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<document id="085AD18DBB556773C26B7BED418B98FD" ID-DOI="10.5281/zenodo.5334580" ID-ISSN="0374-1036" ID-Zenodo-Dep="5334580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630175634131" checkinUser="felipe" docAuthor="Fikáček, Martin, Delgado, Juan A. &amp; Gentili, Elio" docDate="2012" docId="0394878EFFA0FFC84276FAB1FD37840B" docLanguage="en" docName="ActaEntMusNatPra.2012.52.suppl.107-130.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Georissus (Neogeorissus) nemo Fikáček &amp; Delgado &amp; Gentili 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="116" masterDocId="FFADFFF6FFA5FFC14308FFCAFFAB8018" masterDocTitle="The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae)" masterLastPageNumber="130" masterPageNumber="107" pageNumber="112" updateTime="1699175334189" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9D93836DECF1F56A4980D55988BA703D">The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="722071E035E3F551C25AAFB880D13624">Fikáček, Martin</mods:namePart>
<mods:affiliation id="F45BDA3E63D705E6D89F30AA21CA0FF0">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="3494F8138C868F51CBE10A4101B17A07">Delgado, Juan A.</mods:namePart>
<mods:affiliation id="B7E8CD0C4839665BC7A348EA64F97734">Departamento de Zoología, Facultad de Biología, Universidad de Murcia, 30100, Murcia, Spain; e-mail: jdelgado @ um. es</mods:affiliation>
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<mods:namePart id="ED59750488D67451F977E5E693868747">Gentili, Elio</mods:namePart>
<mods:affiliation id="84CC06566722CC34223E20CA2ED1ED56">Via San Gottardo 37, I- 21030 Varese-Rasa, Italy; e-mail: elio. gentili. 32 @ alice. it</mods:affiliation>
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<mods:title id="4DD5018CEA34590B972006112FB3FAE6">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="0C259CF2FE5E9B4E366F65A486D837CD">2012</mods:date>
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<mods:identifier id="359B9F227091AD47E43C1F08E1876D0B" type="ISSN">0374-1036</mods:identifier>
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<treatment id="0394878EFFA0FFC84276FAB1FD37840B" ID-DOI="http://doi.org/10.5281/zenodo.5413319" ID-Zenodo-Dep="5413319" LSID="urn:lsid:plazi:treatment:0394878EFFA0FFC84276FAB1FD37840B" httpUri="http://treatment.plazi.org/id/0394878EFFA0FFC84276FAB1FD37840B" lastPageId="9" lastPageNumber="116" pageId="5" pageNumber="112">
<subSubSection id="C3276513FFA0FFC44276FAB1FC9E858C" box="[382,821,1403,1428]" pageId="5" pageNumber="112" type="nomenclature">
<paragraph id="8B823698FFA0FFC44276FAB1FC9E858C" blockId="5.[382,821,1403,1428]" box="[382,821,1403,1428]" pageId="5" pageNumber="112">
<heading id="D0CA81F4FFA0FFC44276FAB1FC9E858C" bold="true" box="[382,821,1403,1428]" centered="true" fontSize="8" level="2" pageId="5" pageNumber="112" reason="3">
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<taxonomicName id="4C3D4D1BFFA0FFC44276FAB1FD72858B" ID-CoL="3FS2Q" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[382,729,1403,1427]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov." subGenus="Neogeorissus">
<emphasis id="B949EA8AFFA0FFC44276FAB1FE41858B" bold="true" box="[382,490,1403,1427]" italics="true" pageId="5" pageNumber="112">Georissus</emphasis>
(
<emphasis id="B949EA8AFFA0FFC442F1FAB1FD27858B" bold="true" box="[505,652,1403,1427]" italics="true" pageId="5" pageNumber="112">Neogeorissus</emphasis>
)
<emphasis id="B949EA8AFFA0FFC44194FAB6FD72858B" bold="true" box="[668,729,1404,1427]" italics="true" pageId="5" pageNumber="112">nemo</emphasis>
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<taxonomicNameLabel id="A27A57F1FFA0FFC441D7FAB6FC9E858C" box="[735,821,1404,1428]" pageId="5" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA0FFC442F2FA50FD1C85B5" box="[506,695,1434,1453]" pageId="5" pageNumber="112" type="description">
<paragraph id="8B823698FFA0FFC442F2FA50FD1C85B5" blockId="5.[506,695,1434,1453]" box="[506,695,1434,1453]" pageId="5" pageNumber="112">
(
<figureCitation id="13062A1DFFA0FFC44108FA50FD9685B5" box="[512,573,1434,1453]" captionStart="Figs" captionStartId="4.[153,192,1033,1052]" captionTargetBox="[266,1016,226,1012]" captionTargetId="figure-178@4.[262,1016,219,1015]" captionTargetPageId="4" captionText="Figs. 78. Genitalia of Socotran endemic Georissus species. 7 G. maritimus sp. nov. (a, b: two different ventral views of the same aedeagus; c: lateral view); 8 G. nemo sp. nov., ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334586" httpUri="https://zenodo.org/record/5334586/files/figure.png" pageId="5" pageNumber="112">Figs. 8</figureCitation>
,
<figureCitation id="13062A1DFFA0FFC4414EFA50FDD985B5" box="[582,626,1434,1453]" captionStart="Figs" captionStartId="7.[113,152,1482,1501]" captionTargetBox="[113,1087,217,1402]" captionTargetId="figure-13@7.[113,1087,217,1402]" captionTargetPageId="7" captionText="Figs. 914. Georissus nemo sp. nov. (913 paratype; 14 holotype). 9 pronotum and elytra in dorsal view; 10 head, dorsal view; 1112 detail of pronotum (11 dorsal view; 12 dorsolateral view); 13 elytron, dorsolateral view; 14 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334588" httpUri="https://zenodo.org/record/5334588/files/figure.png" pageId="5" pageNumber="112">914</figureCitation>
,
<figureCitation id="13062A1DFFA0FFC44173FA50FD1985B5" box="[635,690,1434,1453]" captionStart="Figs" captionStartId="10.[151,190,1491,1510]" captionTargetBox="[153,1125,221,1476]" captionTargetId="figure-12@10.[147,1128,216,1478]" captionTargetPageId="10" captionText="Figs. 1522.Socotran endemic Georissus species and their habitats. 1516 G. maritimus sp. nov.(15 dorsal view; 16 lateral view). 1718 G. nemo sp. nov. (17 dorsal view; 18 lateral view). 1920 the sand tongue 3 km NE of Shuab, type locality of G. maritimus sp. nov. (19 general view of the habitat; 20 satellite view showing the position of the locality); 2122 the stream at Arhar, type locality of G. nemo sp. nov. (21 general view of the habitat; 22 satellite view showing the position of the locality)." figureDoi="http://doi.org/10.5281/zenodo.5334592" httpUri="https://zenodo.org/record/5334592/files/figure.png" pageId="5" pageNumber="112">1718</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA0FFC74379FA0EFE6B8141" lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112" type="materials_examined">
