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<document id="0E74BBAB18AFB65F638448254AB50BA1" ID-CLB-Dataset="298438" ID-DOI="10.11646/zootaxa.5468.2.2" ID-GBIF-Dataset="7a281190-31de-47e2-8ba9-85a1c6924faf" ID-ISSN="1175-5326" ID-Zenodo-Dep="11615558" ID-ZooBank="0FF33E52-AEAB-4207-935C-4E11F551A96D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1718185369582" checkinUser="plazi" docAuthor="Muilwijk, Jan, Kataev, Boris M. &amp; Wrase, David W." docDate="2024" docId="039487C7FFB1BB66FF66FA14989DF8C7" docLanguage="en" docName="zootaxa.5468.2.2.pdf" docOrigin="Zootaxa 5468 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5468.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyrothorax (Lyrothorax) Chaudoir 1838" docType="key" docVersion="6" lastPageNumber="273" masterDocId="FFADFFBFFFA3BB74FFF1FFDB9D01FFD3" masterDocTitle="Review of the subgenus Lyrothorax of the genus Pterostichus (Coleoptera: Carabidae) in the Western Palaearctic, with descriptions of two new species" masterLastPageNumber="274" masterPageNumber="255" pageNumber="273" updateTime="1718287281004" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="50FCF11D285C0E9E99A0FDF0277C873A">Review of the subgenus Lyrothorax of the genus Pterostichus (Coleoptera: Carabidae) in the Western Palaearctic, with descriptions of two new species</mods:title>
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<mods:namePart id="5D899CBD5C14530677086F59D9D0065C">Muilwijk, Jan</mods:namePart>
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<mods:namePart id="DF80F22E31DA9BCE648BA07229A6F19F">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="EB83D34EB3BD332202A2AE138A975033">Wrase, David W.</mods:namePart>
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<treatment id="039487C7FFB1BB66FF66FA14989DF8C7" ID-DOI="http://doi.org/10.5281/zenodo.11621089" ID-GBIF-Taxon="232746602" ID-Zenodo-Dep="11621089" LSID="urn:lsid:plazi:treatment:039487C7FFB1BB66FF66FA14989DF8C7" httpUri="http://treatment.plazi.org/id/039487C7FFB1BB66FF66FA14989DF8C7" lastPageNumber="273" pageId="18" pageNumber="273">
<subSubSection id="C327655AFFB1BB66FF66FA149987FA3A" box="[151,1158,1486,1513]" pageId="18" pageNumber="273" type="nomenclature">
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<heading id="D0CA81BDFFB1BB66FF66FA149987FA3A" bold="true" box="[151,1158,1486,1513]" fontSize="11" level="1" pageId="18" pageNumber="273" reason="1">
<emphasis id="B949EAC3FFB1BB66FF66FA149987FA3A" bold="true" box="[151,1158,1486,1513]" pageId="18" pageNumber="273">
Key to Western Palaearctic
<taxonomicName id="4C3D4D52FFB1BB66FE02FA159F7AFA3A" authority="Chaudoir, 1838" authorityName="Chaudoir" authorityYear="1838" box="[499,635,1486,1513]" class="Insecta" family="Carabidae" genus="Lyrothorax" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="274" phylum="Arthropoda" rank="subGenus" subGenus="Lyrothorax">
<emphasis id="B949EAC3FFB1BB66FE02FA159F7AFA3A" bold="true" box="[499,635,1486,1513]" italics="true" pageId="18" pageNumber="273">Lyrothorax</emphasis>
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species based on median lobe of aedeagus
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<keyLead id="30CC95E4FFB1BB66FF66F9CF989DF9B7" pageId="18" pageNumber="273">
1 In left lateral view angulate margin of preputial field of the apical orifice (pft) very large, triangular; terminal lamella in lateral view oblique, with a large tooth (tltl) (
<figureCitation id="13062A54FFB1BB66FDA6F9EA9F9EF994" box="[599,671,1585,1607]" captionStart="FIGURE 6" captionStartId="8.[151,250,1669,1693]" captionTargetBox="[153,1433,1100,1643]" captionTargetId="figure-97@8.[151,1435,1097,1645]" captionTargetPageId="8" captionText="FIGURE 6. Median lobe in left lateral view, (a) P. caspius (Ménétriés, 1832) lectotype, (b) P. fritzei sp. n. from Khelard, (c) P. vagus sp. n. from Ramsar Curti, preputial field tooth (pft), terminal lamella (tl), tooth on terminal lamella (tltl). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11615582" httpUri="https://zenodo.org/record/11615582/files/figure.png" pageId="18" pageNumber="273">Fig. 6b</figureCitation>
