treatments-xml/data/03/94/87/0394878DFFB57B20198CFE31FD7BD9AF.xml
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<document ID-DOI="10.5252/geodiversitas2022v44a27" ID-ISSN="1638-9395" ID-ZooBank="urn:lsid:zoobank.org:pub:E4896081-9312-4EA6-AE33-AAC44201748E" checkinTime="1664917095605" checkinUser="admin" docAuthor="Sandoval, José" docDate="2022" docId="0394878DFFB57B20198CFE31FD7BD9AF" docLanguage="en" docName="Geodiversitas.44.27.801-851.pdf.imf" docOrigin="Geodiversitas 44 (27)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Witchelliinae Callomon &amp; Chandler 2006" docType="treatment" docVersion="4" lastPageNumber="822" masterDocId="FFADFFF5FFA27B381A52FFC7FFE0D81E" masterDocTitle="Sonniniidae Ammonitina, Middle Jurassic from Southern Spain: taxonomic, biostratigraphical and palaeobiogeographical analysis" masterLastPageNumber="851" masterPageNumber="801" pageNumber="822" updateTime="1665000352097" updateUser="diego">
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<mods:title>Sonniniidae Ammonitina, Middle Jurassic from Southern Spain: taxonomic, biostratigraphical and palaeobiogeographical analysis</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sandoval, José</mods:namePart>
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<mods:date>2022</mods:date>
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<mods:number>2022-09-15</mods:number>
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<mods:number>44</mods:number>
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<mods:identifier type="DOI">10.5252/geodiversitas2022v44a27</mods:identifier>
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<subSubSection pageId="23" pageNumber="822" type="nomenclature">
<paragraph blockId="23.[976,1292,502,560]" pageId="23" pageNumber="822">
<heading box="[990,1277,502,529]" centered="true" fontSize="9" level="2" pageId="23" pageNumber="822" reason="3">
Subfamily
<taxonomicName authorityName="Callomon &amp; Chandler" authorityYear="2006" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="subFamily" subFamily="Witchelliinae">
WITCHELLIINAE Callomon &amp;
<bibRefCitation author="CHANDLER R. B. &amp; CALLOMON J. H. &amp; KING A. &amp; JEFFREYS K. &amp; VARAH M. &amp; BENTLEY A." box="[1122,1292,534,560]" pageId="23" pageNumber="822" pagination="345 - 375" refId="ref40976" refString="CHANDLER R. B., CALLOMON J. H., KING A., JEFFREYS K., VARAH M. &amp; BENTLEY A. 2006. - The stratigraphy of the Inferior Oolite at South Main Road Quarry, Dundry, Avon. Proceedings of the Geologists' Association 117: 345 - 375. https: // doi. org / 10.1016 / S 0016 - 7878 (06) 80043 - X" type="journal article" year="2006">Chandler, 2006</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="822" type="reference_group">
<paragraph blockId="23.[811,1455,591,642]" pageId="23" pageNumber="822">
<treatmentCitationGroup pageId="23" pageNumber="822">
<taxonomicName authority="Callomon &amp; Chandler" authorityName="Callomon &amp; Chandler" box="[813,1164,591,615]" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="subFamily" subFamily="Witchelliinae">Witchelliinae Callomon &amp; Chandler</taxonomicName>
<emphasis box="[1167,1187,592,615]" italics="true" pageId="23" pageNumber="822">in</emphasis>
<treatmentCitation author="CHANDLER R. B. &amp; CALLOMON J. H. &amp; KING A. &amp; JEFFREYS K. &amp; VARAH M. &amp; BENTLEY A." page="370" pageId="23" pageNumber="822" year="2006">
<bibRefCitation author="CHANDLER R. B. &amp; CALLOMON J. H. &amp; KING A. &amp; JEFFREYS K. &amp; VARAH M. &amp; BENTLEY A." pageId="23" pageNumber="822" pagination="345 - 375" refId="ref40976" refString="CHANDLER R. B., CALLOMON J. H., KING A., JEFFREYS K., VARAH M. &amp; BENTLEY A. 2006. - The stratigraphy of the Inferior Oolite at South Main Road Quarry, Dundry, Avon. Proceedings of the Geologists' Association 117: 345 - 375. https: // doi. org / 10.1016 / S 0016 - 7878 (06) 80043 - X" type="journal article" year="2006">Chandler, Callomon, King, Jeffreys, Varah &amp; Bentley, 2006: 370</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="822" type="description">
<paragraph blockId="23.[811,1458,694,1262]" box="[813,957,694,720]" pageId="23" pageNumber="822">DESCRIPTION</paragraph>
<paragraph blockId="23.[811,1458,694,1262]" pageId="23" pageNumber="822">
Medium to relatively large macroconchs [M], but smaller than those of
<taxonomicName box="[961,1095,757,783]" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="840" phylum="Mollusca" rank="subFamily" subFamily="Sonniniinae">Sonniniinae</taxonomicName>
. Coiling is involute to moderately evolute, with a compressed whorl section, varying from typically rectangular or subtriangular, to even ovate or oxycone, with a more or less developed keel. The ribs, radial or slightly proverse and simple or divided near the umbilical edge, have irregular thickness and usually fade towards the outer whorls, which in some forms become completely smooth or the ornamentation is reduced to striae. The innermost whorls of some forms have small lateral tubercles. The aperture has plain mouth borders. Dimorphic microconchs [m], (until now usually nominated
