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<document id="FE1852F5713152FA633C641FC61C5014" ID-DOI="10.11646/zootaxa.5123.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6399827" ID-ZooBank="036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1648673534065" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard &amp; Janssen, Ronald" docDate="2022" docId="039487D1FFB3FF99FFBAFE1D6AFBF96E" docLanguage="en" docName="zootaxa.5123.1.1.pdf" docOrigin="Zootaxa 5123 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5123.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Granulatocincta contorta Harzhauser &amp; Landau &amp; Janssen 2022, nov. comb." docType="treatment" docVersion="4" lastPageNumber="50" masterDocId="FFADFFA9FF82FFA8FF2DFFAB6872FFC0" masterDocTitle="The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera" masterLastPageNumber="172" masterPageNumber="1" pageNumber="50" updateTime="1709053659529" updateUser="karina">
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<taxonomicName id="4C3D4D44FFB3FF99FFBAFE1D6ACDFE10" ID-CoL="8YMMN" authority="(Svagrovsky, 1958)" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" baseAuthorityName="Svagrovsky" baseAuthorityYear="1958" box="[151,703,438,467]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="contorta" status="comb. nov.">
<emphasis id="B949EAD5FFB3FF99FFBAFE1D69B8FE11" bold="true" box="[151,458,438,465]" italics="true" pageId="49" pageNumber="50">Granulatocincta contorta</emphasis>
(
<bibRefCitation id="EFAC4B36FFB3FF99FEF7FE1C6AC5FE10" author="Svagrovsky, J." box="[474,695,438,467]" pageId="49" pageNumber="50" pagination="5 - 56" refId="ref145176" refString="Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56." type="journal article" year="1958">Švagrovský, 1958</bibRefCitation>
)
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<taxonomicNameLabel id="A27A57AEFFB3FF99FDEBFE1C6B3FFE11" box="[710,845,439,465]" pageId="49" pageNumber="50" rank="species">nov. comb.</taxonomicNameLabel>
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<subSubSection id="C327654CFFB3FF99FFBAFE7069B4FE35" box="[151,454,474,507]" pageId="49" pageNumber="50" type="description">
<paragraph id="8B8236C7FFB3FF99FFBAFE7069B4FE35" blockId="49.[151,845,438,507]" box="[151,454,474,507]" pageId="49" pageNumber="50">
<figureCitation id="13062A42FFB3FF99FFBAFE706888FE35" box="[151,250,475,501]" captionStart="FIGURE 17" captionStartId="52.[151,250,1723,1747]" captionTargetBox="[248,1339,181,1698]" captionTargetId="figure-26@52.[248,1339,181,1698]" captionTargetPageId="52" captionText="FIGURE 17 A1A3. Granulatocincta callim nov. sp., holotype, NHMW 1866/0001/0549, Wiesen (Austria). B1B3. Granulatocincta callim nov.sp.,paralectotype,NHMW 1846/0037/0351b, Wiesen (Austria).C1C3. Granulatocincta callim nov. sp., paralectotype, NHMW 1846/0037/0351, Wiesen (Austria). D1D2. Granulatocincta callim nov. sp., paralectotype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). E. Granulatocincta carasiensis (Jekelius, 1944), MNG P 4965, Soceni (Romania). F1F2. Granulatocincta contorta (Švagrovský, 1958), holotype, Z 3076, Kuzmice (Slovakia). G1G2. Granulatocincta contorta (Švagrovský, 1958), paratype, SNMB Z 3074, Kuzmice (Slovakia). H1H3. Granulatocincta pelliscrocodili nov. sp., holotype, RGM 776 907, Gödet Creek near Tilkikaya (Turkey). I1I2. Granulatocincta pelliscrocodili nov. sp., paratype, RGM 776 906, southwest bank of Gödet Creek near Tilkikaya (Turkey). J1J3. Granulatocincta nataliae (Hoernes &amp; Auinger, 1891), holotype, NHMW 1855/0035/0021, Belgrade/Rakovica (Serbia)." figureDoi="http://doi.org/10.5281/zenodo.6399871" httpUri="https://zenodo.org/record/6399871/files/figure.png" pageId="49" pageNumber="50">Figs 17F</figureCitation>
<figureCitation id="13062A42FFB3FF99FFD7FE46696DFE35" box="[250,287,475,507]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="49" pageNumber="50">
<subScript id="17B93482FFB3FF99FFD7FE466970FE3B" attach="left" box="[250,258,493,507]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
F
</figureCitation>