), in dorsal view short (WL-tl: 2.99–3.41), apical margin incised medially (
<figureCitation id="13062A54FFB1BB66FA84F9EA9DFAF9B6" captionStart="FIGURE 7" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[160,1430,625,1209]" captionTargetId="figure-232@11.[151,1436,618,1211]" captionTargetPageId="11" captionText="FIGURE 7. Median lobe in dorsal view, (a) P. caspius (Ménétriés, 1832) from Astara, Istisu, (b) P. fritzei sp. n. from Kheyroud Forest, (c) P. vagus sp. n., from different localities, c1 from Ramsar Curti, c2 from “Saramsakli”, Māzandarān, c3 from Now Shahr, preputial field tooth (pft), right pigmented band (rpb), terminal lamella (tl). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11615586" httpUri="https://zenodo.org/record/11615586/files/figure.png" pageId="18" pageNumber="273">Fig. 7b</figureCitation>
).....................................................................................
<taxonomicName id="4C3D4D52FFB1BB66FAE0F9949861F9B7" authority="Muilwijk &amp; Kataev &amp; Wrase, 2024" authorityName="Muilwijk &amp; Kataev &amp; Wrase" authorityYear="2024" box="[1297,1376,1615,1636]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="273" phylum="Arthropoda" rank="species" species="fritzei" status="sp. n.">
<emphasis id="B949EAC3FFB1BB66FAE0F9949861F9B7" box="[1297,1376,1615,1636]" italics="true" pageId="18" pageNumber="273">P. fritzei</emphasis>
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<taxonomicNameLabel id="A27A57B8FFB1BB66FA97F994989DF9B7" box="[1382,1436,1615,1636]" pageId="18" pageNumber="273" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8236D1FFB1BB66FF66F9B7989DF952" blockId="18.[151,1437,1556,1813]" box="[151,1436,1644,1666]" pageId="18" pageNumber="273">
<keyLead id="30CC95E4FFB1BB66FF66F9B7989DF952" box="[151,1436,1644,1666]" pageId="18" pageNumber="273">
- In left lateral view angulate margin of preputial field of the apical orifice (pft) short (
<figureCitation id="13062A54FFB1BB66FBFEF9B7996CF952" box="[1039,1133,1644,1666]" captionStart="FIGURE 6" captionStartId="8.[151,250,1669,1693]" captionTargetBox="[153,1433,1100,1643]" captionTargetId="figure-97@8.[151,1435,1097,1645]" captionTargetPageId="8" captionText="FIGURE 6. Median lobe in left lateral view, (a) P. caspius (Ménétriés, 1832) lectotype, (b) P. fritzei sp. n. from Khelard, (c) P. vagus sp. n. from Ramsar Curti, preputial field tooth (pft), terminal lamella (tl), tooth on terminal lamella (tltl). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11615582" httpUri="https://zenodo.org/record/11615582/files/figure.png" pageId="18" pageNumber="273">Figs 6a, c</figureCitation>
)....................... 2
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<keyStep id="30C92E74FFB1BB66FF66F952989DF8C7" pageId="18" pageNumber="273">
<paragraph id="8B8236D1FFB1BB66FF66F952989DF90A" blockId="18.[151,1437,1556,1813]" pageId="18" pageNumber="273">
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2 Terminal lamella in lateral view straight or slightly bent, without a tooth along the left lateral margin (
<figureCitation id="13062A54FFB1BB66FB4DF9529803F94C" box="[1212,1282,1673,1695]" captionStart="FIGURE 6" captionStartId="8.[151,250,1669,1693]" captionTargetBox="[153,1433,1100,1643]" captionTargetId="figure-97@8.[151,1435,1097,1645]" captionTargetPageId="8" captionText="FIGURE 6. Median lobe in left lateral view, (a) P. caspius (Ménétriés, 1832) lectotype, (b) P. fritzei sp. n. from Khelard, (c) P. vagus sp. n. from Ramsar Curti, preputial field tooth (pft), terminal lamella (tl), tooth on terminal lamella (tltl). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11615582" httpUri="https://zenodo.org/record/11615582/files/figure.png" pageId="18" pageNumber="273">Fig. 6c</figureCitation>
