), 2 ‘Pm’ (
<emphasis id="B9688603870CFFEBFBC2FA63616BA223" bold="true" box="[1078,1132,1514,1540]" pageId="10" pageNumber="379">Pm1</emphasis>
, 
<emphasis id="B9688603870CFFEBFB8EFA6361B7A223" bold="true" box="[1146,1200,1514,1540]" pageId="10" pageNumber="379">Pm3</emphasis>
), 6 ‘Pp’ (
<emphasis id="B9688603870CFFEBFAEDFA636063A223" bold="true" box="[1305,1380,1514,1540]" pageId="10" pageNumber="379">Pp1–6</emphasis>
) and 2 ‘Pe’ (
<emphasis id="B9688603870CFFEBFF1CF9876439A20F" bold="true" box="[232,318,1550,1576]" pageId="10" pageNumber="379">Pe3–5?</emphasis>
) chaetae, plus 2 extra mac near to 
<emphasis id="B9688603870CFFEBFD4BF98467D4A200" bold="true" box="[703,723,1549,1575]" pageId="10" pageNumber="379">A</emphasis>
series. Clypeal formula with 2 (
<emphasis id="B9688603870CFFEBFBCDF987616CA20F" bold="true" box="[1081,1131,1550,1576]" pageId="10" pageNumber="379">l1–2</emphasis>
), 9 (
<emphasis id="B9688603870CFFEBFB6BF98461AFA200" bold="true" box="[1183,1192,1549,1575]" pageId="10" pageNumber="379">f</emphasis>
) and 3 (
<emphasis id="B9688603870CFFEBFAF1F984604FA20F" bold="true" box="[1285,1352,1549,1576]" pageId="10" pageNumber="379">pf0–2</emphasis>
) ciliate chaetae, 1 
<emphasis id="B9688603870CFFEBFEF3F9B86417A26C" bold="true" box="[263,272,1585,1611]" pageId="10" pageNumber="379">f</emphasis>
and 
<emphasis id="B9688603870CFFEBFEB2F9B8646AA26C" bold="true" box="[326,365,1585,1611]" pageId="10" pageNumber="379">pf0</emphasis>
unpaired, 
<emphasis id="B9688603870CFFEBFE10F9BB6711A26B" bold="true" box="[484,534,1586,1612]" pageId="10" pageNumber="379">l1–2</emphasis>
acuminate, 
<emphasis id="B9688603870CFFEBFD69F9BB67C8A26B" bold="true" box="[669,719,1586,1612]" pageId="10" pageNumber="379">l1–2</emphasis>
and 
<emphasis id="B9688603870CFFEBFCF1F9B8664FA26B" bold="true" box="[773,840,1585,1612]" pageId="10" pageNumber="379">pf0–1</emphasis>
larger, others subequal (
<figureCitation id="13274694870CFFEBFBAEF9BB61B5A26B" box="[1114,1202,1586,1612]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7C</figureCitation>
). Maxillary palp 
<emphasis id="B9688603870CFFEBFA86F9BB609AA26B" bold="true" box="[1394,1437,1586,1612]" pageId="10" pageNumber="379">b.c.</emphasis>
thicker and 1.35 larger than the 
<emphasis id="B9688603870CFFEBFE08F9DF6726A257" bold="true" box="[508,545,1622,1648]" pageId="10" pageNumber="379">t.a.</emphasis>
Labial papilla E 
<emphasis id="B9688603870CFFEBFD16F9DF6600A257" bold="true" box="[738,775,1622,1648]" pageId="10" pageNumber="379">l.p.</emphasis>
pointed (
<figureCitation id="13274694870CFFEBFC85F9DF66CCA257" box="[881,971,1622,1648]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7D</figureCitation>
). Ventral chaetotaxy with about 29 ciliate chaetae and 3 lateral spine-like mic; postlabial chaetotaxy forming three rows with 4 (
<emphasis id="B9688603870CFFEBFBBCF9F0618FA2B3" bold="true" box="[1096,1160,1657,1684]" pageId="10" pageNumber="379">G1–4</emphasis>
), 3–4 (
<emphasis id="B9688603870CFFEBFB23F9F36012A2B3" bold="true" box="[1239,1301,1658,1684]" pageId="10" pageNumber="379">X2–3</emphasis>
, 
<emphasis id="B9688603870CFFEBFAD6F9F36031A2B3" bold="true" box="[1314,1334,1658,1684]" pageId="10" pageNumber="379">X</emphasis>
, 
<emphasis id="B9688603870CFFEBFAB7F9F36062A2B3" bold="true" box="[1347,1381,1658,1684]" pageId="10" pageNumber="379">X4</emphasis>
) and 4 (
<emphasis id="B9688603870CFFEBFF42F91765F2A29F" bold="true" box="[182,245,1694,1720]" pageId="10" pageNumber="379">H1–4</emphasis>
) chaetae (except 
<emphasis id="B9688603870CFFEBFE4EF91764C9A29F" bold="true" box="[442,462,1694,1720]" pageId="10" pageNumber="379">X</emphasis>
as spine), respectively; cephalic groove with 8 chaetae, 
<emphasis id="B9688603870CFFEBFBB1F9176168A29F" bold="true" box="[1093,1135,1694,1720]" pageId="10" pageNumber="379">b.c.</emphasis>
longer (
<figureCitation id="13274694870CFFEBFB39F9176023A29F" box="[1229,1316,1694,1720]" captionStart="FIGURE 7" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[224,1363,181,1919]" captionTargetId="figure-22@12.[224,1363,181,1919]" captionTargetPageId="12" captionText="FIGURE 7A–E. Lepidocyrtus (Fractocyrtus) andensis sp. nov., head; A, Ant III distal end (lateral view); B, dorsal head chaetotaxy (left side); C, clypeal and prelabral chaetotaxy; D, labial papillae E (right side); E, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent." figureDoi="http://doi.org/10.5281/zenodo.10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="10" pageNumber="379">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="8BA35A11870CFFEBFF33F94B6175A327" blockId="10.[151,1437,649,2009]" pageId="10" pageNumber="379">
<collectorName id="26E93FC7870CFFEBFF33F94B641FA2FB" box="[199,280,1730,1756]" pageId="10" pageNumber="379">Thorax</collectorName>
chaetotaxy (
<figureCitation id="13274694870CFFEBFE51F94B671EA2FB" box="[421,537,1730,1756]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[238,1356,181,1919]" captionTargetId="figure-17@13.[232,1356,181,1919]" captionTargetPageId="13" captionText="FIGURE 8A–D. Lepidocyrtus (Fractocyrtus) andensis sp. nov., dorsal chaetotaxy (left side); A, Th II; B, Th III; C, Abd I; D, Abd II." figureDoi="http://doi.org/10.5281/zenodo.10151356" httpUri="https://zenodo.org/record/10151356/files/figure.png" pageId="10" pageNumber="379">Fig 8A–B</figureCitation>
). 
