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<document id="C6D5B69DCD55E813D6AF22B2EA9C777A" ID-CLB-Dataset="7950" ID-DOI="10.11646/zootaxa.4878.3.3" ID-GBIF-Dataset="b9c711ba-146d-447b-ac2d-e1a0fcce0053" ID-ISSN="1175-5326" ID-Zenodo-Dep="4425178" ID-ZooBank="B660B3B2-7D4A-44C4-92C9-4BE24A1EA950" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610040207257" checkinUser="plazi" docAuthor="Sirenko, Boris I." docDate="2020" docId="039487CEF016FFA5FF5DFAD1FB3DF87E" docLanguage="en" docName="zootaxa.4878.3.3.pdf" docOrigin="Zootaxa 4878 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptochiton longigranum Sirenko 2020, n. sp." docType="treatment" docVersion="8" lastPageNumber="491" masterDocId="FFADFFB6F003FFBDFFCAFF86FFD6FF91" masterDocTitle="New shelf and bathyal Philippine leptochitons (Mollusca: Polyplacophora Lepidopleurida)" masterLastPageNumber="500" masterPageNumber="467" pageNumber="488" updateTime="1698923802988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="DC5099E2941FA8E6E7305B0DCEB46D00">
<mods:title id="32788E4B57769C2C3A04757865025FCD">New shelf and bathyal Philippine leptochitons (Mollusca: Polyplacophora Lepidopleurida)</mods:title>
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<mods:namePart id="1E0781F5686212EA884E78938C515A9D">Sirenko, Boris I.</mods:namePart>
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<mods:title id="A45EECC88C6FDF93D8BF109873EFBEE0">Zootaxa</mods:title>
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<mods:date id="39DA0C8D5CE089E2F3814DED1CF9EC1B">2020</mods:date>
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<mods:number id="7573579F337BF879133B8647E17147A3">2020-11-16</mods:number>
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<mods:identifier id="D11684D9020237F88DB2FEA1D7FA3418" type="DOI">10.11646/zootaxa.4878.3.3</mods:identifier>
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<treatment id="039487CEF016FFA5FF5DFAD1FB3DF87E" ID-DOI="http://doi.org/10.5281/zenodo.4426190" ID-GBIF-Taxon="180324281" ID-Zenodo-Dep="4426190" LSID="urn:lsid:plazi:treatment:039487CEF016FFA5FF5DFAD1FB3DF87E" httpUri="http://treatment.plazi.org/id/039487CEF016FFA5FF5DFAD1FB3DF87E" lastPageId="24" lastPageNumber="491" pageId="21" pageNumber="488">
<subSubSection id="C3276553F016FFA8FF5DFAD1FDC1FAE0" box="[151,535,1367,1394]" pageId="21" pageNumber="488" type="nomenclature">
<paragraph id="8B8236D8F016FFA8FF5DFAD1FDC1FAE0" blockId="21.[151,535,1367,1429]" box="[151,535,1367,1394]" pageId="21" pageNumber="488">
<heading id="D0CA81B4F016FFA8FF5DFAD1FDC1FAE0" bold="true" box="[151,535,1367,1394]" fontSize="11" level="1" pageId="21" pageNumber="488" reason="1">
<emphasis id="B949EACAF016FFA8FF5DFAD1FDC1FAE0" bold="true" box="[151,535,1367,1394]" pageId="21" pageNumber="488">
<taxonomicName id="4C3D4D5BF016FFA8FF5DFAD1FE18FAE3" ID-CoL="473dd3bb-f78c-4054-8c5e-1bfbf3df0b81" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[151,462,1367,1394]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="21" pageNumber="488" phylum="Mollusca" rank="species" species="longigranum" status="sp. nov.">
<emphasis id="B949EACAF016FFA8FF5DFAD1FE18FAE3" bold="true" box="[151,462,1367,1394]" italics="true" pageId="21" pageNumber="488">Leptochiton longigranum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57B1F016FFA8FE1FFAD1FDC1FAE0" box="[469,535,1367,1393]" pageId="21" pageNumber="488" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3276553F016FFA8FF5DFAFDFEF8FA04" box="[151,302,1403,1429]" pageId="21" pageNumber="488" type="description">
<paragraph id="8B8236D8F016FFA8FF5DFAFDFEF8FA04" blockId="21.[151,535,1367,1429]" box="[151,302,1403,1429]" pageId="21" pageNumber="488">
(