), in dorsal view mostly long; but strongly variable (WL-tl: 1.89–3.06) (
<figureCitation id="13062A54FFB1BB66FD18F97C9E33F96F" box="[745,818,1703,1725]" captionStart="FIGURE 7" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[160,1430,625,1209]" captionTargetId="figure-232@11.[151,1436,618,1211]" captionTargetPageId="11" captionText="FIGURE 7. Median lobe in dorsal view, (a) P. caspius (Ménétriés, 1832) from Astara, Istisu, (b) P. fritzei sp. n. from Kheyroud Forest, (c) P. vagus sp. n., from different localities, c1 from Ramsar Curti, c2 from “Saramsakli”, Māzandarān, c3 from Now Shahr, preputial field tooth (pft), right pigmented band (rpb), terminal lamella (tl). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11615586" httpUri="https://zenodo.org/record/11615586/files/figure.png" pageId="18" pageNumber="273">Figs 7c</figureCitation>
<figureCitation id="13062A54FFB1BB66FCC3F97C9E57F96F" box="[818,854,1703,1724]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1516,1540]" captionStartId-1="6.[151,250,1236,1260]" captionStartId-2="7.[151,250,1143,1167]" captionTargetBox-0="[159,1428,641,1484]" captionTargetBox-1="[154,1435,182,1213]" captionTargetBox-2="[153,1433,183,1117]" captionTargetId-0="figure-249@3.[151,1436,633,1493]" captionTargetId-1="figure-29@6.[151,1435,181,1213]" captionTargetId-2="figure-17@7.[151,1435,181,1119]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 1. Habitus of lectotype and paralectotypes of P. caspius (Ménétriés, 1832), with labels including the handwritten bottom label by Ménétriés. Scale bar =10 mm." captionText-1="FIGURE 2. Habitus in dorsal view and pronotum increased, (a) P. caspius (Ménétriés, 1832) from Talysh, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n., from different localities, c1 holotype, c2 from Bylnice, Czech Republic. Scale bar =10 mm." captionText-2="FIGURE 3. Habitus in ventral view, (a) P. caspius (Ménétriés, 1832) from Talysh Mts., Hyrkan, (b) P. fritzei sp. n. holotype, (c) P. vagus sp. n. holotype. Scale bar = 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11615564" figureDoi-1="http://doi.org/10.5281/zenodo.11615568" figureDoi-2="http://doi.org/10.5281/zenodo.11615576" httpUri-0="https://zenodo.org/record/11615564/files/figure.png" httpUri-1="https://zenodo.org/record/11615568/files/figure.png" httpUri-2="https://zenodo.org/record/11615576/files/figure.png" pageId="18" pageNumber="273">1–3</figureCitation>
), Czech specimens from
<collectingRegion id="49F9F833FFB1BB66FBB5F97C9998F96E" box="[1092,1177,1703,1725]" country="Czech Republic" name="Jihomoravsky" pageId="18" pageNumber="273">Moravia</collectingRegion>
: 1.67–2.04, from
<collectingCountry id="F32A7641FFB1BB66FACEF97C9898F96E" box="[1343,1433,1703,1725]" name="Czech Republic" pageId="18" pageNumber="273">Bohemia</collectingCountry>
: 2.37–2.76), mostly apical margin medially incised (
<figureCitation id="13062A54FFB1BB66FD20F91F9E14F90A" box="[721,789,1732,1754]" captionStart="FIGURE 7" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[160,1430,625,1209]" captionTargetId="figure-232@11.[151,1436,618,1211]" captionTargetPageId="11" captionText="FIGURE 7. Median lobe in dorsal view, (a) P. caspius (Ménétriés, 1832) from Astara, Istisu, (b) P. fritzei sp. n. from Kheyroud Forest, (c) P. vagus sp. n., from different localities, c1 from Ramsar Curti, c2 from “Saramsakli”, Māzandarān, c3 from Now Shahr, preputial field tooth (pft), right pigmented band (rpb), terminal lamella (tl). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11615586" httpUri="https://zenodo.org/record/11615586/files/figure.png" pageId="18" pageNumber="273">Fig. 7c</figureCitation>
).........................................