<collectorName id="26E93FC7870CFFEBFDDEF94B6765A2FB" box="[554,610,1730,1756]" pageId="10" pageNumber="379">Th II</collectorName>
<emphasis id="B9688603870CFFEBFD9CF94B6771A2FB" bold="true" box="[616,630,1730,1756]" pageId="10" pageNumber="379">a</emphasis>
, 
<emphasis id="B9688603870CFFEBFD76F94B679EA2FB" bold="true" box="[642,665,1730,1756]" pageId="10" pageNumber="379">m</emphasis>
and 
<emphasis id="B9688603870CFFEBFD3AF94B67D9A2FB" bold="true" box="[718,734,1730,1756]" pageId="10" pageNumber="379">p</emphasis>
series with 8, 7 and 8 mic, respectively. 
<collectorName id="26E93FC7870CFFEBFB6EF94B61BEA2FB" box="[1178,1209,1730,1756]" pageId="10" pageNumber="379">Th</collectorName>
<collectionCode id="ED0DC2D4870CFFEBFB4BF94B61DCA2FB" box="[1215,1243,1730,1756]" pageId="10" pageNumber="379">III</collectionCode>
<emphasis id="B9688603870CFFEBFB15F94B61E8A2FB" bold="true" box="[1249,1263,1730,1756]" pageId="10" pageNumber="379">a</emphasis>
, 
<emphasis id="B9688603870CFFEBFB08F94B6014A2FB" bold="true" box="[1276,1299,1730,1756]" pageId="10" pageNumber="379">m</emphasis>
and 
<emphasis id="B9688603870CFFEBFAB3F94B6050A2FB" bold="true" box="[1351,1367,1730,1756]" pageId="10" pageNumber="379">p</emphasis>
series with 6, 5 and 6 mic, respectively. 
<collectorName id="26E93FC7870CFFEBFDE5F96F6797A327" box="[529,656,1766,1792]" pageId="10" pageNumber="379">Ratio Th II</collectorName>
: 
<collectionCode id="ED0DC2D4870CFFEBFD68F96F67BFA327" box="[668,696,1766,1792]" pageId="10" pageNumber="379">III</collectionCode>
= 3.11–2.34: 1 (n=4), 
<typeStatus id="54A7E4B3870CFFEBFC40F96F6111A327" box="[948,1046,1766,1792]" pageId="10" pageNumber="379" type="holotype">holotype</typeStatus>
2.77: 1.
</paragraph>
<paragraph id="8BA35A11870CFFEAFF33F883671EA4DE" blockId="10.[151,1437,649,2009]" lastBlockId="11.[151,1436,150,321]" lastPageId="11" lastPageNumber="380" pageId="10" pageNumber="379">
Abdomen chaetotaxy (
<figureCitation id="13274694870CFFEBFE3CF883674FA303" box="[456,584,1802,1828]" captionStart="FIGURE 8" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[238,1356,181,1919]" captionTargetId="figure-17@13.[232,1356,181,1919]" captionTargetPageId="13" captionText="FIGURE 8A–D. Lepidocyrtus (Fractocyrtus) andensis sp. nov., dorsal chaetotaxy (left side); A, Th II; B, Th III; C, Abd I; D, Abd II." figureDoi="http://doi.org/10.5281/zenodo.10151356" httpUri="https://zenodo.org/record/10151356/files/figure.png" pageId="10" pageNumber="379">Figs 8C–D</figureCitation>
, 
<figureCitation id="13274694870CFFEBFDA2F883679AA303" box="[598,669,1802,1828]" captionStart="FIGURE 9" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[227,1360,181,1919]" captionTargetId="figure-22@14.[227,1360,181,1919]" captionTargetPageId="14" captionText="FIGURE 9A–C. Lepidocyrtus (Fractocyrtus) andensis sp. nov., dorsal chaetotaxy (left side); A, Abd III; B, Abd IV; C, Abd V." figureDoi="http://doi.org/10.5281/zenodo.10151358" httpUri="https://zenodo.org/record/10151358/files/figure.png" pageId="10" pageNumber="379">9A–C</figureCitation>
). Abd I 
<emphasis id="B9688603870CFFEBFD0FF883660EA303" bold="true" box="[763,777,1802,1828]" pageId="10" pageNumber="379">a</emphasis>
, 
<emphasis id="B9688603870CFFEBFCEDF8836637A303" bold="true" box="[793,816,1802,1828]" pageId="10" pageNumber="379">m</emphasis>
and 
<emphasis id="B9688603870CFFEBFC9FF883667CA303" bold="true" box="[875,891,1802,1828]" pageId="10" pageNumber="379">p</emphasis>
series with 5, 5 and 2 mic, respectively. Abd II with 
<emphasis id="B9688603870CFFEBFF25F8A765D8A36F" bold="true" box="[209,223,1838,1864]" pageId="10" pageNumber="379">a</emphasis>
, 
<emphasis id="B9688603870CFFEBFF19F8A76403A36F" bold="true" box="[237,260,1838,1864]" pageId="10" pageNumber="379">m</emphasis>
and 
<emphasis id="B9688603870CFFEBFEC8F8A7644BA36F" bold="true" box="[316,332,1838,1864]" pageId="10" pageNumber="379">p</emphasis>
series with 5, 7 and 5 chaetae (plus 
<emphasis id="B9688603870CFFEBFD13F8A767FCA36F" bold="true" box="[743,763,1838,1864]" pageId="10" pageNumber="379">el</emphasis>
), respectively; 1 (
<emphasis id="B9688603870CFFEBFC3CF8A766E9A36F" bold="true" box="[968,1006,1838,1864]" pageId="10" pageNumber="379">m3</emphasis>
) inner and 1 lateral (
<emphasis id="B9688603870CFFEBFB28F8A76006A36F" bold="true" box="[1244,1281,1838,1864]" pageId="10" pageNumber="379">m5</emphasis>
) mac; 
<emphasis id="B9688603870CFFEBFAA4F8A7606BA36F" bold="true" box="[1360,1388,1838,1864]" pageId="10" pageNumber="379">a5</emphasis>
and 
<emphasis id="B9688603870CFFEBFF63F8DB65BAA34B" bold="true" box="[151,189,1874,1900]" pageId="10" pageNumber="379">m2</emphasis>
bothriotricha with 5 and 3 fan-shaped chaetae, respectively. Abd III 
<emphasis id="B9688603870CFFEBFC31F8DB66D4A34B" bold="true" box="[965,979,1874,1900]" pageId="10" pageNumber="379">a</emphasis>
, 
<emphasis id="B9688603870CFFEBFC16F8DB66FEA34B" bold="true" box="[994,1017,1874,1900]" pageId="10" pageNumber="379">m</emphasis>