<paragraph id="8B823698FFA0FFC44379FA0EFCE385CF" blockId="5.[113,1089,1475,1575]" box="[113,840,1475,1495]" pageId="5" pageNumber="112">
<emphasis id="B949EA8AFFA0FFC44379FA0EFF4185CF" bold="true" box="[113,234,1475,1495]" pageId="5" pageNumber="112">
<typeStatus id="5486883AFFA0FFC44379FA0EFF3585CF" box="[113,158,1476,1495]" pageId="5" pageNumber="112">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32A7608FFA0FFC443E7FA0EFE8685CF" box="[239,301,1476,1495]" name="Yemen" pageId="5" pageNumber="112">Yemen</collectingCountry>
,
<collectingCountry id="F32A7608FFA0FFC4423DFA0EFE1985CF" box="[309,434,1476,1495]" name="Socotra Island" pageId="5" pageNumber="112">Socotra Island</collectingCountry>
, Hallah Arhar,
<geoCoordinate id="EE09505FFFA0FFC4413BFA0EFD2185CF" box="[563,650,1475,1495]" degrees="12" direction="north" minutes="33.0" orientation="latitude" pageId="5" pageNumber="112" precision="92" value="12.55">12°33.0N</geoCoordinate>
<geoCoordinate id="EE09505FFFA0FFC44198FA09FD4C85CF" box="[656,743,1475,1495]" degrees="54" direction="east" minutes="27.6" orientation="longitude" pageId="5" pageNumber="112" precision="92" value="54.46">54°27.6E</geoCoordinate>
,
<quantity id="4CC59B7DFFA0FFC441E7FA09FCB085CF" box="[751,795,1475,1495]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="112" unit="m" value="15.0">15 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8B823698FFA0FFC74379FA14FE6B8141" blockId="5.[113,1089,1475,1575]" lastBlockId="6.[151,1128,219,346]" lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112">
<emphasis id="B949EA8AFFA0FFC44379FA14FF5D85E9" bold="true" box="[113,246,1501,1521]" pageId="5" pageNumber="112">Type material.</emphasis>
<materialsCitation id="3B553CC5FFA0FFC443F3FA15FEFF8614" collectingDate="2010-11-11" collectionCode="NMPC" collectorName="J. Hajek'" country="Yemen" elevation="15" latitude="12.55" location="Hallah Arhar" longLatPrecision="129" longitude="54.46" pageId="5" pageNumber="112" specimenCount="1" typeStatus="holotype">
<typeStatus id="5486883AFFA0FFC443F3FA15FEE585E8" box="[251,334,1503,1522]" pageId="5" pageNumber="112" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED2CAE5DFFA0FFC4427CFA14FE1D85E9" box="[372,438,1502,1521]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="112">NMPC</collectionCode>
):
<collectingCountry id="F32A7608FFA0FFC442CFFA14FDB185E9" box="[455,538,1502,1521]" name="Yemen" pageId="5" pageNumber="112">YEMEN</collectingCountry>
: Socotra Isl. /
<location id="8EE26043FFA0FFC4419AFA14FCAA85E9" LSID="urn:lsid:plazi:treatment:0394878EFFA0FFC84276FAB1FD37840B:8EE26043FFA0FFC4419AFA14FCAA85E9" box="[658,769,1502,1521]" country="Yemen" latitude="12.55" longLatPrecision="129" longitude="54.46" name="Hallah Arhar" pageId="5" pageNumber="112">Hallah Arhar</location>
(spring) /
<geoCoordinate id="EE09505FFFA0FFC44052FA14FC1A85EA" box="[858,945,1501,1522]" degrees="12" direction="north" minutes="33.0" orientation="latitude" pageId="5" pageNumber="112" precision="92" value="12.55">12°33.0N</geoCoordinate>
<geoCoordinate id="EE09505FFFA0FFC440BEFA14FBA685EA" box="[950,1037,1501,1522]" degrees="54" direction="east" minutes="27.6" orientation="longitude" pageId="5" pageNumber="112" precision="92" value="54.46">54°27.6E</geoCoordinate>
,
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<elevation id="0010D1ABFFA0FFC4471EFA14FBEA85EA" box="[1046,1089,1502,1522]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="112" unit="m" value="15.0">15 m</elevation>
</quantity>
/
<date id="FF831058FFA0FFC44374FA33FF738614" box="[124,216,1529,1548]" pageId="5" pageNumber="112" value="2010-11-11">
<collectingDate id="EFC7E9B0FFA0FFC44374FA33FF738614" box="[124,216,1529,1548]" pageId="5" pageNumber="112" value="2010-11-11">11.xi.2010</collectingDate>
</date>
, leg.
<collectorName id="26C8534EFFA0FFC4420DFA33FEFF8614" box="[261,340,1529,1548]" pageId="5" pageNumber="112">J. Hájek</collectorName>
</materialsCitation>
.
<materialsCitation id="3B553CC5FFA0FFC44253FA33FF33863F" collectingDate="2010-11-11" collectionCode="NMPC, KSEM, NHMW, IRSNB" collectorName="J. Hajek'" country="Yemen" elevation="15" latitude="12.55" location="Hallah Arhar" longLatPrecision="129" longitude="54.46" pageId="5" pageNumber="112" specimenCount="7" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="5486883AFFA0FFC44253FA33FE188613" box="[347,435,1529,1548]" pageId="5" pageNumber="112" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D3BFD11FFA0FFC442B5FA33FE778614" box="[445,476,1523,1548]" pageId="5" pageNumber="112" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D3BFD11FFA0FFC442EEFA33FD898614" box="[486,546,1529,1548]" pageId="5" pageNumber="112" type="generic">6 spec.</specimenCount>
(
<collectionCode id="ED2CAE5DFFA0FFC44123FA33FDC08614" box="[555,619,1529,1548]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="112">NMPC</collectionCode>
,
<collectionCode id="ED2CAE5DFFA0FFC4417BFA33FD1B8614" box="[627,688,1529,1548]" pageId="5" pageNumber="112">KSEM</collectionCode>
,
<collectionCode id="ED2CAE5DFFA0FFC441B1FA33FD548614" box="[697,767,1529,1548]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="112" type="Museum">NHMW</collectionCode>
,
<collectionCode id="ED2CAE5DFFA0FFC4400FFA33FCE18614" box="[775,842,1529,1548]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="5" pageNumber="112">IRSNB</collectionCode>
): same label data as the holotype
</materialsCitation>
;
<materialsCitation id="3B553CC5FFA0FFC743AAF9DEFF5280F6" collectionCode="BMNH, NMPC, KSEM" country="Yemen" lastPageId="6" lastPageNumber="113" location="Halla area" municipality="Arher" pageId="5" pageNumber="112" specimenCount="72" typeStatus="paratype">
<specimenCount id="9D3BFD11FFA0FFC443AAF9DEFF42863F" box="[162,233,1556,1575]" pageId="5" pageNumber="112" type="generic">62 spec.</specimenCount>
(
<collectionCode id="ED2CAE5DFFA0FFC443FCF9DEFE93863F" box="[244,312,1556,1575]" pageId="5" pageNumber="112">BMNH</collectionCode>
;
<collectionCode id="ED2CAE5DFFA0FFC44248F9DEFED4863F" box="[320,383,1556,1575]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="112">NMPC</collectionCode>
,
<collectionCode id="ED2CAE5DFFA0FFC44280F9DEFE6E863F" box="[392,453,1556,1575]" pageId="5" pageNumber="112">KSEM</collectionCode>
,
<specimenCount id="9D3BFD11FFA0FFC442C6F9DEFDBD863F" box="[462,534,1556,1575]" pageId="5" pageNumber="112" type="generic">10 spec.</specimenCount>
in DNA grade in coll.