<figureCitation id="13062A5DF016FFA8FF55FAFDFEF0FA04" box="[159,294,1403,1429]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStartId-0="22.[151,250,1880,1904]" captionStartId-1="23.[151,250,1271,1295]" captionStartId-2="23.[151,250,1978,2002]" captionStartId-3="24.[151,250,1322,1346]" captionTargetBox-0="[218,1373,343,1842]" captionTargetBox-1="[182,1405,188,1240]" captionTargetBox-2="[181,1407,1395,1951]" captionTargetBox-3="[173,1415,189,1288]" captionTargetId-0="figure-147@22.[212,1381,335,1847]" captionTargetId-1="figure-16@23.[176,1411,181,1247]" captionTargetId-2="figure-58@23.[177,1411,1391,1954]" captionTargetId-3="figure-21@24.[164,1423,181,1299]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="23" captionTargetPageId-3="24" captionText-0="FIGURE 24. Leptochiton longigranum n. sp., Philippines, holotype (MNHN-IM-2013-67032), BL 3.5 mm: A. Valve I, dorsal view; B. Valve II, dorsal view; C. Valve IV, dorsal view; D. Valve V, ventral view; E. Valve II, detail of tegmentum in central area; F. Head valve, detail of tegmentum; G. Valve IV, posterior margin with protuberances." captionText-1="FIGURE 25. Leptochiton longigranum n. sp., Philippines, holotype (MNHN-IM-2013-67032), BL 3.5 mm:A. Sutural needles and scales; B. Dorsal scales; C. Dorsal scales and ventral scale with four ribs; D. Smooth ventral and ribbed marginal and dorsal scales." captionText-2="FIGURE 26. Leptochiton longigranum n. sp., Philippines, holotype (MNHN-IM-2013-67032), BL 3.5 mm: A. Radula; B. Central, first lateral, major lateral and spatulate uncinal teeth." captionText-3="FIGURE 27. Leptochiton longigranum n. sp., Philippines, holotype (MNHN-IM-2013-67032), BL 3.5 mm: A. Sutural needle; B. Dorsal scale; C. Marginal needle; D. marginal scale; E. Ventral scale near margin; F. Ventral scale from middle hyponotum; G. Head of major lateral tooth of radula; H. Central and first lateral teeth of radula; I. Aesthete group. Scale bar 100 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4425230" figureDoi-1="http://doi.org/10.5281/zenodo.4425232" figureDoi-2="http://doi.org/10.5281/zenodo.4425236" figureDoi-3="http://doi.org/10.5281/zenodo.4425238" httpUri-0="https://zenodo.org/record/4425230/files/figure.png" httpUri-1="https://zenodo.org/record/4425232/files/figure.png" httpUri-2="https://zenodo.org/record/4425236/files/figure.png" httpUri-3="https://zenodo.org/record/4425238/files/figure.png" pageId="21" pageNumber="488">Figs. 24–27</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3276553F016FFA8FF0DFA45FA89F9D8" pageId="21" pageNumber="488" type="materials_examined">
<paragraph id="8B8236D8F016FFA8FF0DFA45FEAFF9B4" blockId="21.[151,1437,1475,2042]" pageId="21" pageNumber="488">
<emphasis id="B949EACAF016FFA8FF0DFA45FEA3FA4C" bold="true" box="[199,373,1475,1501]" pageId="21" pageNumber="488">Type material.</emphasis>
<materialsCitation id="3B553C85F016FFA8FEB7FA45FEAFF9B4" ID-GBIF-Occurrence="3044017304" collectionCode="II, IV, V" pageId="21" pageNumber="488" specimenCount="1" typeStatus="holotype">
<typeStatus id="5486887AF016FFA8FEB7FA45FE0BFA4C" box="[381,477,1475,1501]" pageId="21" pageNumber="488" type="holotype">Holotype</typeStatus>
(MNHN-IM-2013-67032), a damaged specimen (tail valve was missing), now disarticulated, consisting of valves I,
<collectionCode id="ED2CAE1DF016FFA8FEB3FA6EFE5DF993" box="[377,395,1512,1538]" pageId="21" pageNumber="488">II</collectionCode>
,
<collectionCode id="ED2CAE1DF016FFA8FE5FFA6EFE79F993" box="[405,431,1512,1538]" pageId="21" pageNumber="488">IV</collectionCode>
,
<collectionCode id="ED2CAE1DF016FFA8FE7DFA6EFE1FF993" box="[439,457,1512,1538]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="21" pageNumber="488" type="Museum">V</collectionCode>
, part of perinotum and radula, mount of part of perinotum and radula mounted on SEM-stub and vial with other valves.