<taxonomicName id="4C3D4D52FFB1BB66FAFFF91E9861F909" authority="Muilwijk &amp; Kataev &amp; Wrase, 2024" authorityName="Muilwijk &amp; Kataev &amp; Wrase" authorityYear="2024" box="[1294,1376,1732,1754]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="273" phylum="Arthropoda" rank="species" species="vagus" status="sp. n.">
<emphasis id="B949EAC3FFB1BB66FAFFF91E981CF909" box="[1294,1309,1733,1754]" italics="true" pageId="18" pageNumber="273">P</emphasis>
.
<emphasis id="B949EAC3FFB1BB66FAD9F91E9861F909" box="[1320,1376,1733,1754]" italics="true" pageId="18" pageNumber="273">vagus</emphasis>
</taxonomicName>
<emphasis id="B949EAC3FFB1BB66FA97F91F989DF90A" bold="true" box="[1382,1436,1732,1753]" pageId="18" pageNumber="273">
<taxonomicNameLabel id="A27A57B8FFB1BB66FA97F91F989DF90A" box="[1382,1436,1732,1753]" pageId="18" pageNumber="273" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8236D1FFB1BB66FF66F939989DF8C7" blockId="18.[151,1437,1556,1813]" pageId="18" pageNumber="273">
<keyLead id="30CC95E4FFB1BB66FF66F939989DF8C7" pageId="18" pageNumber="273">
- Terminal lamella in lateral view oblique, with a small tooth (
<figureCitation id="13062A54FFB1BB66FCDEF93A9E74F924" box="[815,885,1761,1783]" captionStart="FIGURE 6" captionStartId="8.[151,250,1669,1693]" captionTargetBox="[153,1433,1100,1643]" captionTargetId="figure-97@8.[151,1435,1097,1645]" captionTargetPageId="8" captionText="FIGURE 6. Median lobe in left lateral view, (a) P. caspius (Ménétriés, 1832) lectotype, (b) P. fritzei sp. n. from Khelard, (c) P. vagus sp. n. from Ramsar Curti, preputial field tooth (pft), terminal lamella (tl), tooth on terminal lamella (tltl). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11615582" httpUri="https://zenodo.org/record/11615582/files/figure.png" pageId="18" pageNumber="273">Fig. 6a</figureCitation>
), in dorsal view medium long (WL-tl: 2.38–2.60), apical margin straight, in some specimens from
<collectingRegion id="49F9F833FFB1BB66FD9EF9249FA5F8C6" box="[623,676,1791,1813]" country="Iran" name="Gilan" pageId="18" pageNumber="273">Gilan</collectingRegion>
rounded (
<figureCitation id="13062A54FFB1BB66FCF7F9249E4BF8C7" box="[774,842,1791,1813]" captionStart="FIGURE 7" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[160,1430,625,1209]" captionTargetId="figure-232@11.[151,1436,618,1211]" captionTargetPageId="11" captionText="FIGURE 7. Median lobe in dorsal view, (a) P. caspius (Ménétriés, 1832) from Astara, Istisu, (b) P. fritzei sp. n. from Kheyroud Forest, (c) P. vagus sp. n., from different localities, c1 from Ramsar Curti, c2 from “Saramsakli”, Māzandarān, c3 from Now Shahr, preputial field tooth (pft), right pigmented band (rpb), terminal lamella (tl). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11615586" httpUri="https://zenodo.org/record/11615586/files/figure.png" pageId="18" pageNumber="273">Fig. 7a</figureCitation>
)..................................
<taxonomicName id="4C3D4D52FFB1BB66FB04F924989DF8C7" ID-CoL="4Q3WG" authority="(Men.)" baseAuthorityName="Menetries" baseAuthorityYear="1832" box="[1269,1436,1791,1812]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="273" phylum="Arthropoda" rank="species" species="caspius">
<emphasis id="B949EAC3FFB1BB66FB04F9249855F8C7" box="[1269,1364,1791,1812]" italics="true" pageId="18" pageNumber="273">P. caspius</emphasis>
(Mén.)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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