and 
<emphasis id="B9688603870CFFEBFBC5F8DB6146A34B" bold="true" box="[1073,1089,1874,1900]" pageId="10" pageNumber="379">p</emphasis>
series with 6, 7 and 6 chaetae, respectively; 5 lateral mac presents (
<emphasis id="B9688603870CFFEBFDC7F8FF676FA3B7" bold="true" box="[563,616,1910,1936]" pageId="10" pageNumber="379">pm6</emphasis>
, 
<emphasis id="B9688603870CFFEBFD83F8FF679BA3B7" bold="true" box="[631,668,1910,1936]" pageId="10" pageNumber="379">m7</emphasis>
, 
<emphasis id="B9688603870CFFEBFD5EF8FF67F5A3B7" bold="true" box="[682,754,1910,1936]" pageId="10" pageNumber="379">p6–8?</emphasis>
); 
<emphasis id="B9688603870CFFEBFCFFF8FF6637A3B7" bold="true" box="[779,816,1910,1936]" pageId="10" pageNumber="379">m2</emphasis>
bothriotrichum with 5, bothriotricha 
<emphasis id="B9688603870CFFEBFB21F8FF61F6A3B7" bold="true" box="[1237,1265,1910,1936]" pageId="10" pageNumber="379">a5</emphasis>
and 
<emphasis id="B9688603870CFFEBFADCF8FF6049A3B7" bold="true" box="[1320,1358,1910,1936]" pageId="10" pageNumber="379">m2</emphasis>
with 9 fan-shaped chaetae between them; lateral tuft with about 27 elongated chaetae. Abd IV series ‘ 
<emphasis id="B9688603870CFFEBFB30F81061DFA394" bold="true" box="[1220,1240,1945,1971]" pageId="10" pageNumber="379">A</emphasis>
’–‘ 
<emphasis id="B9688603870CFFEBFB0DF8136002A393" bold="true" box="[1273,1285,1946,1972]" pageId="10" pageNumber="379">r</emphasis>
’ with 5, 5, 5, 6, 4, 2, 5, 4, 4 and 2–7 chaetae, respectively, plus 
<emphasis id="B9688603870CFFEBFD3EF83467E6A3F0" bold="true" box="[714,737,1981,2007]" pageId="10" pageNumber="379">Si</emphasis>
and 
<emphasis id="B9688603870CFFEBFCEDF8346647A3F0" bold="true" box="[793,832,1981,2007]" pageId="10" pageNumber="379">Sm</emphasis>
mic; inner region with 2 mac (
<emphasis id="B9688603870CFFEBFB6BF83761DBA3FF" bold="true" box="[1183,1244,1982,2008]" pageId="10" pageNumber="379">B5–6</emphasis>
) and lateral with 12 mac (
<emphasis id="B9688603870DFFEAFF0FFF1F643DA497" bold="true" box="[251,314,150,177]" pageId="11" pageNumber="380">D2–3</emphasis>
, 
<emphasis id="B9688603870DFFEAFEBDFF1F6470A497" bold="true" box="[329,375,150,176]" pageId="11" pageNumber="380">De3</emphasis>
, 
<emphasis id="B9688603870DFFEAFE72FF1F64D4A497" bold="true" box="[390,467,150,177]" pageId="11" pageNumber="380">E1–4p</emphasis>
, 
<emphasis id="B9688603870DFFEAFE16FF1F672AA497" bold="true" box="[482,557,150,177]" pageId="11" pageNumber="380">F1–3p</emphasis>
); at least 16 sens (
<emphasis id="B9688603870DFFEAFCF7FF1E661BA496" bold="true" box="[771,796,151,177]" pageId="11" pageNumber="380">as</emphasis>
and 
<emphasis id="B9688603870DFFEAFCA1FF1E6668A496" bold="true" box="[853,879,151,177]" pageId="11" pageNumber="380">ps</emphasis>
type I, others type II) and 29 posterior mes. Abd V 
<emphasis id="B9688603870DFFEAFF46FF3265C7A4F2" bold="true" box="[178,192,187,213]" pageId="11" pageNumber="380">a</emphasis>
, 
<emphasis id="B9688603870DFFEAFF3AFF3265E2A4F2" bold="true" box="[206,229,187,213]" pageId="11" pageNumber="380">m</emphasis>
and 
<emphasis id="B9688603870DFFEAFEE8FF32642BA4F2" bold="true" box="[284,300,187,213]" pageId="11" pageNumber="380">p</emphasis>
series with 3, 3 and 5 mac (plus 2 chaetae of unknow homology), respectively. Ratio Abd III: IV = 1: 5.57–8.74 (n=4), 
<typeStatus id="54A7E4B3870DFFEAFEAFFF5664BAA4DE" box="[347,445,223,249]" pageId="11" pageNumber="380" type="holotype">holotype</typeStatus>
1: 5.72.
</paragraph>
<paragraph id="8BA35A11870DFFEAFF33FE8A66F7A566" blockId="11.[151,1436,150,321]" pageId="11" pageNumber="380">
Body psp distribution (
<figureCitation id="13274694870DFFEAFE3FFE8A6718A53A" box="[459,543,259,285]" captionStart="FIGURE 10" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[194,1394,191,1919]" captionTargetId="figure-17@15.[194,1394,181,1919]" captionTargetPageId="15" captionText="FIGURE 10A–B. Lepidocyrtus (F.) andensis sp. nov., body psp distribution; A, dorsal side; B, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10151360" httpUri="https://zenodo.org/record/10151360/files/figure.png" pageId="11" pageNumber="380">Fig. 10</figureCitation>
). Dorsal psp: dens proximally with 7 (6). Lateral psp: BP4 with 25–29. Ventral psp: Ant II with 1 (rarely 5) and tenaculum posteriorly with 2 (rarely 1 or 0).
</paragraph>
<caption id="DF630A99870DFFEAFF63F8D8664CA3F2" ID-DOI="http://doi.org/10.5281/zenodo.10151352" ID-Zenodo-Dep="10151352" httpUri="https://zenodo.org/record/10151352/files/figure.png" pageId="11" pageNumber="380" startId="11.[151,250,1873,1897]" targetBox="[152,1434,346,1850]" targetPageId="11" targetType="figure">
<paragraph id="8BA35A11870DFFEAFF63F8D8664CA3F2" blockId="11.[151,1437,1873,2005]" pageId="11" pageNumber="380">
<emphasis id="B9688603870DFFEAFF63F8D86445A34E" bold="true" box="[151,322,1873,1897]" pageId="11" pageNumber="380">FIGURE 6A–C</emphasis>
. 