<collectionCode id="ED2CAE5DFFA0FFC441D1F9DEFCB1863F" box="[729,794,1556,1575]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="5" pageNumber="112">NMPC</collectionCode>
):
<collectingCountry id="F32A7608FFA0FFC44024F9DEFCD4863F" box="[812,895,1556,1575]" name="Yemen" pageId="5" pageNumber="112">YEMEN</collectingCountry>
: Socotra island /
<location id="8EE26043FFA0FFC7471AF9DEFF1680F7" LSID="urn:lsid:plazi:treatment:0394878EFFA0FFC84276FAB1FD37840B:8EE26043FFA0FFC7471AF9DEFF1680F7" country="Yemen" lastPageId="6" lastPageNumber="113" municipality="Arher" name="Halla area" pageId="5" pageNumber="112">Halla area</location>
,
<collectingMunicipality id="6BE6ACE2FFA3FFC743CCFF11FF5280F6" box="[196,249,219,238]" pageId="6" pageNumber="113">Arher</collectingMunicipality>
</materialsCitation>
;
<materialsCitation id="3B553CC5FFA3FFC7420AFF11FC628111" collectingDate="2012-06-15" collectorName="Expedition &amp; Bezdek &amp; J. Hajek &amp; V. Hula &amp; Kment &amp; Malenovsky &amp; J. Niedobova &amp; L. Purchart'" country="Yemen" elevation="5" latitude="12.55" location="Socotra" longLatPrecision="129" longitude="54.46" pageId="6" pageNumber="113" specimenCount="1" typeStatus="paratype">
freshwater / spring in sand dune / 9.-10.+
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<collectingDate id="EFC7E9B0FFA3FFC74156FF11FD1680F6" box="[606,701,219,238]" pageId="6" pageNumber="113" value="2012-06-15">15.vi.2012</collectingDate>
</date>
/
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<geoCoordinate id="EE09505FFFA3FFC74023FF11FC2980F7" box="[811,898,219,239]" degrees="54" direction="east" minutes="27.6" orientation="longitude" pageId="6" pageNumber="113" precision="92" value="54.46">54°27.6E</geoCoordinate>
,
<quantity id="4CC59B7DFFA3FFC74083FF11FC0780F7" box="[907,940,219,239]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="113" unit="m" value="5.0">
<elevation id="0010D1ABFFA3FFC74083FF11FC0780F7" box="[907,940,219,239]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="113" unit="m" value="5.0">5 m</elevation>
</quantity>
//
<collectingCountry id="F32A7608FFA3FFC740CAFF16FBA880F7" box="[962,1027,220,239]" name="Yemen" pageId="6" pageNumber="113">Socotra</collectingCountry>
<collectorName id="26C8534EFFA3FFC74701FF11FBCC80F6" box="[1033,1127,219,238]" pageId="6" pageNumber="113">Expedition</collectorName>
2012 / J.
<collectorName id="26C8534EFFA3FFC743E1FF3CFE878111" box="[233,300,246,265]" pageId="6" pageNumber="113">Bezděk</collectorName>
,
<collectorName id="26C8534EFFA3FFC7423DFF3DFED48111" box="[309,383,246,266]" pageId="6" pageNumber="113">J. Hájek</collectorName>
,
<collectorName id="26C8534EFFA3FFC74280FF3DFE618111" box="[392,458,246,266]" pageId="6" pageNumber="113">V. Hula</collectorName>
/ P.
<collectorName id="26C8534EFFA3FFC742FBFF3DFD848112" box="[499,559,247,266]" pageId="6" pageNumber="113">Kment</collectorName>
, I.
<collectorName id="26C8534EFFA3FFC74144FF3CFD138111" box="[588,696,246,265]" pageId="6" pageNumber="113">Malenovský</collectorName>
, /
<collectorName id="26C8534EFFA3FFC741C6FF3DFCEB8111" box="[718,832,246,266]" pageId="6" pageNumber="113">J. Niedobová</collectorName>
&amp;
<collectorName id="26C8534EFFA3FFC74056FF3DFC628111" box="[862,969,246,266]" pageId="6" pageNumber="113">L. Purchart</collectorName>
</materialsCitation>
;
<materialsCitation id="3B553CC5FFA3FFC740DBFF3CFCF2813C" collectionCode="NMPC" country="Yemen" location="Shuab" pageId="6" pageNumber="113" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D3BFD11FFA3FFC740DBFF3CFBBA8112" box="[979,1041,246,266]" pageId="6" pageNumber="113" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED2CAE5DFFA3FFC74714FF3CFBF58111" box="[1052,1118,246,265]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="6" pageNumber="113">NMPC</collectionCode>
):
<collectingCountry id="F32A7608FFA3FFC74392FEDBFF46813C" box="[154,237,273,292]" name="Yemen" pageId="6" pageNumber="113">YEMEN</collectingCountry>
: Socotra Island / ca.
<quantity id="4CC59B7DFFA3FFC742AFFEDBFE7F813C" box="[423,468,273,292]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="113" unit="km" value="3.0">3 km</quantity>
NE of
<location id="8EE26043FFA3FFC7411FFEDBFDE7813C" LSID="urn:lsid:plazi:treatment:0394878EFFA0FFC84276FAB1FD37840B:8EE26043FFA3FFC7411FFEDBFDE7813C" box="[535,588,273,292]" country="Yemen" name="Shuab" pageId="6" pageNumber="113">Shuab</location>
/
<taxonomicName id="4C3D4D1BFFA3FFC74169FED8FD51813C" box="[609,762,274,292]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="6" pageNumber="113" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="B949EA8AFFA3FFC74169FED8FD51813C" box="[609,762,274,292]" italics="true" pageId="6" pageNumber="113">Avicennia marina</emphasis>
</taxonomicName>
mangrove
</materialsCitation>
; /
<materialsCitation id="3B553CC5FFA3FFC74078FEDBFE128141" collectingDate="2012-06-20" collectingDateMax="2012-06-21" collectingDateMin="2012-06-20" collectorName="Bezdek &amp; J. Hajek &amp; Hula &amp; Kment &amp; Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" elevation="3" latitude="12.568334" location="SOCOTRA" longLatPrecision="129" longitude="53.398335" pageId="6" pageNumber="113" specimenCount="1" typeStatus="paratype">
sand dunes,
<date id="FF831058FFA3FFC740D4FEDBFBF2813C" box="[988,1113,273,292]" pageId="6" pageNumber="113" value="2012-06-20" valueMax="2012-06-21" valueMin="2012-06-20">
<collectingDate id="EFC7E9B0FFA3FFC740D4FEDBFBF2813C" box="[988,1113,273,292]" pageId="6" pageNumber="113" value="2012-06-20" valueMax="2012-06-21" valueMin="2012-06-20">20.-21.vi.2012</collectingDate>
</date>
/
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<geoCoordinate id="EE09505FFFA3FFC743FCFEE1FEE08127" box="[244,331,299,319]" degrees="53" direction="east" minutes="23.9" orientation="longitude" pageId="6" pageNumber="113" precision="92" value="53.398335">53°23.9E</geoCoordinate>
,
<quantity id="4CC59B7DFFA3FFC7425BFEE6FEDE8127" box="[339,373,300,319]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="113" unit="m" value="3.0">
<elevation id="0010D1ABFFA3FFC7425BFEE6FEDE8127" box="[339,373,300,319]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="113" unit="m" value="3.0">3 m</elevation>
</quantity>
//
<collectingCountry id="F32A7608FFA3FFC74283FEE6FE468127" box="[395,493,300,319]" name="Yemen" pageId="6" pageNumber="113">SOCOTRA</collectingCountry>
expedition 2012 / J.