<figureCitation id="13062A42FFB3FF99FE0DFE46695AFE3B" box="[288,296,493,507]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="49" pageNumber="50">
<subScript id="17B93482FFB3FF99FE0DFE46695AFE3B" attach="right" box="[288,296,493,507]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
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, G
<figureCitation id="13062A42FFB3FF99FE67FE466907FE35" box="[330,373,475,507]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="49" pageNumber="50">
<subScript id="17B93482FFB3FF99FE67FE466920FE3B" attach="left" box="[330,338,493,507]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
G
</figureCitation>
<figureCitation id="13062A42FFB3FF99FE58FE46690FFE3B" box="[373,381,493,507]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="49" pageNumber="50">
<subScript id="17B93482FFB3FF99FE58FE46690FFE3B" attach="left" box="[373,381,493,507]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
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,
<figureCitation id="13062A42FFB3FF99FEA6FE7069DEFE35" box="[395,428,475,501]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. Clavatula diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. Clavatula mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. Clavatula bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. Clavatula pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. Clavatula romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. Clavatula interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. Clavatula hirmetzli Kovács &amp; Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. Clavatula irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. Clavatula sophiae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. Clavatula veronicae (Hoernes &amp;Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. Clavatula santhai Kovács &amp; Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. Clavatula camillae (Hoernes &amp;Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &amp;Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes &amp; Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes &amp; Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="49" pageNumber="50">3Z</figureCitation>
,
<figureCitation id="13062A42FFB3FF99FE95FE7169B4FE35" box="[440,454,474,501]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="49" pageNumber="50">5</figureCitation>
</paragraph>
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<paragraph id="8B8236C7FFB3FF99FFBAFD8B6BC0FDF7" blockId="49.[151,1029,543,631]" box="[151,946,543,570]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="AB2D11E9FFB3FF99FFBAFD8B6BC0FDF7" box="[151,946,543,570]" pageId="49" pageNumber="50">
<emphasis id="B949EAD5FFB3FF99FFBAFD8B68D6FDF7" box="[151,164,544,567]" italics="true" pageId="49" pageNumber="50">*</emphasis>
<taxonomicName id="4C3D4D44FFB3FF99FF87FDB46994FDF7" ID-CoL="028c732f-b28b-4fd1-bd11-21e34532025d" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[170,486,543,567]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="contorta" status="sp. nov." subGenus="Clavatula">
<emphasis id="B949EAD5FFB3FF99FF87FDB4697CFDF7" box="[170,270,543,567]" italics="true" pageId="49" pageNumber="50">Clavatula</emphasis>
(
<emphasis id="B949EAD5FFB3FF99FE30FDB469F3FDF7" box="[285,385,543,567]" italics="true" pageId="49" pageNumber="50">Clavatula</emphasis>
)
<emphasis id="B949EAD5FFB3FF99FEBDFD8B6994FDF7" box="[400,486,544,567]" italics="true" pageId="49" pageNumber="50">contorta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFFB3FF99FEC1FD8B6A6DFDFA" box="[492,543,544,570]" pageId="49" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
<treatmentCitation id="0A9C10D6FFB3FF99FD14FD8A6B66FDF7" author="Svagrovsky, J." box="[569,788,543,570]" page="15" pageId="49" pageNumber="50" year="1958">
<bibRefCitation id="EFAC4B36FFB3FF99FD14FD8A6B66FDF7" author="Svagrovsky, J." box="[569,788,543,570]" pageId="49" pageNumber="50" pagination="5 - 56" refId="ref145176" refString="Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56." type="journal article" year="1958">Švagrovský 1958: 15</bibRefCitation>
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, pl. 4, figs 37.