</materialsCitation>
</paragraph>
<paragraph id="8B8236D8F016FFA8FF0DF9A9FA89F9D8" blockId="21.[151,1437,1475,2042]" box="[199,1375,1583,1610]" pageId="21" pageNumber="488">
<materialsCitation id="3B553C85F016FFA8FF0DF9A9FA8DF9D8" ID-GBIF-Occurrence="3044017311" box="[199,1371,1583,1610]" collectingDate="2004-02-07" collectorName="Balicasag Id." country="Philippines" elevation="85" latitude="9.52" location="Philippines" longLatPrecision="129" longitude="123.67834" pageId="21" pageNumber="488" specimenCount="1" typeStatus="holotype">
<emphasis id="B949EACAF016FFA8FF0DF9A9FE81F9D8" bold="true" box="[199,343,1583,1609]" pageId="21" pageNumber="488">
<typeStatus id="5486887AF016FFA8FF0DF9A9FF2BF9D8" box="[199,253,1583,1609]" pageId="21" pageNumber="488">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32A7648F016FFA8FE96F9A9FE07F9D8" box="[348,465,1583,1609]" name="Philippines" pageId="21" pageNumber="488">Philippines</collectingCountry>
,
<collectorName id="26C8530EF016FFA8FE11F9A9FDB7F9D8" box="[475,609,1583,1609]" pageId="21" pageNumber="488">Balicasag Id.</collectorName>
,
<geoCoordinate id="EE09501FF016FFA8FDA1F9A9FD1AF9D8" box="[619,716,1583,1609]" degrees="9" direction="north" minutes="31.2" orientation="latitude" pageId="21" pageNumber="488" precision="92" value="9.52">9°31.2’N</geoCoordinate>
,
<geoCoordinate id="EE09501FF016FFA8FD1CF9A9FC9BF9D8" box="[726,845,1583,1609]" degrees="123" direction="east" minutes="40.7" orientation="longitude" pageId="21" pageNumber="488" precision="92" value="123.67834">123°40.7’E</geoCoordinate>
,
<quantity id="4CC59B3DF016FFA8FC92F9A9FC65F9DB" box="[856,947,1583,1610]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="21" pageNumber="488" unit="m" value="85.0" valueMax="90.0" valueMin="80.0">
<elevation id="0010D1EBF016FFA8FC92F9A9FC65F9DB" box="[856,947,1583,1610]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="21" pageNumber="488" unit="m" value="85.0" valueMax="90.0" valueMin="80.0">80–90 m</elevation>
</quantity>
(PANGLAO 2004, stn L42,
<date id="FF831018F016FFA8FB16F9A9FA83F9D8" box="[1244,1365,1583,1609]" pageId="21" pageNumber="488" value="2004-02-07">
<collectingDate id="EFC7E9F0F016FFA8FB16F9A9FA83F9D8" box="[1244,1365,1583,1609]" pageId="21" pageNumber="488" value="2004-02-07">02.07.2004</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F016FFA8FF0DF9D5FDC2F900" pageId="21" pageNumber="488" type="etymology">
<paragraph id="8B8236D8F016FFA8FF0DF9D5FDC2F900" blockId="21.[151,1437,1475,2042]" pageId="21" pageNumber="488">
<emphasis id="B949EACAF016FFA8FF0DF9D5FEE9F9FC" bold="true" box="[199,319,1619,1645]" pageId="21" pageNumber="488">Etymology</emphasis>
. From the Latin longus, long, and granum, granule, referring to the several long granules in posterior margin of intermediate valves in this species.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F016FFA8FF0DF91DFDE4F924" box="[199,562,1691,1717]" pageId="21" pageNumber="488" type="materials_examined">
<paragraph id="8B8236D8F016FFA8FF0DF91DFDE4F924" blockId="21.[151,1437,1475,2042]" box="[199,562,1691,1717]" pageId="21" pageNumber="488">
<emphasis id="B949EACAF016FFA8FF0DF91DFE49F924" bold="true" box="[199,415,1691,1717]" pageId="21" pageNumber="488">Material examined.</emphasis>
The
<typeStatus id="5486887AF016FFA8FE1BF91DFDF8F924" box="[465,558,1691,1717]" pageId="21" pageNumber="488" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F016FFA8FF0DF939FD48F948" box="[199,670,1727,1754]" pageId="21" pageNumber="488" type="distribution">
<paragraph id="8B8236D8F016FFA8FF0DF939FD48F948" blockId="21.[151,1437,1475,2042]" box="[199,670,1727,1754]" pageId="21" pageNumber="488">
<emphasis id="B949EACAF016FFA8FF0DF939FE9AF948" bold="true" box="[199,332,1727,1753]" pageId="21" pageNumber="488">Distribution</emphasis>
. Known only from the
<typeStatus id="5486887AF016FFA8FDF6F939FD4FF948" box="[572,665,1727,1753]" pageId="21" pageNumber="488" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F016FFA8FF0DF965FABCF8FB" pageId="21" pageNumber="488" type="diagnosis">