<collectorName id="26C8534EFFA3FFC741A8FEE1FD488126" box="[672,739,299,319]" pageId="6" pageNumber="113">Bezděk</collectorName>
,
<collectorName id="26C8534EFFA3FFC741E3FEE6FC988126" box="[747,819,299,319]" pageId="6" pageNumber="113">J. Hájek</collectorName>
, V.
<collectorName id="26C8534EFFA3FFC7405BFEE1FCD78126" box="[851,892,299,318]" pageId="6" pageNumber="113">Hula</collectorName>
/ P.
<collectorName id="26C8534EFFA3FFC740A9FEE6FC768127" box="[929,989,300,319]" pageId="6" pageNumber="113">Kment</collectorName>
, I.
<collectorName id="26C8534EFFA3FFC740F0FEE1FBCF8126" box="[1016,1124,299,318]" pageId="6" pageNumber="113">Malenovský</collectorName>
, /
<collectorName id="26C8534EFFA3FFC743AAFE8DFEB88141" box="[162,275,326,346]" pageId="6" pageNumber="113">J. Niedobová</collectorName>
&amp;
<collectorName id="26C8534EFFA3FFC74226FE8DFE258141" box="[302,398,326,346]" pageId="6" pageNumber="113">L. Purchart</collectorName>
leg.
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA3FFC9439FFEBBFEF1828B" lastPageId="8" lastPageNumber="115" pageId="6" pageNumber="113" type="description">
<paragraph id="8B823698FFA3FFC7439FFEBBFEF98212" blockId="6.[151,1128,369,1578]" pageId="6" pageNumber="113">
<emphasis id="B949EA8AFFA3FFC7439FFEBBFE898191" bold="true" box="[151,290,369,393]" pageId="6" pageNumber="113">Description.</emphasis>
Body weakly elongate, lowly convex in lateral view. Body length
<quantity id="4CC59B7DFFA3FFC740EFFEBBFBCD8191" box="[999,1126,369,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="6" pageNumber="113" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.31.5 mm</quantity>
(
<typeStatus id="5486883AFFA3FFC74396FE58FF5681B2" box="[158,253,402,426]" pageId="6" pageNumber="113" type="holotype">holotype</typeStatus>
<quantity id="4CC59B7DFFA3FFC7420DFE58FEF781B2" box="[261,348,402,426]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="113" unit="mm" value="1.4">1.4 mm</quantity>
), width of head
<quantity id="4CC59B7DFFA3FFC74105FE5BFD0281B1" box="[525,681,401,425]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="6" pageNumber="113" unit="mm" value="0.4" valueMax="0.45" valueMin="0.35">0.350.45 mm</quantity>
(
<typeStatus id="5486883AFFA3FFC741B1FE58FCB381B2" box="[697,792,402,426]" pageId="6" pageNumber="113" type="holotype">holotype</typeStatus>
<quantity id="4CC59B7DFFA3FFC74028FE58FC2881B2" box="[800,899,402,426]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="113" unit="mm" value="0.37">0.37 mm</quantity>
), maximum width of pronotum
<quantity id="4CC59B7DFFA3FFC7420BFE7BFE2A81D1" box="[259,385,433,457]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="113" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50.6 mm</quantity>
(
<typeStatus id="5486883AFFA3FFC74285FE78FE4781D2" box="[397,492,434,458]" pageId="6" pageNumber="113" type="holotype">holotype</typeStatus>
<quantity id="4CC59B7DFFA3FFC742F9FE7BFDEF81D2" box="[497,580,433,458]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="113" unit="mm" value="0.5">0.5 mm</quantity>
), maximum width of elytra
<quantity id="4CC59B7DFFA3FFC7406BFE7BFC5081D1" box="[867,1019,433,457]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.5" metricValueMin="7.5" pageId="6" pageNumber="113" unit="mm" value="0.85" valueMax="0.95" valueMin="0.75">0.750.95 mm</quantity>
(
<typeStatus id="5486883AFFA3FFC74700FE78FBCC81D2" box="[1032,1127,434,458]" pageId="6" pageNumber="113" type="holotype">holotype</typeStatus>
<quantity id="4CC59B7DFFA3FFC7439FFE18FF5781F2" box="[151,252,466,490]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="113" unit="mm" value="0.8">0.80 mm</quantity>
). Coloration elytra and pronotum reddish brown to brown with olive reflections, head dark brown.
</paragraph>
<paragraph id="8B823698FFA3FFC743BDFDD8FE078352" blockId="6.[151,1128,369,1578]" pageId="6" pageNumber="113">
Head (
<figureCitation id="13062A1DFFA3FFC743F2FDD8FEEC8232" box="[250,327,530,554]" captionStart="Figs" captionStartId="7.[113,152,1482,1501]" captionTargetBox="[113,1087,217,1402]" captionTargetId="figure-13@7.[113,1087,217,1402]" captionTargetPageId="7" captionText="Figs. 914. Georissus nemo sp. nov. (913 paratype; 14 holotype). 9 pronotum and elytra in dorsal view; 10 head, dorsal view; 1112 detail of pronotum (11 dorsal view; 12 dorsolateral view); 13 elytron, dorsolateral view; 14 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334588" httpUri="https://zenodo.org/record/5334588/files/figure.png" pageId="6" pageNumber="113">Fig. 10</figureCitation>
). Clypeus weakly convex, with scattered sparsely arranged granules, anterior margin with marginal row of densely arranged granules; each posterolateral portion of clypeus anteriorly to eye declined, divided from mesal portion by ridge arising from frons, ridge high and sharp posteriorly, but more weakly defined anteriorly before joining with marginal clypeal row of granules. Frons with central elongate depression surrounded by elevated ridges each bearing several weakly pronounced granules; anterior portion of frons with sharp sublateral ridge without granules at midlength between central depression and inner margin of each eye (this ridges continues to clypeus more anteriorly); submesal portion with transverse blunt ridge connecting midlength of central ridge with posterior portion of sublateral ridge. Eyes large, oval in lateral view.