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</paragraph>
<paragraph id="8B8236C7FFB3FF99FFBAFD956BA2FD96" blockId="49.[151,1029,543,631]" box="[151,976,574,600]" pageId="49" pageNumber="50">
<taxonomicName id="4C3D4D44FFB3FF99FFBAFD956A22FD98" ID-CoL="028c732f-b28b-4fd1-bd11-21e34532025d" authority="Svagrovsky" authorityName="Svagrovsky" authorityYear="1958" box="[151,592,574,600]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="contorta" subGenus="Clavatula">
<emphasis id="B949EAD5FFB3FF99FFBAFD956889FD96" box="[151,251,574,598]" italics="true" pageId="49" pageNumber="50">Clavatula</emphasis>
(
<emphasis id="B949EAD5FFB3FF99FE27FD95691CFD96" box="[266,366,574,598]" italics="true" pageId="49" pageNumber="50">Clavatula</emphasis>
)
<emphasis id="B949EAD5FFB3FF99FE50FD9469A1FD96" box="[381,467,575,598]" italics="true" pageId="49" pageNumber="50">contorta</emphasis>
Švagrovský
</taxonomicName>
—Švagrovský 1959: 223, pl. 8, fig. 6.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFB3FF99FFBAFDF76C77FDB5" box="[151,1029,604,631]" pageId="49" pageNumber="50" type="description">
<paragraph id="8B8236C7FFB3FF99FFBAFDF76C77FDB5" blockId="49.[151,1029,543,631]" box="[151,1029,604,631]" pageId="49" pageNumber="50">
<taxonomicName id="4C3D4D44FFB3FF99FFBAFDF76A22FDB7" authority="Svagrovsky" authorityName="Svagrovsky" authorityYear="1958" box="[151,592,604,631]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="contorta" subGenus="Clavatula">
<emphasis id="B949EAD5FFB3FF99FFBAFDF76889FDB4" box="[151,251,604,628]" italics="true" pageId="49" pageNumber="50">Clavatula</emphasis>
(
<emphasis id="B949EAD5FFB3FF99FE27FDF7691CFDB4" box="[266,366,604,628]" italics="true" pageId="49" pageNumber="50">Clavatula</emphasis>
)
<emphasis id="B949EAD5FFB3FF99FE50FDF669A1FDB4" box="[381,467,605,628]" italics="true" pageId="49" pageNumber="50">contorta</emphasis>
Švagrovský
</taxonomicName>
—Švagrovský 1960a: 102, pl. 13, figs 23.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFB3FF99FFBAFD086A5CFAE1" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8B8236C7FFB3FF99FFBAFD086C1CFCEA" blockId="49.[151,1437,674,1711]" pageId="49" pageNumber="50">
<emphasis id="B949EAD5FFB3FF99FFBAFD086931FD7C" bold="true" box="[151,323,674,701]" pageId="49" pageNumber="50">Type material.</emphasis>
<typeStatus id="54868865FFB3FF99FE65FD0869C7FD7D" box="[328,437,675,701]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
: Z 3076, Kuzmice (
<collectingCountry id="F32A7657FFB3FF99FDA6FD086A81FD7E" box="[651,755,675,702]" name="Slovakia" pageId="49" pageNumber="50">Slovakia</collectingCountry>
), SL:
<quantity id="4CC59B22FFB3FF99FC1CFD096BE4FD7D" box="[817,918,674,701]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="49" pageNumber="50" unit="mm" value="15.3">15.3 mm</quantity>
,
<collectionCode id="ED2CAE02FFB3FF99FC8DFD086BA3FD7D" box="[928,977,675,701]" country="Germany" name="Museum Donaueschingen" pageId="49" pageNumber="50">MD</collectionCode>
:
<quantity id="4CC59B22FFB3FF99FCF7FD086C42FD7D" box="[986,1072,675,701]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="49" pageNumber="50" unit="mm" value="6.9">6.9 mm</quantity>
, illustrated in
<bibRefCitation id="EFAC4B36FFB3FF99FBE7FD0F6DE5FD7D" author="Svagrovsky, J." box="[1226,1431,674,703]" pageId="49" pageNumber="50" pagination="5 - 56" refId="ref145176" refString="Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56." type="journal article" year="1958">Švagrovský (1958</bibRefCitation>
, pl. 4, fig. 5), stored in the Slovak National Museum (Bratislava,
<collectingCountry id="F32A7657FFB3FF99FC52FD6C6B97FD22" box="[895,997,711,738]" name="Slovakia" pageId="49" pageNumber="50">Slovakia</collectingCountry>
), figs 17F
<subScript id="17B93482FFB3FF99FB74FD726C13FD27" attach="left" box="[1113,1121,729,743]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
F
<subScript id="17B93482FFB3FF99FB53FD726CF4FD27" attach="left" box="[1150,1158,729,743]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
, 3Z.
