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<mods:title id="FE9F7D0DF66ECE112AC5DBFCABB607CD">First Record Of Calcareous Green Algae (Dasycladales, Halimedaceae) From The Paleocene Chehel Kaman Formation Of North-Eastern Iran (Kopet-Dagh Basin)</mods:title>
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<mods:namePart id="89C438A1FF5F1AE1545A92F894D79B80">SCHLAGINTWEIT, FELIX</mods:namePart>
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<mods:namePart id="0771BFF0EDFC2E11DC665314F935D330">RASHIDI, KOOROSH</mods:namePart>
<mods:affiliation id="A0D3D8B4E487FA52F401DDFCA26B717A">2 Department of Geology, Yazd University, 89195 - 741 Yazd, Iran</mods:affiliation>
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<mods:namePart id="5B6A6FC570BFCC6339D750AF61779219">MOSAVINIA, ABDOLMAJID</mods:namePart>
<mods:affiliation id="1EB48BA8694F2B8A7A2A3452E6A931D1">Department of Geology, Payame Noor University, Tehran, Iran, Po Box 19395 - 3697</mods:affiliation>
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<treatment id="0394879D143BA31FFCAA389BF438FBA6" LSID="urn:lsid:plazi:treatment:0394879D143BA31FFCAA389BF438FBA6" httpUri="http://treatment.plazi.org/id/0394879D143BA31FFCAA389BF438FBA6" lastPageNumber="53" pageId="2" pageNumber="53">
<subSubSection id="C3276500143BA31FFCAA389BF4D3FF04" box="[816,1195,191,215]" pageId="2" pageNumber="53" type="nomenclature">
<paragraph id="8B82368B143BA31FFCAA389BF4D3FF04" blockId="2.[816,1195,191,248]" box="[816,1195,191,215]" pageId="2" pageNumber="53">
<taxonomicName id="4C3D4D08143BA31FFCAA389BF4D3FF04" authority="(Elliott, 1968)" baseAuthorityName="Elliott" baseAuthorityYear="1968" box="[816,1195,191,215]" class="Ulvophyceae" family="Dasycladaceae" genus="Uteria" isUncertain="true" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="species" species="merienda">
<emphasis id="B949EA99143BA31FFCAA389BF30DFF04" box="[816,885,192,215]" italics="true" pageId="2" pageNumber="53">Uteria</emphasis>
aff.
<emphasis id="B949EA99143BA31FFC3238E4F475FF04" box="[936,1037,191,215]" italics="true" pageId="2" pageNumber="53">merienda</emphasis>
(
<bibRefCitation id="EFAC4B7A143BA31FFB8138E4F4DAFF04" author="Elliott" box="[1051,1186,191,215]" firstAuthor="Elliott" pageId="2" pageNumber="53" pagination="1 - 111" refId="ref6430" refString="Elliott, G. F., 1968. Permian to Paleocene calcareous algae (Dasycladaceae) of the Middle-East. Bulletin of the British Museum, Natural History, Geology, supplement, 4: 1 - 111." type="journal article" year="1968">Elliott, 1968</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3276500143BA31FFCAA38BBF3DCFF2B" box="[816,932,224,248]" pageId="2" pageNumber="53" type="description">
<paragraph id="8B82368B143BA31FFCAA38BBF3DCFF2B" blockId="2.[816,1195,191,248]" box="[816,932,224,248]" pageId="2" pageNumber="53">
<figureCitation id="13062A0E143BA31FFCAA38BBF307FF2B" box="[816,895,224,248]" captionStart="Fig" captionStartId="1.[222,257,1967,1988]" captionTargetBox="[235,1353,1131,1929]" captionTargetId="figure-122@1.[235,1353,1131,1929]" captionTargetPageId="1" captionText="Fig. 2 Wackestone with numerous specimens of the calcareous benthic foraminifera Rahaghia khorassanica (Rahaghi) from the Chehel Kaman Formation, outcrops along the road Mashhad-Sarakhs. Note the verticil of Uteria aff. merienda (Elliott) in b and fragment of Neomeris in d with two partially preserved pedunculated fertile ampullae and one secondary lateral. Scale bars 0.5 mm in bd. Thin-sections Nf 8 (a, d), Nf 5 (b), Nf 13 (c)." figureDoi="http://doi.org/10.5281/zenodo.10520825" httpUri="https://zenodo.org/record/10520825/files/figure.png" pageId="2" pageNumber="53">Fig. 2b</figureCitation>