</paragraph>
<paragraph id="8B823698FFA3FFC743BDFC98FDAF8491" blockId="6.[151,1128,369,1578]" pageId="6" pageNumber="113">
Prothorax (
<figureCitation id="13062A1DFFA3FFC74223FC98FE008372" box="[299,427,850,874]" captionStart="Figs" captionStartId="7.[113,152,1482,1501]" captionTargetBox="[113,1087,217,1402]" captionTargetId="figure-13@7.[113,1087,217,1402]" captionTargetPageId="7" captionText="Figs. 914. Georissus nemo sp. nov. (913 paratype; 14 holotype). 9 pronotum and elytra in dorsal view; 10 head, dorsal view; 1112 detail of pronotum (11 dorsal view; 12 dorsolateral view); 13 elytron, dorsolateral view; 14 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334588" httpUri="https://zenodo.org/record/5334588/files/figure.png" pageId="6" pageNumber="113">Figs. 1112</figureCitation>
). Pronotum 1.1× as wide as long, with maximum width at posterior 0.4; lateral portions very small, slightly projecting laterad as blunt elongate lobes only. Anterior portion with two closely situated submedian ridges delimiting shallow median groove, each side laterally of ridges weakly convex, with scattered granules. Central portion of pronotum with large but rather shallow rhomboid depression delimited by wide granuliferous ridges anteriorly and narrow ones posteriorly, posterior ridges not joining but converging to posterior margin; posterolaterally of central depression with pair of large but shallow impressions delimited laterally by small granuliferous bulge; posterolateral portion of pronotum each with high granuliferous protuberance. Posterolateral pits absent. Ventral morphology of prothorax corresponding with
<taxonomicName id="4C3D4D1BFFA3FFC74264FBB9FE568492" baseAuthorityName="Rossi" baseAuthorityYear="1794" box="[364,509,1139,1162]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="113" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B949EA8AFFA3FFC74264FBB9FE568492" box="[364,509,1139,1162]" italics="true" pageId="6" pageNumber="113">G. crenulatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B823698FFA3FFC743BDFB58FC1E8612" blockId="6.[151,1128,369,1578]" pageId="6" pageNumber="113">
Mesothorax. Elytra (
<figureCitation id="13062A1DFFA3FFC74286FB58FE5684B2" box="[398,509,1170,1194]" captionStart="Figs" captionStartId="7.[113,152,1482,1501]" captionTargetBox="[113,1087,217,1402]" captionTargetId="figure-13@7.[113,1087,217,1402]" captionTargetPageId="7" captionText="Figs. 914. Georissus nemo sp. nov. (913 paratype; 14 holotype). 9 pronotum and elytra in dorsal view; 10 head, dorsal view; 1112 detail of pronotum (11 dorsal view; 12 dorsolateral view); 13 elytron, dorsolateral view; 14 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334588" httpUri="https://zenodo.org/record/5334588/files/figure.png" pageId="6" pageNumber="113">Figs. 9, 13</figureCitation>
) combined 1.2× as long as wide, 2.2× as long as pronotum; base of elytra ca. as wide as maximum width of pronotum, maximum width of elytra between anterior 0.10.5, elytra gradually narrowing in posterior 0.50.2, apex strongly narrowed. Elytral suture and intervals 2, 4 and 6 elevated into high narrow ridges, ridge on interval 6 arising from large humeral protuberance; lateral portion with Y-shaped structure formed by highly elevate interval 8 and anterior portion of interval 9; odd intervals (1, 3, 5) flat, not elevate, interval 7 flat anteriorly and becoming slightly elevate posteriorly. All ridges and humeral protuberance with very weak and low granules only, hence the elytral ridges nearly smooth; odd elytral intervals only with scattered and extremely low granules, hence appearing nearly totally flat. Elytral series regular, serial punctures small but sharply impressed. Lateral-most portion of elytron declined, hence elytral laterally without clear projections. Median pentagonal protuberance of mesoventrite flat, without distinct pits.
</paragraph>
<paragraph id="8B823698FFA3FFC943BDF9D8FDE9810C" blockId="6.[151,1128,369,1578]" lastBlockId="8.[151,1127,219,1556]" lastPageId="8" lastPageNumber="115" pageId="6" pageNumber="113">Metathorax. Metaventrite ca. 2× as long as mesoventrite, flat, only with few scattered indistinct granules, without distinct median discrimen. Metathoracic wings present in specimens examined for this character (n=2).</paragraph>
<caption id="DF426610FFA2FFC64379FA00FE8D860A" ID-DOI="http://doi.org/10.5281/zenodo.5334588" ID-Zenodo-Dep="5334588" httpUri="https://zenodo.org/record/5334588/files/figure.png" pageId="7" pageNumber="114" startId="7.[113,152,1482,1501]" targetBox="[113,1087,217,1402]" targetPageId="7">
<paragraph id="8B823698FFA2FFC64379FA00FE8D860A" blockId="7.[113,1090,1482,1555]" pageId="7" pageNumber="114">
Figs. 914.
<taxonomicName id="4C3D4D1BFFA2FFC643DFFA01FECB85C5" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[215,352,1483,1501]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFA2FFC643DFFA01FECB85C5" box="[215,352,1483,1501]" italics="true" pageId="7" pageNumber="114">Georissus nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA2FFC6426EFA00FE0285C5" box="[358,425,1482,1501]" pageId="7" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
(913 paratype; 14 holotype). 9 pronotum and elytra in dorsal view; 10 head, dorsal view; 1112 detail of pronotum (11 dorsal view; 12 dorsolateral view); 13 elytron, dorsolateral view; 14 abdomen.
</paragraph>
</caption>
<paragraph id="8B823698FFADFFC943BDFED1FEE4818B" blockId="8.[151,1127,219,1556]" pageId="8" pageNumber="115">
Abdomen (
<figureCitation id="13062A1DFFADFFC94238FED1FE2B812C" box="[304,384,283,308]" captionStart="Figs" captionStartId="7.[113,152,1482,1501]" captionTargetBox="[113,1087,217,1402]" captionTargetId="figure-13@7.[113,1087,217,1402]" captionTargetPageId="7" captionText="Figs. 914. Georissus nemo sp. nov. (913 paratype; 14 holotype). 9 pronotum and elytra in dorsal view; 10 head, dorsal view; 1112 detail of pronotum (11 dorsal view; 12 dorsolateral view); 13 elytron, dorsolateral view; 14 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334588" httpUri="https://zenodo.org/record/5334588/files/figure.png" pageId="8" pageNumber="115">Fig. 14</figureCitation>
) gradually narrowing posteriad. Ventrites 1 flat, only with sparsely arranged indistinct granules especially in posterior third (anterior portion of some specimens totally bare), ventrites 24 without granules, ventrite 5 flat with few indistinct granules along posterior margin.
</paragraph>
<paragraph id="8B823698FFADFFC943BDFE51FEF1828B" blockId="8.[151,1127,219,1556]" pageId="8" pageNumber="115">
Male genitalia (
<figureCitation id="13062A1DFFADFFC94268FE51FE0981AC" box="[352,418,411,436]" captionStart="Figs" captionStartId="4.[153,192,1033,1052]" captionTargetBox="[266,1016,226,1012]" captionTargetId="figure-178@4.[262,1016,219,1015]" captionTargetPageId="4" captionText="Figs. 78. Genitalia of Socotran endemic Georissus species. 7 G. maritimus sp. nov. (a, b: two different ventral views of the same aedeagus; c: lateral view); 8 G. nemo sp. nov., ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334586" httpUri="https://zenodo.org/record/5334586/files/figure.png" pageId="8" pageNumber="115">Fig. 8</figureCitation>
). Aedeagus
<quantity id="4CC59B7DFFADFFC9412FFE56FDD281AB" box="[551,633,412,436]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="115" unit="mm" value="0.4">0.4 mm</quantity>
long (measured in
<typeStatus id="5486883AFFADFFC94044FE51FC0281AB" box="[844,937,411,435]" pageId="8" pageNumber="115" type="holotype">holotype</typeStatus>
only). Parameres 0.85× as long as phallobase, their combined width slightly narrower than maximum width of phallobase; lateral margins of parameres very slightly convergent in basal 0.6, arcuately bent inward apically; apex widely obtusely pointed; inner margins straight, well sclerotized. Median lobe 0.75× length of parameres, narrowly triangular apically, apex sharp, gonopore situated subapical, struts ca. 0.4× as long as apical portion of median lobe. Phallobase slightly widening posteriad, with wide but indistinct latero-posterior band along margins, without posterior opening.