<paragraph id="8B8236D8F016FFA8FF0DF965FABCF8FB" blockId="21.[151,1437,1475,2042]" pageId="21" pageNumber="488">
<emphasis id="B949EACAF016FFA8FF0DF965FEE1F96C" bold="true" box="[199,311,1763,1789]" pageId="21" pageNumber="488">Diagnosis.</emphasis>
Small, shell rounded. Tegmentum sculptured with well raised, round or oval granules arranged in quincunx, at the posterior margin of the lateral areas, there are six or more protuberances standing up about four times the height of the larger granules. Dorsal scales pointed, with 4–6 longitudinal ribs. Central tooth of radula short with sharp base, major lateral tooth with bidentate cusp, inner denticle much smaller than outer one. Four gills on each side extending from valve VII to near anus.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F016FFA5FF0DF8F5FC85F8AA" lastPageId="24" lastPageNumber="491" pageId="21" pageNumber="488" type="description">
<paragraph id="8B8236D8F016FFA8FF0DF8F5FEB9F820" blockId="21.[151,1437,1475,2042]" pageId="21" pageNumber="488">
<emphasis id="B949EACAF016FFA8FF0DF8F5FE87F81C" bold="true" box="[199,337,1907,1933]" pageId="21" pageNumber="488">Description.</emphasis>
Small, BL
<quantity id="4CC59B3DF016FFA8FE00F8F5FDCAF81F" box="[458,540,1907,1934]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="488" unit="mm" value="2.5">2.5 mm</quantity>
(probably adult), oval, valves only little elevated, rounded, not beaked. Color of tegmentum dirty white.
</paragraph>
<paragraph id="8B8236D8F016FFA8FF0DF83DFE3BF868" blockId="21.[151,1437,1475,2042]" pageId="21" pageNumber="488">Head valve semicircular. Intermediate valves rectangular, anterior and posterior margin nearly straight, side margins rounded, lateral areas not raised.</paragraph>
<paragraph id="8B8236D8F015FFABFF0DFF11FCC8FE8C" blockId="22.[151,1437,151,285]" pageId="22" pageNumber="489">Tegmentum sculptured with well raised, round or oval granules arranged in quincunx, granules of central area flattened, those of other areas convex.At the posterior margin of the lateral areas, there are six or more protuberances standing up about four times the height of the larger granules. Each granule with one megalaesthete and seven to nine micraesthetes, all pores arranged at the periphery of flattened top of granule.</paragraph>
<caption id="DF426650F015FFABFF5DF8DEFC2EF829" ID-DOI="http://doi.org/10.5281/zenodo.4425230" ID-Zenodo-Dep="4425230" httpUri="https://zenodo.org/record/4425230/files/figure.png" pageId="22" pageNumber="489" startId="22.[151,250,1880,1904]" targetBox="[218,1373,343,1842]" targetPageId="22">
<paragraph id="8B8236D8F015FFABFF5DF8DEFC2EF829" blockId="22.[151,1437,1880,1977]" pageId="22" pageNumber="489">
<emphasis id="B949EACAF015FFABFF5DF8DEFEF7F8E1" bold="true" box="[151,289,1880,1904]" pageId="22" pageNumber="489">FIGURE 24.</emphasis>
<taxonomicName id="4C3D4D5BF015FFABFEE2F8DEFDFFF8E1" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[296,553,1880,1904]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="22" pageNumber="489" phylum="Mollusca" rank="species" species="longigranum" status="sp. nov.">
<emphasis id="B949EACAF015FFABFEE2F8DEFDFFF8E1" box="[296,553,1880,1904]" italics="true" pageId="22" pageNumber="489">Leptochiton longigranum</emphasis>
</taxonomicName>
<emphasis id="B949EACAF015FFABFDFAF8DFFDBFF8E1" bold="true" box="[560,617,1881,1905]" pageId="22" pageNumber="489">
<taxonomicNameLabel id="A27A57B1F015FFABFDFAF8DFFDBFF8E1" box="[560,617,1881,1905]" pageId="22" pageNumber="489" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Philippines, holotype (MNHN-IM-2013-67032), BL 3.5 mm: A. Valve I, dorsal view; B. Valve II, dorsal view; C. Valve IV, dorsal view; D. Valve V, ventral view; E. Valve II, detail of tegmentum in central area; F. Head valve, detail of tegmentum; G. Valve IV, posterior margin with protuberances.