<taxonomicName id="4C1C2192870DFFEAFEA7F8D867DDA34E" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[339,730,1873,1897]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="11" pageNumber="380" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B9688603870DFFEAFEA7F8D864D0A34E" box="[339,471,1873,1897]" italics="true" pageId="11" pageNumber="380">Lepidocyrtus</emphasis>
(
<emphasis id="B9688603870DFFEAFE1FF8DB6769A34E" box="[491,622,1874,1897]" italics="true" pageId="11" pageNumber="380">Fractocyrtus</emphasis>
) 
<emphasis id="B9688603870DFFEAFD75F8D867DDA34E" box="[641,730,1873,1897]" italics="true" pageId="11" pageNumber="380">andensis</emphasis>
</taxonomicName>
<emphasis id="B9688603870DFFEAFD10F8D8663EA34E" bold="true" box="[740,825,1873,1897]" pageId="11" pageNumber="380">
<taxonomicNameLabel id="A25B3B78870DFFEAFD10F8D8663EA34E" box="[740,825,1873,1897]" pageId="11" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus of specimens fixed in ethanol (lateral view); 
<emphasis id="B9688603870DFFEAFA70F8D86091A34E" bold="true" box="[1412,1430,1873,1897]" pageId="11" pageNumber="380">A</emphasis>
, specimen with weak lateral spot on Abd IV (frequent color pattern); 
<emphasis id="B9688603870DFFEAFC96F8FC6674A3AA" bold="true" box="[866,883,1909,1933]" pageId="11" pageNumber="380">B</emphasis>
, specimen with strong lateral spot on Th II (frequent color pattern); 
<emphasis id="B9688603870DFFEAFED9F8106438A396" bold="true" box="[301,319,1945,1969]" pageId="11" pageNumber="380">C</emphasis>
, specimen with both spots on Th II and Abd IV, including a small spot between the Ant I base and eyepatches, and coxae I–III pigmented (atypical color pattern). Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF630A99870AFFEDFF63F81E6151A3D0" ID-DOI="http://doi.org/10.5281/zenodo.10151354" ID-Zenodo-Dep="10151354" httpUri="https://zenodo.org/record/10151354/files/figure.png" pageId="12" pageNumber="381" startId="12.[151,250,1943,1967]" targetBox="[224,1363,181,1919]" targetPageId="12" targetType="figure">
<paragraph id="8BA35A11870AFFEDFF63F81E6151A3D0" blockId="12.[151,1436,1943,2039]" pageId="12" pageNumber="381">
<emphasis id="B9688603870AFFEDFF63F81E6440A388" bold="true" box="[151,327,1943,1967]" pageId="12" pageNumber="381">FIGURE 7A–E.</emphasis>
<taxonomicName id="4C1C2192870AFFEDFEA4F81E67D2A388" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[336,725,1943,1968]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="12" pageNumber="381" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B9688603870AFFEDFEA4F81E64D3A388" box="[336,468,1943,1967]" italics="true" pageId="12" pageNumber="381">Lepidocyrtus</emphasis>
(
<emphasis id="B9688603870AFFEDFE13F811676DA388" box="[487,618,1944,1967]" italics="true" pageId="12" pageNumber="381">Fractocyrtus</emphasis>
) 
<emphasis id="B9688603870AFFEDFD88F81E67D2A388" box="[636,725,1943,1967]" italics="true" pageId="12" pageNumber="381">andensis</emphasis>
</taxonomicName>
<emphasis id="B9688603870AFFEDFD2BF81E6634A388" bold="true" box="[735,819,1943,1967]" pageId="12" pageNumber="381">
<taxonomicNameLabel id="A25B3B78870AFFEDFD2BF81E6634A388" box="[735,819,1943,1967]" pageId="12" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, head; 
<emphasis id="B9688603870AFFEDFC70F81E6691A388" bold="true" box="[900,918,1943,1967]" pageId="12" pageNumber="381">A</emphasis>
, Ant III distal end (lateral view); 
<emphasis id="B9688603870AFFEDFAF0F81E6012A388" bold="true" box="[1284,1301,1943,1967]" pageId="12" pageNumber="381">B</emphasis>
, dorsal head chaetotaxy (left side); 
<emphasis id="B9688603870AFFEDFE8CF832648DA3F4" bold="true" box="[376,394,1979,2003]" pageId="12" pageNumber="381">C</emphasis>
, clypeal and prelabral chaetotaxy; 
<emphasis id="B9688603870AFFEDFD1DF83267FCA3F4" bold="true" box="[745,763,1979,2003]" pageId="12" pageNumber="381">D</emphasis>
, labial papillae E (right side); 
<emphasis id="B9688603870AFFEDFBDFF832613BA3F4" bold="true" box="[1067,1084,1979,2003]" pageId="12" pageNumber="381">E</emphasis>
, basomedian and basolateral labial fields and complete postlabial chaetotaxy (left side), arrow indicates chaetae present or absent.
</paragraph>
</caption>
<caption id="DF630A99870BFFECFF63F81E65E6A3F4" ID-DOI="http://doi.org/10.5281/zenodo.10151356" ID-Zenodo-Dep="10151356" httpUri="https://zenodo.org/record/10151356/files/figure.png" pageId="13" pageNumber="382" startId="13.[151,250,1943,1967]" targetBox="[238,1356,181,1919]" targetPageId="13" targetType="figure">
<paragraph id="8BA35A11870BFFECFF63F81E65E6A3F4" blockId="13.[151,1436,1943,2003]" pageId="13" pageNumber="382">
<emphasis id="B9688603870BFFECFF63F81E6442A388" bold="true" box="[151,325,1943,1967]" pageId="13" pageNumber="382">FIGURE 8A–D.</emphasis>
<taxonomicName id="4C1C2192870BFFECFEBFF81E67CEA388" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[331,713,1943,1967]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="13" pageNumber="382" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B9688603870BFFECFEBFF81E64C8A388" box="[331,463,1943,1967]" italics="true" pageId="13" pageNumber="382">Lepidocyrtus</emphasis>
(
<emphasis id="B9688603870BFFECFE2AF8116766A388" box="[478,609,1944,1967]" italics="true" pageId="13" pageNumber="382">Fractocyrtus</emphasis>
) 
<emphasis id="B9688603870BFFECFD84F81E67CEA388" box="[624,713,1943,1967]" italics="true" pageId="13" pageNumber="382">andensis</emphasis>
</taxonomicName>
<emphasis id="B9688603870BFFECFD24F81E6627A388" bold="true" box="[720,800,1943,1967]" pageId="13" pageNumber="382">
<taxonomicNameLabel id="A25B3B78870BFFECFD24F81E6627A388" box="[720,800,1943,1967]" pageId="13" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal chaetotaxy (left side); 
<emphasis id="B9688603870BFFECFBA2F81E616FA388" bold="true" box="[1110,1128,1943,1967]" pageId="13" pageNumber="382">A</emphasis>
, Th II; 
<emphasis id="B9688603870BFFECFB42F81E61C0A388" bold="true" box="[1206,1223,1943,1967]" pageId="13" pageNumber="382">B</emphasis>
, Th III; 
<emphasis id="B9688603870BFFECFAE9F81E6028A388" bold="true" box="[1309,1327,1943,1967]" pageId="13" pageNumber="382">C</emphasis>
, Abd I; 
<emphasis id="B9688603870BFFECFA70F81E6091A388" bold="true" box="[1412,1430,1943,1967]" pageId="13" pageNumber="382">D</emphasis>
, Abd II.
</paragraph>
</caption>
<caption id="DF630A998708FFEFFF63F81E65A9A3F4" ID-DOI="http://doi.org/10.5281/zenodo.10151358" ID-Zenodo-Dep="10151358" httpUri="https://zenodo.org/record/10151358/files/figure.png" pageId="14" pageNumber="383" startId="14.[151,250,1943,1967]" targetBox="[227,1360,181,1919]" targetPageId="14" targetType="figure">
<paragraph id="8BA35A118708FFEFFF63F81E65A9A3F4" blockId="14.[151,1437,1943,2003]" pageId="14" pageNumber="383">
<emphasis id="B96886038708FFEFFF63F81E6442A388" bold="true" box="[151,325,1943,1967]" pageId="14" pageNumber="383">FIGURE 9A–C.</emphasis>
<taxonomicName id="4C1C21928708FFEFFEB9F81167CAA397" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[333,717,1944,1968]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="14" pageNumber="383" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B96886038708FFEFFEB9F81164D6A397" box="[333,465,1944,1968]" italics="true" pageId="14" pageNumber="383">Lepidocyrtus</emphasis>
(
<emphasis id="B96886038708FFEFFE15F8116763A388" box="[481,612,1944,1967]" italics="true" pageId="14" pageNumber="383">Fractocyrtus</emphasis>
) 
<emphasis id="B96886038708FFEFFD80F81167CAA397" box="[628,717,1944,1968]" italics="true" pageId="14" pageNumber="383">andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038708FFEFFD21F8116621A397" bold="true" box="[725,806,1944,1968]" pageId="14" pageNumber="383">
<taxonomicNameLabel id="A25B3B788708FFEFFD21F8116621A397" box="[725,806,1944,1968]" pageId="14" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal chaetotaxy (left side); 
<emphasis id="B96886038708FFEFFB94F81E6175A388" bold="true" box="[1120,1138,1943,1967]" pageId="14" pageNumber="383">A</emphasis>
, Abd III; 
<emphasis id="B96886038708FFEFFB2EF81161ECA397" bold="true" box="[1242,1259,1944,1968]" pageId="14" pageNumber="383">B</emphasis>
, Abd IV; 
<emphasis id="B96886038708FFEFFAA6F81E6063A388" bold="true" box="[1362,1380,1943,1967]" pageId="14" pageNumber="383">C</emphasis>
, Abd V.