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFADFFC9439FFD51FBE9858B" pageId="8" pageNumber="115" type="diagnosis">
<paragraph id="8B823698FFADFFC9439FFD51FBAE848B" blockId="8.[151,1127,219,1556]" pageId="8" pageNumber="115">
<emphasis id="B949EA8AFFADFFC9439FFD51FE2682AB" bold="true" box="[151,397,667,691]" pageId="8" pageNumber="115">Differential diagnosis.</emphasis>
Based on the general sculpture of the pronotum (i.e., rhomboid central depression and low submedian and higher lateral tubercles,
<figureCitation id="13062A1DFFADFFC9401DFD71FCCA82CC" box="[789,865,699,724]" captionStart="Figs" captionStartId="7.[113,152,1482,1501]" captionTargetBox="[113,1087,217,1402]" captionTargetId="figure-13@7.[113,1087,217,1402]" captionTargetPageId="7" captionText="Figs. 914. Georissus nemo sp. nov. (913 paratype; 14 holotype). 9 pronotum and elytra in dorsal view; 10 head, dorsal view; 1112 detail of pronotum (11 dorsal view; 12 dorsolateral view); 13 elytron, dorsolateral view; 14 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334588" httpUri="https://zenodo.org/record/5334588/files/figure.png" pageId="8" pageNumber="115">Fig. 11</figureCitation>
,
<figureCitation id="13062A1DFFADFFC94065FD76FC2382CC" box="[877,904,700,724]" captionStart="Figs" captionStartId="10.[151,190,1491,1510]" captionTargetBox="[153,1125,221,1476]" captionTargetId="figure-12@10.[147,1128,216,1478]" captionTargetPageId="10" captionText="Figs. 1522.Socotran endemic Georissus species and their habitats. 1516 G. maritimus sp. nov.(15 dorsal view; 16 lateral view). 1718 G. nemo sp. nov. (17 dorsal view; 18 lateral view). 1920 the sand tongue 3 km NE of Shuab, type locality of G. maritimus sp. nov. (19 general view of the habitat; 20 satellite view showing the position of the locality); 2122 the stream at Arhar, type locality of G. nemo sp. nov. (21 general view of the habitat; 22 satellite view showing the position of the locality)." figureDoi="http://doi.org/10.5281/zenodo.5334592" httpUri="https://zenodo.org/record/5334592/files/figure.png" pageId="8" pageNumber="115">17</figureCitation>
) and elytra (i.e., odd intervals more elevated than even ones),
<taxonomicName id="4C3D4D1BFFADFFC94142FD17FD5E82EC" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[586,757,733,756]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFADFFC94142FD17FD5E82EC" box="[586,757,733,756]" italics="true" pageId="8" pageNumber="115">Georissus nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFADFFC941F3FD16FCE582EB" box="[763,846,732,755]" pageId="8" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C3D4D1BFFADFFC940E7FD17FBCD82EC" authorityName="Laporte de Castelnau" authorityYear="1840" box="[1007,1126,733,756]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B949EA8AFFADFFC940E7FD17FBCD82EC" box="[1007,1126,733,756]" italics="true" pageId="8" pageNumber="115">G. costatus</emphasis>
</taxonomicName>
species group sensu
<bibRefCitation id="EFAC4B69FFADFFC94262FD36FDA0830B" author="DELEVE J." box="[362,523,763,787]" pageId="8" pageNumber="115" pagination="1 - 23" refId="ref12544" refString="DELEVE J. 1967 a: Notes sur les Georissus d'Afrique et descriptions d'especes nouvelles (Coleoptera Georissidae). Bulletin de l ' Institute Royal des Sciences Naturelles de Belgique 43 (26): 1 - 23." type="journal article" year="1967">DELÈVE (1967a</bibRefCitation>
,b). It may be distinguished from other species of the group by the combination of the following characters: elytral ridges highly elevated, very weakly denticulate in lateral view; intervals 2, 4 and 6 nearly completely flat with very few indistinct granules; lateral portions of pronotum not projecting into acute lobes; abdomen with very indistinct granules on ventrites 1 and 5 only; phallobase slightly longer than parameres, only indistinctly widened posteriad; parameres widely arcuate apically. By elytral morphology,
<taxonomicName id="4C3D4D1BFFADFFC9439FFC77FF5983CC" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[151,242,957,980]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFADFFC9439FFC77FF5983CC" box="[151,242,957,980]" italics="true" pageId="8" pageNumber="115">G. nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFADFFC943F1FC76FEE783CB" box="[249,332,956,979]" pageId="8" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
especially resembles the African species
<taxonomicName id="4C3D4D1BFFADFFC94000FC77FB9283CC" authority="Grouvelle, 1909" authorityName="Grouvelle" authorityYear="1909" box="[776,1081,955,980]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="alticosta">
<emphasis id="B949EA8AFFADFFC94000FC77FC2E83CC" box="[776,901,957,980]" italics="true" pageId="8" pageNumber="115">G. alticosta</emphasis>
Grouvelle, 1909
</taxonomicName>
and
<taxonomicName id="4C3D4D1BFFADFFC9439FFC17FDB683EC" authority="Paulian &amp; Legros, 1943" authorityName="Paulian &amp; Legros" authorityYear="1943" box="[151,541,987,1012]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="B949EA8AFFADFFC9439FFC17FEB983EC" box="[151,274,989,1012]" italics="true" pageId="8" pageNumber="115">G. decorsei</emphasis>
Paulian &amp; Legros, 1943
</taxonomicName>
, and
<taxonomicName id="4C3D4D1BFFADFFC9415DFC17FCDE83EC" authority="Deleve, 1972" authorityName="Deleve" authorityYear="1972" box="[597,885,987,1012]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="decoratus">
<emphasis id="B949EA8AFFADFFC9415DFC17FD4B83EC" box="[597,736,989,1012]" italics="true" pageId="8" pageNumber="115">G. decoratus</emphasis>
<bibRefCitation id="EFAC4B69FFADFFC941EFFC11FCDE83EC" author="DELEVE J." box="[743,885,987,1012]" pageId="8" pageNumber="115" pagination="149 - 165" refId="ref12618" refString="DELEVE J. 1972: Les Georissidae (Coleoptera) de Ceylan. Bulletin et Annales de la Societe Royal de la Societe Entomologique de Belgique 108: 149 - 165." type="journal article" year="1972">Delève, 1972</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F32A7608FFADFFC940B0FC11FB8883EB" box="[952,1059,987,1011]" name="Sri Lanka" pageId="8" pageNumber="115">Sri Lanka</collectingCountry>
, all of which may be easily distinguished by the morphology of the aedeagus which is much narrower in both African species (
<bibRefCitation id="EFAC4B69FFADFFC9429AFBD6FD81842B" author="DELEVE J." box="[402,554,1051,1076]" pageId="8" pageNumber="115" pagination="1 - 23" refId="ref12544" refString="DELEVE J. 1967 a: Notes sur les Georissus d'Afrique et descriptions d'especes nouvelles (Coleoptera Georissidae). Bulletin de l ' Institute Royal des Sciences Naturelles de Belgique 43 (26): 1 - 23." type="journal article" year="1967">DELÈVE 1967a</bibRefCitation>
,
<figureCitation id="13062A1DFFADFFC9413BFBD1FDC4842C" box="[563,623,1051,1076]" captionStart="Figs" captionStartId="4.[153,192,1033,1052]" captionTargetBox="[266,1016,226,1012]" captionTargetId="figure-178@4.[262,1016,219,1015]" captionTargetPageId="4" captionText="Figs. 78. Genitalia of Socotran endemic Georissus species. 7 G. maritimus sp. nov. (a, b: two different ventral views of the same aedeagus; c: lateral view); 8 G. nemo sp. nov., ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334586" httpUri="https://zenodo.org/record/5334586/files/figure.png" pageId="8" pageNumber="115">Fig. 8</figureCitation>