<typeStatus id="54868865FFB3FF99FBEBFD6C6D45FD21" box="[1222,1335,711,737]" pageId="49" pageNumber="50" type="paratype">Paratypes</typeStatus>
: Z 3074, illustrated in
<bibRefCitation id="EFAC4B36FFB3FF99FE02FD476A72FCC5" author="Svagrovsky, J." box="[303,512,746,775]" pageId="49" pageNumber="50" pagination="5 - 56" refId="ref145176" refString="Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56." type="journal article" year="1958">Švagrovský (1958</bibRefCitation>
, pl. 4, figs, 3), figs 16G
<subScript id="17B93482FFB3FF99FC3BFD566B6CFCCB" attach="left" box="[790,798,765,779]" fontSize="6" pageId="49" pageNumber="50">1</subScript>
G
<subScript id="17B93482FFB3FF99FC6CFD566B3BFCCB" attach="left" box="[833,841,765,779]" fontSize="6" pageId="49" pageNumber="50">2</subScript>
; Z 3075, Z 3077, Z 3078, illustrated in
<bibRefCitation id="EFAC4B36FFB3FF99FA35FD4768A8FCE9" author="Svagrovsky, J." pageId="49" pageNumber="50" pagination="5 - 56" refId="ref145176" refString="Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56." type="journal article" year="1958">Švagrovský (1958</bibRefCitation>
, pl. 4, figs, 4, 6, 7), stored in the Slovak National Museum (Bratislava,
<collectingCountry id="F32A7657FFB3FF99FCD6FCA46C10FCEA" box="[1019,1122,783,810]" name="Slovakia" pageId="49" pageNumber="50">Slovakia</collectingCountry>
).
</paragraph>
<paragraph id="8B8236C7FFB3FF99FFEAFC996A5CFAE1" blockId="49.[151,1437,674,1711]" pageId="49" pageNumber="50">
<emphasis id="B949EAD5FFB3FF99FFEAFC9969CEFC8C" bold="true" box="[199,444,818,844]" pageId="49" pageNumber="50">Revised description.</emphasis>
Shell small, solid, moderately broad buccinoid with conical spire; apical angle ~43°. Protoconch paucispiral of one whorl. Teleoconch of up to nine whorls. Early teleoconch whorls flat-sided, bearing weakly opisthocline axial ribs with tripartite sculpture; two beaded adsutural spiral cords, separated by narrower beaded central cord. Subsutural cord becoming coarsely tuberculose, central and suprasutural band with smaller tubercles, further finely granular secondary spirals intercalated. Suture narrowly impressed, undulating. Last whorl 65% of total height, subcylindrical to weakly ovoid; subsutural collar broad, poorly delimited, weakly tuberculose. Subsutural ramp weakly concave, narrow, with beaded cord running at mid-ramp. Shoulder poorly defined by beaded cord, weakly convex below, weakly angular at peribasal cord. Base weakly constricted. Siphonal fasciole moderately swollen, twisted. Peribasal and perifasciolar cords weakly tuberculose, the tubercles aligned axially to form indistinct axial riblets, with numerous secondary cords of irregular strength intercalated over entire whorl. Aperture wide, ovate. Outer lip not thickened, smooth within. Anal sinus wide, deep, symmetrically U-shaped. Apex coinciding with row of beads mid-ramp. Siphonal canal moderately short, shallowly notched at tip. Columella weakly excavated in upper third, straight below, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming broad callus rim.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFB3FF99FFEAFA816BB8F9A6" pageId="49" pageNumber="50" type="discussion">
<paragraph id="8B8236C7FFB3FF99FFEAFA816C51FA4D" blockId="49.[151,1437,674,1711]" pageId="49" pageNumber="50">
<emphasis id="B949EAD5FFB3FF99FFEAFA81693EFA84" bold="true" box="[199,332,1322,1348]" pageId="49" pageNumber="50">Discussion.</emphasis>
<taxonomicName id="4C3D4D44FFB3FF99FE7BFA806B13FA85" authority="(Svagrovsky, 1958)" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" baseAuthorityName="Svagrovsky" baseAuthorityYear="1958" box="[342,865,1322,1351]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="contorta">
<emphasis id="B949EAD5FFB3FF99FE7BFA806A0AFA84" box="[342,632,1323,1349]" italics="true" pageId="49" pageNumber="50">Granulatocincta contorta</emphasis>
(
<bibRefCitation id="EFAC4B36FFB3FF99FDA4FA876B2BFA85" author="Svagrovsky, J." box="[649,857,1322,1351]" pageId="49" pageNumber="50" pagination="5 - 56" refId="ref145176" refString="Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56." type="journal article" year="1958">Švagrovský, 1958</bibRefCitation>
)
</taxonomicName>
is slightly reminiscent of