</paragraph>
</caption>
<caption id="DF630A998709FFEEFF63F81E61D8A388" ID-DOI="http://doi.org/10.5281/zenodo.10151360" ID-Zenodo-Dep="10151360" box="[151,1247,1943,1968]" httpUri="https://zenodo.org/record/10151360/files/figure.png" pageId="15" pageNumber="384" startId="15.[151,250,1943,1967]" targetBox="[194,1394,191,1919]" targetPageId="15" targetType="figure">
<paragraph id="8BA35A118709FFEEFF63F81E61D8A388" blockId="15.[151,1247,1943,1968]" box="[151,1247,1943,1968]" pageId="15" pageNumber="384">
<emphasis id="B96886038709FFEEFF63F81E6448A397" bold="true" box="[151,335,1943,1968]" pageId="15" pageNumber="384">FIGURE 10A–B.</emphasis>
<taxonomicName id="4C1C21928709FFEEFEA2F81E6764A388" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[342,611,1943,1968]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="15" pageNumber="384" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B96886038709FFEEFEA2F81E64DDA388" box="[342,474,1943,1967]" italics="true" pageId="15" pageNumber="384">Lepidocyrtus</emphasis>
(
<emphasis id="B96886038709FFEEFE1DF81164FAA388" box="[489,509,1944,1967]" italics="true" pageId="15" pageNumber="384">F.</emphasis>
) 
<emphasis id="B96886038709FFEEFDFEF81E6764A388" box="[522,611,1943,1967]" italics="true" pageId="15" pageNumber="384">andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038709FFEEFD9DF81167BDA397" bold="true" box="[617,698,1944,1968]" pageId="15" pageNumber="384">
<taxonomicNameLabel id="A25B3B788709FFEEFD9DF81167BDA397" box="[617,698,1944,1968]" pageId="15" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, body psp distribution; 
<emphasis id="B96886038709FFEEFC58F81E66B9A388" bold="true" box="[940,958,1943,1967]" pageId="15" pageNumber="384">A</emphasis>
, dorsal side; 
<emphasis id="B96886038709FFEEFBB1F8116151A397" bold="true" box="[1093,1110,1944,1968]" pageId="15" pageNumber="384">B</emphasis>
, ventral side.
</paragraph>
</caption>
<caption id="DF630A998716FFF1FF63FA226135A1CF" ID-DOI="http://doi.org/10.5281/zenodo.10151362" ID-Zenodo-Dep="10151362" httpUri="https://zenodo.org/record/10151362/files/figure.png" pageId="16" pageNumber="385" startId="16.[151,250,1451,1475]" targetBox="[156,1436,181,1418]" targetPageId="16" targetType="figure">
<paragraph id="8BA35A118716FFF1FF63FA226135A1CF" blockId="16.[151,1436,1451,1512]" pageId="16" pageNumber="385">
<emphasis id="B96886038716FFF1FF63FA226457A1E3" bold="true" box="[151,336,1451,1476]" pageId="16" pageNumber="385">FIGURE 11A–E.</emphasis>
<taxonomicName id="4C1C21928716FFF1FEADFA25676DA1E3" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[345,618,1452,1476]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="16" pageNumber="385" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B96886038716FFF1FEADFA2564DAA1E3" box="[345,477,1452,1476]" italics="true" pageId="16" pageNumber="385">Lepidocyrtus</emphasis>
(
<emphasis id="B96886038716FFF1FE1AFA256705A1E4" box="[494,514,1452,1475]" italics="true" pageId="16" pageNumber="385">F.</emphasis>
) 
<emphasis id="B96886038716FFF1FDE5FA25676DA1E3" box="[529,618,1452,1476]" italics="true" pageId="16" pageNumber="385">andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038716FFF1FD86FA2567C2A1E3" bold="true" box="[626,709,1452,1476]" pageId="16" pageNumber="385">
<taxonomicNameLabel id="A25B3B788716FFF1FD86FA2567C2A1E3" box="[626,709,1452,1476]" pageId="16" pageNumber="385" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, legs; 
<emphasis id="B96886038716FFF1FCF8FA22663AA1E4" bold="true" box="[780,829,1451,1475]" pageId="16" pageNumber="385">A–C</emphasis>
, chaetotaxy of subcoxa I–III, respectively; 
<emphasis id="B96886038716FFF1FAF6FA256013A1E3" bold="true" box="[1282,1300,1452,1476]" pageId="16" pageNumber="385">D</emphasis>
, trochanteral organ (posterior view); 
<emphasis id="B96886038716FFF1FE73FA59649FA1CF" bold="true" box="[391,408,1488,1512]" pageId="16" pageNumber="385">E</emphasis>
, distal end of tibiotarsus and empodial complex I (anterior view).