) and much wider in
<taxonomicName id="4C3D4D1BFFADFFC9404BFBD7FC61842C" authorityName="Deleve" authorityYear="1972" box="[835,970,1053,1076]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="decoratus">
<emphasis id="B949EA8AFFADFFC9404BFBD7FC61842C" box="[835,970,1053,1076]" italics="true" pageId="8" pageNumber="115">G. decoratus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAC4B69FFADFFC940DFFBD6FBC8842C" author="DELEVE J." box="[983,1123,1052,1076]" pageId="8" pageNumber="115" pagination="149 - 165" refId="ref12618" refString="DELEVE J. 1972: Les Georissidae (Coleoptera) de Ceylan. Bulletin et Annales de la Societe Royal de la Societe Entomologique de Belgique 108: 149 - 165." type="journal article" year="1972">DELÈVE 1972</bibRefCitation>
,
<figureCitation id="13062A1DFFADFFC9439FFBF1FF43844B" box="[151,232,1083,1107]" captionStart="Figs" captionStartId="10.[151,190,1491,1510]" captionTargetBox="[153,1125,221,1476]" captionTargetId="figure-12@10.[147,1128,216,1478]" captionTargetPageId="10" captionText="Figs. 1522.Socotran endemic Georissus species and their habitats. 1516 G. maritimus sp. nov.(15 dorsal view; 16 lateral view). 1718 G. nemo sp. nov. (17 dorsal view; 18 lateral view). 1920 the sand tongue 3 km NE of Shuab, type locality of G. maritimus sp. nov. (19 general view of the habitat; 20 satellite view showing the position of the locality); 2122 the stream at Arhar, type locality of G. nemo sp. nov. (21 general view of the habitat; 22 satellite view showing the position of the locality)." figureDoi="http://doi.org/10.5281/zenodo.5334592" httpUri="https://zenodo.org/record/5334592/files/figure.png" pageId="8" pageNumber="115">Fig. 16</figureCitation>
).
<taxonomicName id="4C3D4D1BFFADFFC943F4FBF7FE35844C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[252,414,1085,1108]" class="Magnoliopsida" family="Avicenniaceae" genus="Georisus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="8" pageNumber="115" phylum="Tracheophyta" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFADFFC943F4FBF7FE35844C" box="[252,414,1085,1108]" italics="true" pageId="8" pageNumber="115">Georisus nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFADFFC942AEFBF6FE51844B" box="[422,506,1084,1107]" pageId="8" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
also differs from both African species by the indistinctly denticulate elytral costae (costae are totally smooth in
<taxonomicName id="4C3D4D1BFFADFFC941DAFB97FCE6846C" authorityName="Grouvelle" authorityYear="1909" box="[722,845,1117,1140]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="alticosta">
<emphasis id="B949EA8AFFADFFC941DAFB97FCE6846C" box="[722,845,1117,1140]" italics="true" pageId="8" pageNumber="115">G. alticosta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3D4D1BFFADFFC94077FB97FC53846C" authorityName="Paulian &amp; Legros" authorityYear="1943" box="[895,1016,1117,1140]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="decorsei">
<emphasis id="B949EA8AFFADFFC94077FB97FC53846C" box="[895,1016,1117,1140]" italics="true" pageId="8" pageNumber="115">G. decorsei</emphasis>
</taxonomicName>
) and from
<taxonomicName id="4C3D4D1BFFADFFC9439FFBB7FE8A848C" authorityName="Deleve" authorityYear="1972" box="[151,289,1149,1172]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="decoratus">
<emphasis id="B949EA8AFFADFFC9439FFBB7FE8A848C" box="[151,289,1149,1172]" italics="true" pageId="8" pageNumber="115">G. decoratus</emphasis>
</taxonomicName>
by flat elytral intervals 2 and 4 (slightly convex in the latter species).
</paragraph>
<paragraph id="8B823698FFADFFC943BDFB57FBE9858B" blockId="8.[151,1127,219,1556]" pageId="8" pageNumber="115">
<taxonomicName id="4C3D4D1BFFADFFC943BDFB57FECE84AC" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[181,357,1181,1204]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFADFFC943BDFB57FECE84AC" box="[181,357,1181,1204]" italics="true" pageId="8" pageNumber="115">Georissus nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFADFFC94267FB56FE6E84AB" box="[367,453,1180,1203]" pageId="8" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
may be easily distinguished from the following unidentified species from
<collectingCountry id="F32A7608FFADFFC9422DFB76FE1584CB" box="[293,446,1211,1236]" name="Socotra Island" pageId="8" pageNumber="115">Socotra Island</collectingCountry>
by its smaller body size (the other species is 1.82.0 mm long), elytral costae with very indistinct denticulation (strongly denticulate in the other species), elytral intervals 2 and 4 flat (convex and bearing many distinct granules in the other species), abdominal ventrites with extremely indistinct granulation (with very distict granulation on whole ventrites 1 and
<quantity id="4CC59B7DFFADFFC94276FAF1FE0E854B" box="[382,421,1339,1363]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="115" unit="in" value="5.0">5 in</quantity>
the other species), and by the weakly metallic coloration (head and pronotum strongly and elytra moderately metallic in the other species). From the third Socotran species of
<taxonomicName id="4C3D4D1BFFADFFC94200FAB7FED8858C" authorityName="Latreille" authorityYear="1809" box="[264,371,1405,1428]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA8AFFADFFC94200FAB7FED8858C" box="[264,371,1405,1428]" italics="true" pageId="8" pageNumber="115">Georissus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3D4D1BFFADFFC94288FAB7FDA7858C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[384,524,1405,1428]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFADFFC94288FAB7FDA7858C" box="[384,524,1405,1428]" italics="true" pageId="8" pageNumber="115">G. maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFADFFC9411BFAB6FDC3858B" box="[531,616,1404,1427]" pageId="8" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
, it differs by its costate odd elytral intervals.
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFADFFC8439FFA56FE2F822B" lastPageId="9" lastPageNumber="116" pageId="8" pageNumber="115" type="etymology">
<paragraph id="8B823698FFADFFC9439FFA56FCFD860C" blockId="8.[151,1127,219,1556]" pageId="8" pageNumber="115">
<emphasis id="B949EA8AFFADFFC9439FFA56FEB385AC" bold="true" box="[151,280,1436,1460]" pageId="8" pageNumber="115">Etymology.</emphasis>
The species name refers to Captain Nemo, a fictional character of two novels by Jules Verne, who lived underseas (in a submarine Nautilus), hence in an environment unusual for a human. This resembles specimens of
<taxonomicName id="4C3D4D1BFFADFFC94156FA17FCA285EC" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[606,777,1501,1524]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="115" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFADFFC94156FA17FCA285EC" box="[606,777,1501,1524]" italics="true" pageId="8" pageNumber="115">Georissus nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFADFFC94007FA16FCC985EB" box="[783,866,1500,1523]" pageId="8" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
collected in 2010 which were found underwater, in an environment unusual for this genus.