<taxonomicName id="4C3D4D44FFB3FF99FBBEFA806DEEFA85" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1171,1436,1323,1349]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="callim" status="sp. nov.">
<emphasis id="B949EAD5FFB3FF99FBBEFA806DEEFA85" box="[1171,1436,1323,1349]" italics="true" pageId="49" pageNumber="50">Granulatocincta callim</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFFB3FF99FFBAFAE4689FFAA9" box="[151,237,1359,1385]" pageId="49" pageNumber="50" rank="species">nov. sp.</taxonomicNameLabel>
The general mode of sculpture is similar in the two species, but
<taxonomicName id="4C3D4D44FFB3FF99FCEDFAE46C5AFAA9" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[960,1064,1359,1385]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="callim">
<emphasis id="B949EAD5FFB3FF99FCEDFAE46C5AFAA9" box="[960,1064,1359,1385]" italics="true" pageId="49" pageNumber="50">G. callim</emphasis>
</taxonomicName>
differs in its more subcylindrical last whorl and the sculpture of
<taxonomicName id="4C3D4D44FFB3FF99FEDFFAD86A28FA4D" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[498,602,1395,1421]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="callim">
<emphasis id="B949EAD5FFB3FF99FEDFFAD86A28FA4D" box="[498,602,1395,1421]" italics="true" pageId="49" pageNumber="50">G. callim</emphasis>
</taxonomicName>
is distinctly more crowded and stronger.
</paragraph>
<paragraph id="8B8236C7FFB3FF99FFEAFA3C6CAFF981" blockId="49.[151,1437,674,1711]" pageId="49" pageNumber="50">
The Tortonian, Proto-Mediterranean
<taxonomicName id="4C3D4D44FFB3FF99FD43FA3C6C29FA71" authority="(Michelotti, 1847)" baseAuthorityName="Michelotti" baseAuthorityYear="1847" box="[622,1115,1431,1457]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="sotterii">
<emphasis id="B949EAD5FFB3FF99FD43FA3C6BF2FA70" box="[622,896,1431,1457]" italics="true" pageId="49" pageNumber="50">Granulatocincta sotterii</emphasis>
(
<bibRefCitation id="EFAC4B36FFB3FF99FCBFFA3C6C21FA71" author="Michelotti, G." box="[914,1107,1431,1457]" pageId="49" pageNumber="50" pagination="1 - 408" refId="ref138690" refString="Michelotti, G. (1847) Description des fossiles des terrains miocenes de l'Italie septentrionale. Natuurkundige Verhandelingen van de Hollandsche Maatschappij der Wetenschappen te Haarlem, Series 2, 3 (2), 1 - 408. [https: // books. google. be / books? id = aQVqpL 58 HrsC &amp; ots = rhEPJ _ C 6 hy &amp; dq = Michelotti % 2 C % 20 G. % 091847 % 09 Description % 20 des % 20 fossiles % 20 de s % 20 terrains % 20 mioc % C 3 % A 8 nes % 20 de % 20 l'Italie % 20 septentrionale. &amp; lr &amp; pg = PP 5 # v = onepage &amp; q = Michelotti, % 20 G. % 091847 % 09 Description % 20 des % 2]" type="journal article" year="1847">Michelotti, 1847</bibRefCitation>
)
</taxonomicName>
is similar, but differs in its more crowded sculpture, much more prominent beads and more convex last whorl (see
<bibRefCitation id="EFAC4B36FFB3FF99FB51FA106D00FA15" author="Montanaro, E." box="[1148,1394,1467,1493]" pageId="49" pageNumber="50" pagination="115 - 191" refId="ref139628" refString="Montanaro, E. (1937) Studi monografici sulla malacologia miocenica modenese. Parte I. I mollusci tortoniani di Montegibbio. Palaeontographia Italica, New Series 7, 37, 115 - 191." type="journal article" year="1937">Montanaro 1937: 117</bibRefCitation>
, pl. 5, figs 218;
<bibRefCitation id="EFAC4B36FFB3FF99FE06FA746A4CFA39" author="Ruggieri, C. &amp; Davoli, F." box="[299,574,1503,1529]" pageId="49" pageNumber="50" pagination="41 - 85" refId="ref142914" refString="Ruggieri, C. &amp; Davoli, F. (1984) Malacofauna di Casa Nova Calisese (Sogliano, Forli), Palaeontographia Italica, 73, 41 - 85." type="journal article" year="1984">Ruggieri &amp; Davoli 1984</bibRefCitation>
, pl. 4, figs 1, 6, 1314, 1819,
<bibRefCitation id="EFAC4B36FFB3FF99FCB4FA746C55FA39" author="Davoli, F." box="[921,1063,1503,1529]" pageId="49" pageNumber="50" pagination="27 - 109" refId="ref131183" refString="Davoli, F. (1990) La collezione di &quot; Fossili Miocenici di Sogliano &quot; di Ludovico Foresti: revisione ed illustrazione. Atti della Societa dei Naturalisti e Matematici di Modena, 121, 27 - 109." type="journal article" year="1990">Davoli 1990</bibRefCitation>
, pl. 8, figs 28).