</paragraph>
</caption>
<paragraph id="8BA35A118716FFF1FF33F99F666DA2E7" blockId="16.[151,1437,1558,1980]" pageId="16" pageNumber="385">
Legs. Subcoxa I with one row of 7–8 chaetae; subcoxa II with 14 chaetae on row ‘a’ and about 16 microchaetae anteriorly, ‘p’ row with 9 chaetae; subcoxa III with one row of 21–22 chaetae and 4 anterior chaetae (
<figureCitation id="132746948716FFF1FAF7F9B36097A273" box="[1283,1424,1594,1620]" captionStart="FIGURE 11" captionStartId="16.[151,250,1451,1475]" captionTargetBox="[156,1436,181,1418]" captionTargetId="figure-22@16.[151,1436,181,1428]" captionTargetPageId="16" captionText="FIGURE 11A–E. Lepidocyrtus (F.) andensis sp. nov., legs; A–C, chaetotaxy of subcoxa I–III, respectively; D, trochanteral organ (posterior view); E, distal end of tibiotarsus and empodial complex I (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.10151362" httpUri="https://zenodo.org/record/10151362/files/figure.png" pageId="16" pageNumber="385">Figs 11A–C</figureCitation>
). Trochanteral organ with about 34 spine-like chaetae, 6 anterior, 7 posterior, 16 internal, 1 apical and 4 distal arm (
<figureCitation id="132746948716FFF1FF6BF90B6401A2BB" box="[159,262,1666,1692]" captionStart="FIGURE 11" captionStartId="16.[151,250,1451,1475]" captionTargetBox="[156,1436,181,1418]" captionTargetId="figure-22@16.[151,1436,181,1428]" captionTargetPageId="16" captionText="FIGURE 11A–E. Lepidocyrtus (F.) andensis sp. nov., legs; A–C, chaetotaxy of subcoxa I–III, respectively; D, trochanteral organ (posterior view); E, distal end of tibiotarsus and empodial complex I (anterior view)." figureDoi="http://doi.org/10.5281/zenodo.10151362" httpUri="https://zenodo.org/record/10151362/files/figure.png" pageId="16" pageNumber="385">Fig. 11D</figureCitation>
). Tenent hair 0.77 smaller than outer edge of unguis; tibiotarsal smooth chaeta 0.72 smaller than unguiculus length. Ratio unguis III: unguiculus III = 1: 
<quantity id="4CE4F7F48716FFF1FD5FF92F67FEA2E7" box="[683,761,1702,1728]" metricMagnitude="-2" metricUnit="m" metricValue="1.5493999999999999" pageId="16" pageNumber="385" unit="in" value="0.61">0.61 in</quantity>
<typeStatus id="54A7E4B38716FFF1FCF4F92F6661A2E7" box="[768,870,1702,1728]" pageId="16" pageNumber="385" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BA35A118716FFF1FF33F9436126A30B" blockId="16.[151,1437,1558,1980]" pageId="16" pageNumber="385">
Collophore (
<figureCitation id="132746948716FFF1FEA1F94364B8A2C3" box="[341,447,1738,1764]" captionStart="FIGURE 12" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[317,1272,187,1885]" captionTargetId="figure-17@17.[316,1272,181,1885]" captionTargetPageId="17" captionText="FIGURE 12A–D. Lepidocyrtus (F.) andensis sp. nov., abdominal appendages; A, collophore (lateral view); B, distal end of manubrium (anterior view), arrow shows elongated scale; C, distal end of manubrium and proximal end of dens (lateral view), arrows on manubrial plate identify chaetae of different lengths, arrows on dens indicate dental spines and pseudopore present or absent; D, distal end of dens and mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10151364" httpUri="https://zenodo.org/record/10151364/files/figure.png" pageId="16" pageNumber="385">Fig. 12A</figureCitation>
). Anterior side with about 34 ciliate chaetae, 3 mac, 4 mes, 2 thicker chaetae and about 25 thin chaetae of different lengths; posterior side with about 36 ciliated (3 unpaired) of different length and 1 distal smooth chaeta per side; lateral flap with 8 smooth chaetae and 18 ciliate chaetae.
</paragraph>
<paragraph id="8BA35A118716FFF1FF33F8BF66D7A3BF" blockId="16.[151,1437,1558,1980]" pageId="16" pageNumber="385">
Furcula. Manubrium ventrally with 2 longer subapical and about 22 apical scales, 4 of which are elongated (
<figureCitation id="132746948716FFF1FA9BF8BF65CEA353" captionStart="FIGURE 12" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[317,1272,187,1885]" captionTargetId="figure-17@17.[316,1272,181,1885]" captionTargetPageId="17" captionText="FIGURE 12A–D. Lepidocyrtus (F.) andensis sp. nov., abdominal appendages; A, collophore (lateral view); B, distal end of manubrium (anterior view), arrow shows elongated scale; C, distal end of manubrium and proximal end of dens (lateral view), arrows on manubrial plate identify chaetae of different lengths, arrows on dens indicate dental spines and pseudopore present or absent; D, distal end of dens and mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10151364" httpUri="https://zenodo.org/record/10151364/files/figure.png" pageId="16" pageNumber="385">Fig. 12B</figureCitation>
); manubrial plate with 10–15 ciliate chaetae, 4 inner as mac and 2–4 smaller, others subequal (
<figureCitation id="132746948716FFF1FB1DF8D36057A353" box="[1257,1360,1882,1908]" captionStart="FIGURE 12" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[317,1272,187,1885]" captionTargetId="figure-17@17.[316,1272,181,1885]" captionTargetPageId="17" captionText="FIGURE 12A–D. Lepidocyrtus (F.) andensis sp. nov., abdominal appendages; A, collophore (lateral view); B, distal end of manubrium (anterior view), arrow shows elongated scale; C, distal end of manubrium and proximal end of dens (lateral view), arrows on manubrial plate identify chaetae of different lengths, arrows on dens indicate dental spines and pseudopore present or absent; D, distal end of dens and mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10151364" httpUri="https://zenodo.org/record/10151364/files/figure.png" pageId="16" pageNumber="385">Fig. 12C</figureCitation>
). Dens dorsally with 10–12 proximal spines, 
<typeStatus id="54A7E4B38716FFF1FDCAF8F767A7A3BF" box="[574,672,1918,1944]" pageId="16" pageNumber="385" type="holotype">holotype</typeStatus>
with 10 and 11 (
<figureCitation id="132746948716FFF1FCA9F8F766C3A3BF" box="[861,964,1918,1944]" captionStart="FIGURE 12" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[317,1272,187,1885]" captionTargetId="figure-17@17.[316,1272,181,1885]" captionTargetPageId="17" captionText="FIGURE 12A–D. Lepidocyrtus (F.) andensis sp. nov., abdominal appendages; A, collophore (lateral view); B, distal end of manubrium (anterior view), arrow shows elongated scale; C, distal end of manubrium and proximal end of dens (lateral view), arrows on manubrial plate identify chaetae of different lengths, arrows on dens indicate dental spines and pseudopore present or absent; D, distal end of dens and mucro (outer view)." figureDoi="http://doi.org/10.5281/zenodo.10151364" httpUri="https://zenodo.org/record/10151364/files/figure.png" pageId="16" pageNumber="385">Fig. 12C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C306099A8716FFF1FF33F82B61FFA39B" box="[199,1272,1954,1980]" pageId="16" pageNumber="385" type="etymology">
<paragraph id="8BA35A118716FFF1FF33F82B61FFA39B" blockId="16.[151,1437,1558,1980]" box="[199,1272,1954,1980]" pageId="16" pageNumber="385">
<emphasis id="B96886038716FFF1FF33F82B644FA39B" bold="true" box="[199,328,1954,1980]" pageId="16" pageNumber="385">Etymology</emphasis>
. Refers to the region where the new species was found, Colombian Andes (