</paragraph>
</caption>
<caption id="DF426650F014FFAAFF5DFB71FF0DFAC6" ID-DOI="http://doi.org/10.5281/zenodo.4425232" ID-Zenodo-Dep="4425232" httpUri="https://zenodo.org/record/4425232/files/figure.png" pageId="23" pageNumber="490" startId="23.[151,250,1271,1295]" targetBox="[182,1405,188,1240]" targetPageId="23">
<paragraph id="8B8236D8F014FFAAFF5DFB71FF0DFAC6" blockId="23.[151,1436,1271,1367]" pageId="23" pageNumber="490">
<emphasis id="B949EACAF014FFAAFF5DFB71FEF6FA9E" bold="true" box="[151,288,1271,1295]" pageId="23" pageNumber="490">FIGURE 25.</emphasis>
<taxonomicName id="4C3D4D5BF014FFAAFEEFFB71FDF1FA9E" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[293,551,1271,1295]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="23" pageNumber="490" phylum="Mollusca" rank="species" species="longigranum" status="sp. nov.">
<emphasis id="B949EACAF014FFAAFEEFFB71FDF1FA9E" box="[293,551,1271,1295]" italics="true" pageId="23" pageNumber="490">Leptochiton longigranum</emphasis>
</taxonomicName>
<emphasis id="B949EACAF014FFAAFDE7FB71FDB0FA9E" bold="true" box="[557,614,1271,1295]" pageId="23" pageNumber="490">
<taxonomicNameLabel id="A27A57B1F014FFAAFDE7FB71FDB0FA9E" box="[557,614,1271,1295]" pageId="23" pageNumber="490" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Philippines, holotype (MNHN-IM-2013-67032), BL 3.5 mm: A. Sutural needles and scales; B. Dorsal scales; C. Dorsal scales and ventral scale with four ribs; D. Smooth ventral and ribbed marginal and dorsal scales.
</paragraph>
</caption>
<caption id="DF426650F014FFAAFF5DF83CFCD3F866" ID-DOI="http://doi.org/10.5281/zenodo.4425236" ID-Zenodo-Dep="4425236" httpUri="https://zenodo.org/record/4425236/files/figure.png" pageId="23" pageNumber="490" startId="23.[151,250,1978,2002]" targetBox="[181,1407,1395,1951]" targetPageId="23">
<paragraph id="8B8236D8F014FFAAFF5DF83CFCD3F866" blockId="23.[151,1436,1978,2039]" pageId="23" pageNumber="490">
<emphasis id="B949EACAF014FFAAFF5DF83CFEF5F843" bold="true" box="[151,291,1978,2002]" pageId="23" pageNumber="490">FIGURE 26.</emphasis>
<taxonomicName id="4C3D4D5BF014FFAAFEE6F83DFDE4F842" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[300,562,1979,2003]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="23" pageNumber="490" phylum="Mollusca" rank="species" species="longigranum" status="sp. nov.">
<emphasis id="B949EACAF014FFAAFEE6F83DFDE4F842" box="[300,562,1979,2003]" italics="true" pageId="23" pageNumber="490">Leptochiton longigranum</emphasis>
</taxonomicName>
<emphasis id="B949EACAF014FFAAFDF1F83DFDA1F842" bold="true" box="[571,631,1979,2003]" pageId="23" pageNumber="490">
<taxonomicNameLabel id="A27A57B1F014FFAAFDF1F83DFDA1F842" box="[571,631,1979,2003]" pageId="23" pageNumber="490" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Philippines, holotype (MNHN-IM-2013-67032), BL 3.5 mm: A. Radula; B. Central, first lateral, major lateral and spatulate uncinal teeth.