,
<figureCitation id="13062A0E143BA31FFC1038BBF3DCFF2B" box="[906,932,224,248]" captionStart="Fig" captionStartId="5.[151,186,1907,1928]" captionTargetBox="[151,771,1531,1881]" captionTargetId="figure-25@5.[151,771,1531,1881]" captionTargetPageId="5" captionText="Fig. 5 Thyrsoporella/Trinocladus, tangential section (c), Uteria aff. merienda (Elliott) (b), oblique longitudinal section from Paleocene carbonates of the Chehel Kaman Formation, northeastern Iran. Thin-sections: Nf 13 (a), NF 4 (b)." figureDoi="http://doi.org/10.5281/zenodo.10520831" httpUri="https://zenodo.org/record/10520831/files/figure.png" pageId="2" pageNumber="53">5b</figureCitation>
</paragraph>
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<subSubSection id="C3276500143BA31FFCAA3979F5F0FE6C" pageId="2" pageNumber="53" type="reference_group">
<paragraph id="8B82368B143BA31FFCAA3979F3B9FE88" blockId="2.[816,1437,290,447]" pageId="2" pageNumber="53">
*1968
<taxonomicName id="4C3D4D08143BA31FFCE53979F428FEE9" authority="SCHLAGINTWEIT &amp; RASHIDI &amp; MOSAVINIA, 2021" authorityName="SCHLAGINTWEIT &amp; RASHIDI &amp; MOSAVINIA" authorityYear="2021" box="[895,1104,290,314]" class="Ulvophyceae" family="Dasycladaceae" genus="Clypeina" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="species" species="merienda" status="sp. nov.">
<emphasis id="B949EA99143BA31FFCE53979F428FEE9" box="[895,1104,290,314]" italics="true" pageId="2" pageNumber="53">Clypeina merienda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57E2143BA31FFBC63978F4E2FEE9" box="[1116,1178,291,314]" pageId="2" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
Elliott, p. 35, pl. 4, figs. 2-3, 7-8.
</paragraph>
<paragraph id="8B82368B143BA31FFCAA393FF38DFE4E" blockId="2.[816,1437,290,447]" pageId="2" pageNumber="53">
1982
<taxonomicName id="4C3D4D08143BA31FFCF5393FF4FFFEAF" authority="Elliott" authorityName="Elliott" box="[879,1159,356,380]" class="Ulvophyceae" family="Dasycladaceae" genus="Clypeina" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="species" species="merienda">
<emphasis id="B949EA99143BA31FFCF5393FF444FEAF" box="[879,1084,356,380]" italics="true" pageId="2" pageNumber="53">Clypeina merienda</emphasis>
Elliott
</taxonomicName>
Deloffre and Génot, p. 51, pl. 3, figs. 6-7.
</paragraph>
<paragraph id="8B82368B143BA31FFCAA39FCF5F0FE6C" blockId="2.[816,1437,290,447]" box="[816,1416,423,447]" pageId="2" pageNumber="53">
1990
<taxonomicName id="4C3D4D08143BA31FFCF739F3F400FE6C" authority="(Elliott)" baseAuthorityName="Elliott" box="[877,1144,423,447]" class="Ulvophyceae" family="Dasycladaceae" genus="Uteria" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="species" species="merienda">
<emphasis id="B949EA99143BA31FFCF739F3F466FE6C" box="[877,1054,423,447]" italics="true" pageId="2" pageNumber="53">Uteria merienda</emphasis>
(Elliott)
</taxonomicName>
Radoičić, pl. 10, fig. 5.