<figureCitation id="132746948716FFF1FB6CF82B61ECA39B" box="[1176,1259,1954,1980]" captionStart="FIGURE 19" captionStartId="26.[151,250,885,909]" captionTargetBox="[163,1425,181,861]" captionTargetId="figure-22@26.[163,1425,181,861]" captionTargetPageId="26" captionText="FIGURE 19. Records map of Lepidocyrtus (Fractocyrtus) species; stars represent type localities and circles additional localities." figureDoi="http://doi.org/10.5281/zenodo.10151380" httpUri="https://zenodo.org/record/10151380/files/figure.png" pageId="16" pageNumber="385">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF630A998717FFF0FF63F8FD67B5A3DE" ID-DOI="http://doi.org/10.5281/zenodo.10151364" ID-Zenodo-Dep="10151364" httpUri="https://zenodo.org/record/10151364/files/figure.png" pageId="17" pageNumber="386" startId="17.[151,250,1908,1932]" targetBox="[317,1272,187,1885]" targetPageId="17" targetType="figure">
<paragraph id="8BA35A118717FFF0FF63F8FD67B5A3DE" blockId="17.[151,1436,1908,2041]" pageId="17" pageNumber="386">
<emphasis id="B96886038717FFF0FF63F8FD6454A3AA" bold="true" box="[151,339,1908,1933]" pageId="17" pageNumber="386">FIGURE 12A–D.</emphasis>
<taxonomicName id="4C1C21928717FFF0FEAFF8FC676BA3AA" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[347,620,1909,1933]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="17" pageNumber="386" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B96886038717FFF0FEAFF8FC64D8A3AA" box="[347,479,1909,1933]" italics="true" pageId="17" pageNumber="386">Lepidocyrtus</emphasis>
(
<emphasis id="B96886038717FFF0FE04F8FC6703A3AB" box="[496,516,1909,1932]" italics="true" pageId="17" pageNumber="386">F.</emphasis>
) 
<emphasis id="B96886038717FFF0FDE7F8FC676BA3AA" box="[531,620,1909,1933]" italics="true" pageId="17" pageNumber="386">andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038717FFF0FD80F8FC67C1A3AA" bold="true" box="[628,710,1909,1933]" pageId="17" pageNumber="386">
<taxonomicNameLabel id="A25B3B788717FFF0FD80F8FC67C1A3AA" box="[628,710,1909,1933]" pageId="17" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, abdominal appendages; 
<emphasis id="B96886038717FFF0FC3AF8FD66E7A3AB" bold="true" box="[974,992,1908,1932]" pageId="17" pageNumber="386">A</emphasis>
, collophore (lateral view); 
<emphasis id="B96886038717FFF0FB08F8FC600AA3AA" bold="true" box="[1276,1293,1909,1933]" pageId="17" pageNumber="386">B</emphasis>
, distal end of manubrium (anterior view), arrow shows elongated scale; 
<emphasis id="B96886038717FFF0FD13F81167FEA397" bold="true" box="[743,761,1944,1968]" pageId="17" pageNumber="386">C</emphasis>
, distal end of manubrium and proximal end of dens (lateral view), arrows on manubrial plate identify chaetae of different lengths, arrows on dens indicate dental spines and pseudopore present or absent; 
<emphasis id="B96886038717FFF0FF11F86865F0A3DE" bold="true" box="[229,247,2017,2041]" pageId="17" pageNumber="386">D</emphasis>
, distal end of dens and mucro (outer view).
</paragraph>
</caption>
<subSubSection id="C306099A8714FFF3FF33FF1F67B4A6A3" pageId="18" pageNumber="387" type="discussion">
<paragraph id="8BA35A118714FFF3FF33FF1F67B4A6A3" blockId="18.[151,1437,150,645]" pageId="18" pageNumber="387">
<emphasis id="B96886038714FFF3FF33FF1F643CA497" bold="true" box="[199,315,150,176]" pageId="18" pageNumber="387">Remarks.</emphasis>
<taxonomicName id="4C1C21928714FFF3FEB0FF1F67EAA497" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[324,749,150,176]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov." subGenus="Fractocyrtus">
<emphasis id="B96886038714FFF3FEB0FF1F64D1A497" box="[324,470,150,176]" italics="true" pageId="18" pageNumber="387">Lepidocyrtus</emphasis>
(
<emphasis id="B96886038714FFF3FE1CFF1E677EA497" box="[488,633,151,176]" italics="true" pageId="18" pageNumber="387">Fractocyrtus</emphasis>
) 
<emphasis id="B96886038714FFF3FD7FFF1F67EAA497" box="[651,749,150,176]" italics="true" pageId="18" pageNumber="387">andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038714FFF3FD02FF1F6656A497" bold="true" box="[758,849,150,176]" pageId="18" pageNumber="387">
<taxonomicNameLabel id="A25B3B788714FFF3FD02FF1F6656A497" box="[758,849,150,176]" pageId="18" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C1C21928714FFF3FC27FF1E617EA497" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[979,1145,150,176]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi" subGenus="Fractocyrtus">
<emphasis id="B96886038714FFF3FC27FF1E66EDA497" box="[979,1002,151,176]" italics="true" pageId="18" pageNumber="387">L.</emphasis>
(
<emphasis id="B96886038714FFF3FC08FF1E6116A497" box="[1020,1041,151,176]" italics="true" pageId="18" pageNumber="387">F.</emphasis>
) 
<emphasis id="B96886038714FFF3FBD6FF1E617EA497" box="[1058,1145,151,176]" italics="true" pageId="18" pageNumber="387">mateosi</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EF8D27E08714FFF3FB4CFF1F65C4A4F3" author="Cipola, N. G. &amp; Morais, J. W. &amp; Bellini, B. C." pageId="18" pageNumber="387" pagination="189 - 234" refId="ref12839" refString="Cipola, N. G., Morais, J. W. &amp; Bellini, B. C. (2018 b) New subgenus and four species of Lepidocyrtus Bourlet (Collembola, Entomobryidae, Lepidocyrtinae) from Amazon. Insect systematics &amp; Evolution, 50 (2), 189 - 234. https: // doi. org / 10.1163 / 1876312 X- 00002184" type="journal article" year="2018">
Cipola 
<emphasis id="B96886038714FFF3FAF8FF1E6041A497" box="[1292,1350,150,176]" italics="true" pageId="18" pageNumber="387">et al.</emphasis>
2018b: 222
</bibRefCitation>
) in that the main sensilla of the Ant III sense organ are club-shape, interocular chaetotaxy only with ciliate chaetae, head series 
<emphasis id="B96886038714FFF3FEB3FF57645CA4DF" bold="true" box="[327,347,222,248]" pageId="18" pageNumber="387">A</emphasis>
with 2 extra mac, postlabial chaetotaxy with 3 spines and trochanteral organ with about 34 spine-like chaetae (
<tableCitation id="C69E6FAA8714FFF3FEDEFE8B6479A53B" box="[298,382,258,284]" captionStart="TABLE" captionStartId="7.[152,244,872,896]" captionTargetBox="[163,1398,918,1952]" captionTargetPageId="7" captionText="TABLE. 1. Comparison among Lepidocyrtus (Fractocyrtus) species." httpUri="http://table.plazi.org/id/DF630A998701FFE6FF6CFCE1667AA7A7" pageId="18" pageNumber="387" tableUuid="DF630A998701FFE6FF6CFCE1667AA7A7">Table 1</tableCitation>
). However, 
<taxonomicName id="4C1C21928714FFF3FDFEFE8A678EA53B" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[522,649,258,284]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov.">
<emphasis id="B96886038714FFF3FDFEFE8A678EA53B" box="[522,649,258,284]" italics="true" pageId="18" pageNumber="387">L. andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038714FFF3FD64FE8B67EDA53B" bold="true" box="[656,746,258,284]" pageId="18" pageNumber="387">
<taxonomicNameLabel id="A25B3B788714FFF3FD64FE8B67EDA53B" box="[656,746,258,284]" pageId="18" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from this species by the reduced body color pattern or with pigments restricted on Th II–III laterally and coxae, while 
<taxonomicName id="4C1C21928714FFF3FC9AFEAE66E3A567" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[878,996,295,320]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FC9AFEAE66E3A567" box="[878,996,295,320]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
has pigmented on head to Abd III and manubrium. On head, 
<taxonomicName id="4C1C21928714FFF3FE61FEC26712A543" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[405,533,330,356]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov.">