</paragraph>
</caption>
<caption id="DF426650F01BFFA5FF5DFAACFA8CFA1A" ID-DOI="http://doi.org/10.5281/zenodo.4425238" ID-Zenodo-Dep="4425238" httpUri="https://zenodo.org/record/4425238/files/figure.png" pageId="24" pageNumber="491" startId="24.[151,250,1322,1346]" targetBox="[173,1415,189,1288]" targetPageId="24">
<paragraph id="8B8236D8F01BFFA5FF5DFAACFA8CFA1A" blockId="24.[151,1436,1322,1419]" pageId="24" pageNumber="491">
<emphasis id="B949EACAF01BFFA5FF5DFAACFEF6FAD3" bold="true" box="[151,288,1322,1346]" pageId="24" pageNumber="491">FIGURE 27.</emphasis>
<taxonomicName id="4C3D4D5BF01BFFA5FEECFAADFDFEFAD2" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[294,552,1323,1347]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="24" pageNumber="491" phylum="Mollusca" rank="species" species="longigranum" status="sp. nov.">
<emphasis id="B949EACAF01BFFA5FEECFAADFDFEFAD2" box="[294,552,1323,1347]" italics="true" pageId="24" pageNumber="491">Leptochiton longigranum</emphasis>
</taxonomicName>
<emphasis id="B949EACAF01BFFA5FDE4FAADFDB1FAD2" bold="true" box="[558,615,1323,1347]" pageId="24" pageNumber="491">
<taxonomicNameLabel id="A27A57B1F01BFFA5FDE4FAADFDB1FAD2" box="[558,615,1323,1347]" pageId="24" pageNumber="491" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Philippines, holotype (MNHN-IM-2013-67032), BL 3.5 mm: A. Sutural needle; B. Dorsal scale; C. Marginal needle; D. marginal scale; E. Ventral scale near margin; F. Ventral scale from middle hyponotum; G. Head of major lateral tooth of radula; H. Central and first lateral teeth of radula; I. Aesthete group. Scale bar 100 µm.
</paragraph>
</caption>
<paragraph id="8B8236D8F01BFFA5FF0DFA3FFBB6FA42" blockId="24.[151,1437,1465,2031]" box="[199,1120,1465,1491]" pageId="24" pageNumber="491">Articulamentum white, weakly developed, apophyses broadly rounded, triangular.</paragraph>
<paragraph id="8B8236D8F01BFFA5FF0DFA5BFBC6F93A" blockId="24.[151,1437,1465,2031]" pageId="24" pageNumber="491">
Girdle very narrow, densely covered with small, imbricating, conical scales (50 x
<quantity id="4CC59B3DF01BFFA5FB9CFA5BFB74FA66" box="[1110,1186,1501,1527]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="24" pageNumber="491" unit="mm" value="26.0">26 µm</quantity>
), sometimes truncated at the top, always marked with 4–6 sharp ribs, which are twice less than interstices; intersegmental needles very long and smooth (350 x
<quantity id="4CC59B3DF01BFFA5FE67F9A3FE2DF9AE" box="[429,507,1573,1599]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="491" unit="mm" value="17.0">17 µm</quantity>
). Margin armed with two different kinds of elements: 1) few long smooth needles similar to the intersegmental ones (350 x
<quantity id="4CC59B3DF01BFFA5FDA4F9CFFD6DF9F2" box="[622,699,1609,1635]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="491" unit="mm" value="17.0">17 µm</quantity>
), 2) dorsally finely ribbed (6–7 ribs) sharply pointed scales (80 x 18 μm) Ventral side near margin with one row of small (50–60 x 20 μm), oval, distally obtusely pointed scales with four short ribs in upper half; other ventral scales smooth, pointed (46 x
<quantity id="4CC59B3DF01BFFA5FC7DF917FBD2F93A" box="[951,1028,1681,1708]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="491" unit="mm" value="15.0">15 µm</quantity>
).