</paragraph>
</subSubSection>
<subSubSection id="C3276500143BA31FFCAA39B2F438FBA6" pageId="2" pageNumber="53" type="discussion">
<paragraph id="8B82368B143BA31FFCAA39B2F438FBA6" blockId="2.[816,1437,489,1142]" pageId="2" pageNumber="53">
<emphasis id="B949EA99143BA31FFCAA39B2F3D9FDD2" bold="true" box="[816,929,489,513]" pageId="2" pageNumber="53">Remarks:</emphasis>
As the sections illustrated by
<bibRefCitation id="EFAC4B7A143BA31FFA9C39B2F5E4FDD2" author="Elliott" box="[1286,1436,489,513]" firstAuthor="Elliott" pageId="2" pageNumber="53" pagination="1 - 111" refId="ref6430" refString="Elliott, G. F., 1968. Permian to Paleocene calcareous algae (Dasycladaceae) of the Middle-East. Bulletin of the British Museum, Natural History, Geology, supplement, 4: 1 - 111." type="journal article" year="1968">Elliott (1968)</bibRefCitation>
from the Paleocene Sinjar Formation of
<collectingCountry id="F32A761B143BA31FFA9E3A50F557FDF1" box="[1284,1327,523,546]" name="Iraq" pageId="2" pageNumber="53">Iraq</collectingCountry>
were assigned to the genus
<taxonomicName id="4C3D4D08143BA31FFB883A70F4A4FD90" authority="Michelin" authorityName="Michelin" authorityYear="1844" box="[1042,1244,555,579]" class="Ulvophyceae" family="Dasycladaceae" genus="Clypeina" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="genus">
<emphasis id="B949EA99143BA31FFB883A70F40AFD90" box="[1042,1138,555,579]" italics="true" pageId="2" pageNumber="53">Clypeina</emphasis>
Michelin
</taxonomicName>
, they were interpreted as belonging to fertile laterals. “Reminiscent of
<taxonomicName id="4C3D4D08143BA31FFCAA3A36F3E8FD56" authorityName="Michelin" authorityYear="1844" box="[816,912,621,645]" class="Ulvophyceae" family="Dasycladaceae" genus="Clypeina" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="genus">
<emphasis id="B949EA99143BA31FFCAA3A36F3E8FD56" box="[816,912,621,645]" italics="true" pageId="2" pageNumber="53">Clypeina</emphasis>
</taxonomicName>
”, they were referred to the sterile laterals of the genus
<taxonomicName id="4C3D4D08143BA31FFCE73AD4F449FD75" authority="Michelin" authorityName="Michelin" authorityYear="1845" box="[893,1073,654,678]" class="Ulvophyceae" family="Dasycladaceae" genus="Uteria" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="genus">
<emphasis id="B949EA99143BA31FFCE73AD4F3BAFD75" box="[893,962,655,678]" italics="true" pageId="2" pageNumber="53">Uteria</emphasis>
Michelin
</taxonomicName>
showing
<specimenCount id="9D3BFD02143BA31FFB323AD4F56EFD74" box="[1192,1302,655,679]" count="2" pageId="2" pageNumber="53" type="generic" typeStatus="types">two types</specimenCount>
of laterals, sterile and fertile (
<bibRefCitation id="EFAC4B7A143BA31FFC673AF4F4DEFD14" author="Radoicic" box="[1021,1190,687,711]" firstAuthor="Radoicic" pageId="2" pageNumber="53" pagination="91 - 103" refId="ref7226" refString="Radoicic, R., 1990. Paleogene dasycladalean algae from the subsurface of the western Iraqi Desert. Bulletin de l'Academie Serbe des Sciences et des Arts (Sciences mathematiques et naturelles, Sciences naturelles), t. CII, 32: 91 - 103." type="journal article" year="1990">Radoičić, 1990</bibRefCitation>
;
<bibRefCitation id="EFAC4B7A143BA31FFB2F3AF4F502FD14" author="Dieni" box="[1205,1402,687,711]" etAl="et al." firstAuthor="Dieni" pageId="2" pageNumber="53" pagination="1 - 77" refId="ref6335" refString="Dieni, I., Massari, F., Radoicic, R., 1985. Palaeocene Dasycladalean algae from Orosei vicinity (Sardinia). Memorie di Scienze Geologiche, 38: 1 - 77." type="journal article" year="1985">Dieni et al., 1985</bibRefCitation>
, p. 26). This
<typeStatus id="54868829143BA31FFC3A3A8AF3B6FD3A" box="[928,974,721,745]" pageId="2" pageNumber="53">type</typeStatus>
of thallus structure is well discernible in the longitudinal section illustrated in
<figureCitation id="13062A0E143BA31FFB503AA9F523FCDA" box="[1226,1371,754,778]" captionStart="Fig" captionStartId="10.[216,251,1782,1803]" captionTargetBox="[216,1422,154,1740]" captionTargetId="figure-53@10.[216,1422,154,1740]" captionTargetPageId="10" captionText="Fig. 11 Selection of Paleocene Dasycladales from different parts of Iran: Eastern Iran-Sistan Suture Zone (a, c-f, i), Central Iran (g), and Southwestern Iran-Zagros Zone (b, h). a, Uteria merienda (Elliott), longitudinal section, thin-section BN 59. b, Rostroporella oviformis Segonzac, oblique section, thin-section Qs 59. c, Dissocladella savitriae Pia (above), longitudinal section, and Cymopolia aff. morelletorum Beckman (below), oblique section, thin-section 2pb 76. d, Triploporella apenninica Baretti, longitudinal-oblique section, thin-section 2pb 39. e, Cymopolia sp., longitudinal section of an article, thinsection 2pb 3. f, Clypeina socaensis Deloffre &amp; Radoičić, oblique section, thin-section 2pb 27. g, Orioporella sinensis (Wang), thin-section Ah 161. h, Thyrsoporella sp., thin-section Qs 7. i, Jodotella sloveniaensis Deloffre &amp; Radoičić, oblique longitudinal section, thin-section 2pb 40. Scale bars 1 mm (a-d, g) and 0.5 mm (e-f, h-i)." figureDoi="http://doi.org/10.5281/zenodo.10520843" httpUri="https://zenodo.org/record/10520843/files/figure.png" pageId="2" pageNumber="53">Figure 11a, a</figureCitation>
specimen from Eastern
<collectingCountry id="F32A761B143BA31FFB9E3B48F457FCF9" box="[1028,1071,787,810]" name="Iran" pageId="2" pageNumber="53">Iran</collectingCountry>
(Sistan Suture Zone). There may be one or two fertile whorls between the disc-like whorls of sterile laterals. The specimens from the Chehel Kaman Formation refer exclusively to fragments of sterile laterals.