</paragraph>
<paragraph id="8B823698FFACFFC84379FF11FE2F822B" blockId="9.[113,1089,219,1044]" pageId="9" pageNumber="116">
<emphasis id="B949EA8AFFACFFC84379FF11FE3880EB" bold="true" box="[113,403,219,243]" pageId="9" pageNumber="116">Collection circumstances.</emphasis>
The vast majority of specimens in the
<typeStatus id="5486883AFFACFFC84027FF16FCF680EC" box="[815,861,220,244]" pageId="9" pageNumber="116">type</typeStatus>
series were collected in Arhar along the permanent stream rising below the sand dunes on the base of rock cliffs of the
<collectingCountry id="F32A7608FFACFFC84391FED6FF41812C" box="[153,234,284,308]" name="Yemen" pageId="9" pageNumber="116">Socotra</collectingCountry>
Plateau falling to the sea coast. The specimens inhabited the sandy waterlogged surroundings of the stream partly overgrown by short-grazed lawn of few undetermined
<taxonomicName id="4C3D4D1BFFACFFC84379FE96FF63816B" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[113,200,348,371]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
and
<taxonomicName id="4C3D4D1BFFACFFC8420BFE96FED4816C" authorityName="de Jussieu" authorityYear="1789" box="[259,383,348,372]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
and surrounded by shrubs of
<taxonomicName id="4C3D4D1BFFACFFC841C7FE97FEB2818B" authority="(Ehrenb.) Bunge" authorityName="Bunge" baseAuthorityName="Ehrenb." class="Magnoliopsida" family="Tamaricaceae" genus="Tamarix" kingdom="Plantae" order="Caryophyllales" pageId="9" pageNumber="116" phylum="Tracheophyta" rank="species" species="nilotica">
<emphasis id="B949EA8AFFACFFC841C7FE97FC2A816C" box="[719,897,349,372]" italics="true" pageId="9" pageNumber="116">Tamarix nilotica</emphasis>
(Ehrenb.) Bunge (Tamaricaceae)
</taxonomicName>
. The majority of the specimens were collected at night, creeping on the bare wet sand, while a portion of the specimens were attracted at light trap installed close to the stream. In 2010, when the locality was only shortly visited in the daytime, few specimens were found on the submerged underside of the stones directly in the stream. One specimen was also found among a large number of
<taxonomicName id="4C3D4D1BFFACFFC84122FE37FD1C820C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[554,695,509,532]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFACFFC84122FE37FD1C820C" box="[554,695,509,532]" italics="true" pageId="9" pageNumber="116">G. maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFACFFC841B6FE36FCBB820B" box="[702,784,508,531]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
from the
<typeStatus id="5486883AFFACFFC84070FE36FC0D820C" box="[888,934,508,532]" pageId="9" pageNumber="116">type</typeStatus>
locality of the latter species (see above).
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFACFFC84379FDF1FD37840B" pageId="9" pageNumber="116" type="distribution">
<paragraph id="8B823698FFACFFC84379FDF1FD45826B" blockId="9.[113,1089,219,1044]" pageId="9" pageNumber="116">
<emphasis id="B949EA8AFFACFFC84379FDF1FEA8824B" bold="true" box="[113,259,571,595]" pageId="9" pageNumber="116">Distribution.</emphasis>
Known from two distant localities on
<collectingCountry id="F32A7608FFACFFC84198FDF6FC81824B" box="[656,810,571,596]" name="Socotra Island" pageId="9" pageNumber="116">Socotra Island</collectingCountry>
, indicating that the species may be widely distributed on suitable habitats of the island.
</paragraph>
<paragraph id="8B823698FFACFFC84379FDB6FD37840B" blockId="9.[113,1089,219,1044]" pageId="9" pageNumber="116">
<emphasis id="B949EA8AFFACFFC84379FDB6FF13828C" bold="true" box="[113,184,636,660]" pageId="9" pageNumber="116">Notes.</emphasis>
Although the relationships between the species of
<taxonomicName id="4C3D4D1BFFACFFC841CFFDB7FC7A828C" authority="Latreille, 1809" authorityName="Latreille" authorityYear="1809" box="[711,977,635,660]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA8AFFACFFC841CFFDB7FC9A828C" box="[711,817,637,660]" italics="true" pageId="9" pageNumber="116">Georissus</emphasis>
Latreille, 1809
</taxonomicName>
are poorly known and the species groups defined by the pronotal and elytral sculpture may be easily polyphyletic, the strong resemblance of
<taxonomicName id="4C3D4D1BFFACFFC8412DFD77FD2A82CC" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[549,641,701,724]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFACFFC8412DFD77FD2A82CC" box="[549,641,701,724]" italics="true" pageId="9" pageNumber="116">G. nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFACFFC84181FD76FD7682CB" box="[649,733,700,723]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
to some African and Sri Lankan species (see Differential diagnosis) is rather striking. Moreover, there is a long series of an undescribed
<taxonomicName id="4C3D4D1BFFACFFC843F0FD37FE1E830B" box="[248,437,763,788]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B949EA8AFFACFFC843F0FD37FEC8830C" box="[248,355,765,788]" italics="true" pageId="9" pageNumber="116">Georissus</emphasis>
species
</taxonomicName>
in NMPC which was collected in southern
<collectingCountry id="F32A7608FFACFFC84076FD31FC1E830B" box="[894,949,763,787]" name="India" pageId="9" pageNumber="116">India</collectingCountry>
(
<collectingRegion id="49F9F87AFFACFFC840C9FD31FF0F832B" country="India" name="Tamil Nadu" pageId="9" pageNumber="116">Tamil Nadu state</collectingRegion>
, S of Tuticorin) in salt marshes at the Támbrapathi river estuary, hence in a habitat somewhat resembling that of
<taxonomicName id="4C3D4D1BFFACFFC842A2FCF7FDAF834C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[426,516,829,852]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFACFFC842A2FCF7FDAF834C" box="[426,516,829,852]" italics="true" pageId="9" pageNumber="116">G. nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFACFFC84102FCF6FDF6834C" box="[522,605,828,852]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
The Indian species is very similar to
<taxonomicName id="4C3D4D1BFFACFFC840EFFCF7FBEA834C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[999,1089,829,852]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFACFFC840EFFCF7FBEA834C" box="[999,1089,829,852]" italics="true" pageId="9" pageNumber="116">G. nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFACFFC84379FC96FF6A836B" box="[113,193,860,883]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
by dorsal sculpture, male genitalia and highly reduced abdominal granulation, although it shows some weak differences from
<taxonomicName id="4C3D4D1BFFACFFC842F7FCB7FDF2838C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[511,601,893,916]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFACFFC842F7FCB7FDF2838C" box="[511,601,893,916]" italics="true" pageId="9" pageNumber="116">G. nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFACFFC84156FCB6FD1B838B" box="[606,688,892,915]" pageId="9" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
(elytral intervals 2 and 4 slightly convex, body coloration strongly metallic) and therefore seems to represent a separate species. Even though further studies of the Indian specimens are needed to completely understand their identity, the strong resemblance between both taxa suggests that
<taxonomicName id="4C3D4D1BFFACFFC8405CFC17FC0583EC" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[852,942,989,1012]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="116" phylum="Arthropoda" rank="species" species="nemo">
<emphasis id="B949EA8AFFACFFC8405CFC17FC0583EC" box="[852,942,989,1012]" italics="true" pageId="9" pageNumber="116">G. nemo</emphasis>
</taxonomicName>
may possibly represent a South Indian element of Socotran fauna.
</paragraph>
</subSubSection>
</treatment>
</document>