</paragraph>
<paragraph id="8B8236D8F01BFFA5FF0DF933FE4EF886" blockId="24.[151,1437,1465,2031]" pageId="24" pageNumber="491">
Radula
<quantity id="4CC59B3DF01BFFA5FED7F933FEA5F95E" box="[285,371,1717,1743]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="491" unit="mm" value="0.7">0.7 mm</quantity>
long with 25 transverse rows of mature teeth. Central tooth short, rectangular, with sharp base, first lateral tooth short, triangular, without a blade, major lateral tooth with bidentate cusp, inner denticle much smaller than outer one.
</paragraph>
<paragraph id="8B8236D8F01BFFA5FF0DF8A7FC85F8AA" blockId="24.[151,1437,1465,2031]" box="[199,851,1825,1851]" pageId="24" pageNumber="491">There are 4 gills per side arranged from valve VII to anus.</paragraph>
</subSubSection>
<subSubSection id="C3276553F01BFFA5FF0DF8C3FB3DF87E" pageId="24" pageNumber="491" type="discussion">
<paragraph id="8B8236D8F01BFFA5FF0DF8C3FB3DF87E" blockId="24.[151,1437,1465,2031]" pageId="24" pageNumber="491">
<emphasis id="B949EACAF01BFFA5FF0DF8C3FEEDF8CE" bold="true" box="[199,315,1861,1887]" pageId="24" pageNumber="491">Remarks.</emphasis>
The new species can be easily recognized by the protuberances at the posterior margin of the intermediate valves.
<taxonomicName id="4C3D4D5BF01BFFA5FE9BF8EFFDA7F812" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[337,625,1897,1923]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="24" pageNumber="491" phylum="Mollusca" rank="species" species="longigranum" status="sp. nov.">
<emphasis id="B949EACAF01BFFA5FE9BF8EFFDA7F812" box="[337,625,1897,1923]" italics="true" pageId="24" pageNumber="491">Leptochiton longigranum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57B1F01BFFA5FDB3F8EFFD6EF812" box="[633,696,1897,1923]" pageId="24" pageNumber="491" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="4C3D4D5BF01BFFA5FCF0F8EFFB6DF812" authority="(Ashby, 1929)" baseAuthorityName="Ashby" baseAuthorityYear="1929" box="[826,1211,1897,1923]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="24" pageNumber="491" phylum="Mollusca" rank="species" species="finlayi">
<emphasis id="B949EACAF01BFFA5FCF0F8EFFBC7F812" box="[826,1041,1897,1923]" italics="true" pageId="24" pageNumber="491">Leptochiton finlayi</emphasis>
(
<bibRefCitation id="EFAC4B29F01BFFA5FBEBF8EFFB65F812" author="Ashby, E." box="[1057,1203,1897,1923]" pageId="24" pageNumber="491" pagination="366 - 369" refId="ref10535" refString="Ashby, E. (1929) New Zealand fossil Polyplacophora (Chitons). Transactions and Proceedings of the Royal Society of New Zealand, 60, 366 - 369, pl. 32." type="journal article" year="1929">Ashby, 1929</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="F32A7648F01BFFA5FACEF8EFFA4AF812" box="[1284,1436,1897,1923]" name="New Zealand" pageId="24" pageNumber="491">New Zealand</collectingCountry>
which also has protuberances at the posterior margin of the lateral areas, but differs from by having granules arranged in quincunx (vs. granules in head valve, central areas of intermediate valves and antemucronal area of tail valve arranged in longitudinal rows in
<taxonomicName id="4C3D4D5BF01BFFA5FD83F850FD79F87E" baseAuthorityName="Ashby" baseAuthorityYear="1929" box="[585,687,2005,2031]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="24" pageNumber="491" phylum="Mollusca" rank="species" species="finlayi">
<emphasis id="B949EACAF01BFFA5FD83F850FD79F87E" box="[585,687,2005,2031]" italics="true" pageId="24" pageNumber="491">L. finlayi</emphasis>
</taxonomicName>
) and four gills per side (vs. nine gills in
<taxonomicName id="4C3D4D5BF01BFFA5FBBFF850FB0DF87E" baseAuthorityName="Ashby" baseAuthorityYear="1929" box="[1141,1243,2005,2031]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="24" pageNumber="491" phylum="Mollusca" rank="species" species="finlayi">
<emphasis id="B949EACAF01BFFA5FBBFF850FB0DF87E" box="[1141,1243,2005,2031]" italics="true" pageId="24" pageNumber="491">L. finlayi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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