<emphasis id="B96886038714FFF3FE61FEC26712A543" box="[405,533,330,356]" italics="true" pageId="18" pageNumber="387">L. andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038714FFF3FDE9FEC36770A543" bold="true" box="[541,631,330,356]" pageId="18" pageNumber="387">
<taxonomicNameLabel id="A25B3B788714FFF3FDE9FEC36770A543" box="[541,631,330,356]" pageId="18" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs by clypeus with 9 
<emphasis id="B96886038714FFF3FC6CFEC366A6A543" bold="true" box="[920,929,330,356]" pageId="18" pageNumber="387">f</emphasis>
and 3 
<emphasis id="B96886038714FFF3FC13FEC36107A543" bold="true" box="[999,1024,330,356]" pageId="18" pageNumber="387">pf</emphasis>
chaetae (12 
<emphasis id="B96886038714FFF3FB7DFEC36195A543" bold="true" box="[1161,1170,330,356]" pageId="18" pageNumber="387">f</emphasis>
and 2 
<emphasis id="B96886038714FFF3FB2CFEC361F6A543" bold="true" box="[1240,1265,330,356]" pageId="18" pageNumber="387">pf</emphasis>
in 
<taxonomicName id="4C1C21928714FFF3FAE2FEC2608BA543" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[1302,1420,331,356]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FAE2FEC2608BA543" box="[1302,1420,331,356]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
), labial papilla E 
<emphasis id="B96886038714FFF3FEBCFEE7646AA5AF" bold="true" box="[328,365,366,392]" pageId="18" pageNumber="387">l.p.</emphasis>
pointed (finger-shape in 
<taxonomicName id="4C1C21928714FFF3FD73FEE667FBA5AF" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[647,764,367,392]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FD73FEE667FBA5AF" box="[647,764,367,392]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
) and cephalic groove with 7 chaetae (
<quantity id="4CE4F7F48714FFF3FB52FEE761D5A5AF" box="[1190,1234,366,393]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="18" pageNumber="387" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4C1C21928714FFF3FB2CFEE6604AA5AF" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[1240,1357,367,392]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FB2CFEE6604AA5AF" box="[1240,1357,367,392]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
). Concerning the abdominal dorsal chaetotaxy of 
<taxonomicName id="4C1C21928714FFF3FD74FE1A67F8A58B" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[640,767,402,428]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov.">
<emphasis id="B96886038714FFF3FD74FE1A67F8A58B" box="[640,767,402,428]" italics="true" pageId="18" pageNumber="387">L. andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038714FFF3FCF2FE1B6658A58B" bold="true" box="[774,863,402,428]" pageId="18" pageNumber="387">
<taxonomicNameLabel id="A25B3B788714FFF3FCF2FE1B6658A58B" box="[774,863,402,428]" pageId="18" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs by having Abd III with 5 lateral mac and 27 elongated chaetae on lateral tuft (3 and 9 respectively in 
<taxonomicName id="4C1C21928714FFF3FCF9FE3E6686A5F7" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[781,897,439,464]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FCF9FE3E6686A5F7" box="[781,897,439,464]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
), Abd IV has 12 lateral mac and 13 median sens (14 and 3 respectively in 
<taxonomicName id="4C1C21928714FFF3FE46FE526720A5D3" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[434,551,475,500]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FE46FE526720A5D3" box="[434,551,475,500]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
), BP4 region with 25–29 psp (
<quantity id="4CE4F7F48714FFF3FC8BFE5366E4A5D3" box="[895,995,474,501]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="6.35" metricValueMin="5.08" pageId="18" pageNumber="387" unit="in" value="22.5" valueMax="25.0" valueMin="20.0">20–25 in</quantity>
<taxonomicName id="4C1C21928714FFF3FC1EFE526158A5D3" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[1002,1119,475,500]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FC1EFE526158A5D3" box="[1002,1119,475,500]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
), and Abd V with 11 mac (
<quantity id="4CE4F7F48714FFF3FA7AFE5365AAA63F" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="18" pageNumber="387" unit="in" value="7.0">7 in</quantity>
<taxonomicName id="4C1C21928714FFF3FF41FE76642CA63F" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[181,299,511,536]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FF41FE76642CA63F" box="[181,299,511,536]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C1C21928714FFF3FEB7FE776738A63F" authority="Cipola &amp; Viana, 2023" authorityName="Cipola &amp; Viana" authorityYear="2023" box="[323,575,510,536]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="andensis" status="sp. nov.">
<emphasis id="B96886038714FFF3FEB7FE776738A63F" box="[323,575,510,536]" italics="true" pageId="18" pageNumber="387">Lepidocyrtus andensis</emphasis>
</taxonomicName>
<emphasis id="B96886038714FFF3FDB3FE7767A5A63F" bold="true" box="[583,674,510,536]" pageId="18" pageNumber="387">
<taxonomicNameLabel id="A25B3B788714FFF3FDB3FE7767A5A63F" box="[583,674,510,536]" pageId="18" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differs by having 10–12 spines dorsally on the dens (
<quantity id="4CE4F7F48714FFF3FAC3FE77609BA63F" box="[1335,1436,510,536]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.556" pageId="18" pageNumber="387" unit="in" value="15.0" valueMax="16.0" valueMin="14.0">14–16 in</quantity>
<taxonomicName id="4C1C21928714FFF3FF63FDAA640BA61B" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[151,268,547,572]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FF63FDAA640BA61B" box="[151,268,547,572]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
), collophore anteriorly with 34 anterior and 36 posterior ciliate chaetae (25 and 
<quantity id="4CE4F7F48714FFF3FB7FFDAB61C3A61B" box="[1163,1220,546,572]" metricMagnitude="0" metricUnit="m" metricValue="1.0922" pageId="18" pageNumber="387" unit="in" value="43.0">43 in</quantity>
<taxonomicName id="4C1C21928714FFF3FB3FFDAA6038A61B" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[1227,1343,547,572]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FB3FFDAA6038A61B" box="[1227,1343,547,572]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
), lateral flaps with 18 ciliate and 8 smooth chaetae, while 
<taxonomicName id="4C1C21928714FFF3FD4FFDCE6628A647" authorityName="Cipola &amp; Bellini" authorityYear="2018" box="[699,815,583,608]" class="Collembola" family="Entomobryidae" genus="Lepidocyrtus" kingdom="Animalia" order="Entomobryomorpha" pageId="18" pageNumber="387" phylum="Arthropoda" rank="species" species="mateosi">
<emphasis id="B96886038714FFF3FD4FFDCE6628A647" box="[699,815,583,608]" italics="true" pageId="18" pageNumber="387">L. mateosi</emphasis>
</taxonomicName>
has 12 ciliate and 15 smooth chaetae. Other differences among the four species are presented in 
<tableCitation id="C69E6FAA8714FFF3FDADFDE367AAA6A3" box="[601,685,618,644]" captionStart="TABLE" captionStartId="7.[152,244,872,896]" captionTargetBox="[163,1398,918,1952]" captionTargetPageId="7" captionText="TABLE. 1. Comparison among Lepidocyrtus (Fractocyrtus) species." httpUri="http://table.plazi.org/id/DF630A998701FFE6FF6CFCE1667AA7A7" pageId="18" pageNumber="387" tableUuid="DF630A998701FFE6FF6CFCE1667AA7A7">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>