<collectionCode id="ED2CAE4E143BA31FFCEB3BCCF3FCFC7C" box="[881,900,919,943]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="53" type="Herbarium">A</collectionCode>
comparative table of the different
<taxonomicName id="4C3D4D08143BA31FFB663BC3F5E0FC7C" box="[1276,1432,919,943]" pageId="2" pageNumber="53">
<emphasis id="B949EA99143BA31FFB663BC3F539FC7C" box="[1276,1345,920,943]" italics="true" pageId="2" pageNumber="53">Uteria</emphasis>
species
</taxonomicName>
, excluding
<taxonomicName id="4C3D4D08143BA31FFC383BE2F451FC03" box="[930,1065,952,976]" class="Ulvophyceae" family="Dasycladaceae" genus="Uteria" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="species" species="merienda">
<emphasis id="B949EA99143BA31FFC383BE2F451FC03" box="[930,1065,952,976]" italics="true" pageId="2" pageNumber="53">U. merienda</emphasis>
</taxonomicName>
, has been provided by
<bibRefCitation id="EFAC4B7A143BA31FFABD3BE3F3EEFC22" author="Deloffre, R. &amp; Fleury, J. - J. &amp; Fourcade, E. &amp; Michaud, F." pageId="2" pageNumber="53" pagination="3 - 15" refId="ref6122" refString="Deloffre, R., Fleury, J. - J., Fourcade, E., Michaud, F., 1989. Dasycladales (Algues Vertes) du Paleocene-Eocene inferieur du Mexique. Revue de Micropaleontologie, 32 (1): 3 - 15." type="journal article" year="1989">Deloffre et al. (1989</bibRefCitation>
, tab. 1). In fact, the new species described by
<bibRefCitation id="EFAC4B7A143BA31FFCAA3BA1F460FBC1" author="Deloffre, R. &amp; Fleury, J. - J. &amp; Fourcade, E. &amp; Michaud, F." box="[816,1048,1018,1042]" pageId="2" pageNumber="53" pagination="3 - 15" refId="ref6122" refString="Deloffre, R., Fleury, J. - J., Fourcade, E., Michaud, F., 1989. Dasycladales (Algues Vertes) du Paleocene-Eocene inferieur du Mexique. Revue de Micropaleontologie, 32 (1): 3 - 15." type="journal article" year="1989">Deloffre et al. (1989)</bibRefCitation>
as
<taxonomicName id="4C3D4D08143BA31FFBA53BA0F48BFBC1" box="[1087,1267,1019,1042]" class="Ulvophyceae" family="Dasycladaceae" genus="Uteria" kingdom="Plantae" order="Dasycladales" pageId="2" pageNumber="53" phylum="Chlorophyta" rank="species" species="mexicana">
<emphasis id="B949EA99143BA31FFBA53BA0F48BFBC1" box="[1087,1267,1019,1042]" italics="true" pageId="2" pageNumber="53">Uteria mexicana</emphasis>
</taxonomicName>
from the upper Paleocene can in our opinion not be differentiated distinctly from the Iraqi species and might be a possible junior synonym of the latter.
</paragraph>
</subSubSection>
</treatment>
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