<taxonomicName id="4C3D4D44FFB3FF99FBCEFA74694AF9DD" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="pelliscrocodili" status="sp. nov.">
<emphasis id="B949EAD5FFB3FF99FBCEFA74694AF9DD" italics="true" pageId="49" pageNumber="50">Granulatocincta pelliscrocodili</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFFB3FF99FE12F9A869E5F9DD" box="[319,407,1539,1565]" pageId="49" pageNumber="50" rank="species">nov. sp.</taxonomicNameLabel>
, from the Serravallian of
<collectingCountry id="F32A7657FFB3FF99FD95F9A86B7BF9DD" box="[696,777,1539,1565]" name="Turkey" pageId="49" pageNumber="50">Turkey</collectingCountry>
, might be a related species, but differs from
<taxonomicName id="4C3D4D44FFB3FF99FBD3F9A86D0DF9DC" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" baseAuthorityName="Svagrovsky" baseAuthorityYear="1958" box="[1278,1407,1539,1565]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="contorta">
<emphasis id="B949EAD5FFB3FF99FBD3F9A86D0DF9DC" box="[1278,1407,1539,1565]" italics="true" pageId="49" pageNumber="50">G. contorta</emphasis>
</taxonomicName>
in its stout shape, the even weaker beads and the delicate spiral sculpture, which overruns the beads.
</paragraph>
<paragraph id="8B8236C7FFB3FF99FFEAF9E16BB8F9A6" blockId="49.[151,1437,674,1711]" box="[199,970,1610,1639]" pageId="49" pageNumber="50">
<emphasis id="B949EAD5FFB3FF99FFEAF9E169DAF9A4" bold="true" box="[199,424,1610,1636]" pageId="49" pageNumber="50">Paleoenvironment.</emphasis>
Inner neritic environments (
<bibRefCitation id="EFAC4B36FFB3FF99FDCBF9E76BCCF9A6" author="Svagrovsky, J." box="[742,958,1611,1639]" pageId="49" pageNumber="50" pagination="1 - 216" refId="ref145266" refString="Svagrovsky, J. (1960 b) Biostratigrafia a fauna makkysov vrchneho Tortonu vychodneho upatia Slanskych hor. Geologicke Prace, 57, 1 - 216." type="journal article" year="1960">Švagrovský 1960b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFB3FF99FFEAF9C56AFBF96E" pageId="49" pageNumber="50" type="distribution">
<paragraph id="8B8236C7FFB3FF99FFEAF9C56AFBF96E" blockId="49.[151,1437,674,1711]" pageId="49" pageNumber="50">
<emphasis id="B949EAD5FFB3FF99FFEAF9C56A0FF949" bold="true" box="[199,637,1646,1673]" pageId="49" pageNumber="50">Distribution in Central Paratethys.</emphasis>
Late Badenian (middle Miocene):
<emphasis id="B949EAD5FFB3FF99FB0AF9C46D50F948" box="[1063,1314,1647,1673]" italics="true" pageId="49" pageNumber="50">East Slovakian Basin</emphasis>
: Kuzmice (
<collectingCountry id="F32A7657FFB3FF99FFB2F9386976F96E" box="[159,260,1683,1710]" name="Slovakia" pageId="49" pageNumber="50">Slovakia</collectingCountry>
) (
<bibRefCitation id="EFAC4B36FFB3FF99FE37F93F6990F96D" author="Svagrovsky, J." box="[282,482,1682,1711]" pageId="49" pageNumber="50" pagination="5 - 56" refId="ref145176" refString="Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56." type="journal article" year="1958">Švagrovský 1958</bibRefCitation>
, 1959, 1960a).
</paragraph>
</subSubSection>
</treatment>
</document>