<taxonomicName authorityName="Buckman" authorityYear="1923" box="[1021,1131,1076,1102]" genus="Pelekodites" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[1021,1131,1076,1102]" italics="true" pageId="23" pageNumber="822">Pelekodites</emphasis>
</taxonomicName>
), small adult at
<quantity box="[1309,1426,1076,1103]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="23" pageNumber="822" unit="mm" value="40.0" valueMax="60.0" valueMin="20.0">20-60 mm</quantity>
in diameter, are usually ribbed to the end of the BC and have final adult mouth borders with long, spatulate lappets in some forms. The dimorphic size ratio is modest, between 2:1 and 3:1. The septal suture is simple, with L wide and short; scarcely retracted umbilical lobe, and barely trimmed saddles.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="822" type="discussion">
<paragraph blockId="23.[811,1458,1300,1836]" box="[813,913,1300,1326]" pageId="23" pageNumber="822">REMARKS</paragraph>
<paragraph blockId="23.[811,1458,1300,1836]" pageId="23" pageNumber="822">
As indicated above,
<bibRefCitation author="HOWARTH M. K." box="[1042,1225,1331,1358]" pageId="23" pageNumber="822" refId="ref43758" refString="HOWARTH M. K. 2013. - Treatease Online No. 57, Part L, Revised, Volume 3 B, Chapter 4: Psiloceratoidea, Eodoceratoidea, Hildoceratoidea. The University of Kansas, Lawrence, Kansas, 139 p. https: // doi. org / 10.17161 / to. v 0 i 0.4441" type="book" year="2013">Howarth (2013)</bibRefCitation>
included the genera
<taxonomicName authority="Buckman, 1899" authorityName="Buckman" authorityYear="1899" box="[815,1127,1363,1389]" class="Cephalopoda" family="Sonniniidae" genus="Asthenoceras" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[815,943,1363,1389]" italics="true" pageId="23" pageNumber="822">Asthenoceras</emphasis>
Buckman, 1899
</taxonomicName>
,
<taxonomicName authority="Buckman, 1902" authorityName="Buckman" authorityYear="1902" box="[1141,1451,1363,1389]" class="Cephalopoda" family="Sonniniidae" genus="Fontannesia" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[1141,1266,1364,1389]" italics="true" pageId="23" pageNumber="822">Fontannesia</emphasis>
Buckman, 1902
</taxonomicName>
,
<taxonomicName authority="Hall, 1989" authorityName="Hall" authorityYear="1989" box="[814,1151,1395,1421]" class="Cephalopoda" family="Sonniniidae" genus="Newmarracarroceras" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[814,1025,1396,1421]" italics="true" pageId="23" pageNumber="822">Newmarracarroceras</emphasis>
<bibRefCitation author="HALL R. L." box="[1031,1151,1395,1421]" pageId="23" pageNumber="822" pagination="1 - 20" refId="ref43376" refString="HALL R. L. 1989. - Lower Bajocian ammonites (Middle Jurassic; Sonniniidae) from the Newmarracarra Limestone, Western Australia. Alcheringa 13: 1 - 20. https: // doi. org / 10.1080 / 03115518908619122" type="journal article" year="1989">Hall, 1989</bibRefCitation>
</taxonomicName>
,
<taxonomicName authority="Buckman, 1889" authorityName="Buckman" authorityYear="1889" box="[1161,1451,1395,1421]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[1161,1267,1395,1421]" italics="true" pageId="23" pageNumber="822">Witchellia</emphasis>
<bibRefCitation author="BUCKMAN S. S." box="[1275,1451,1395,1421]" pageId="23" pageNumber="822" pagination="651 - 663" refId="ref40442" refString="BUCKMAN S. S. 1889. - The descent of Sonninia and Hammatoceras. Quarterly Journal of the Geological Society of London 45: 651 - 663. https: // doi. org / 10.1144 / GSL. JGS. 1889.045.01 - 04.43" type="journal article" year="1889">Buckman, 1889</bibRefCitation>
</taxonomicName>
, and
<taxonomicName authority="McLearn, 1926" authorityName="McLearn" authorityYear="1926" box="[862,1141,1427,1454]" class="Cephalopoda" family="Sonniniidae" genus="Guhsania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[862,963,1427,1453]" italics="true" pageId="23" pageNumber="822">Guhsania</emphasis>
<bibRefCitation author="MCLEARN F. H." box="[971,1141,1427,1454]" pageId="23" pageNumber="822" pagination="136 - 149" refId="ref44644" refString="MCLEARN F. H. 1926. - New Jurassic species from the Hazelton Group of British Columbia. Bulletin of the Geological Survey of Canada 44: 89 - 99, 136 - 149. https: // doi. org / 10.4095 / 105021" type="journal article" year="1926">
McLearn,
<quantity box="[1085,1172,1427,1453]" metricMagnitude="1" metricUnit="m" metricValue="4.89204" pageId="23" pageNumber="822" unit="in" value="1926.0">1926 in</quantity>
</bibRefCitation>
</taxonomicName>
the subfamily
<taxonomicName class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="subFamily" subFamily="Witchelliinae">Witchelliinae</taxonomicName>
but placed
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[978,1096,1460,1485]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[978,1096,1460,1485]" italics="true" pageId="23" pageNumber="822">Dorsetensia</emphasis>
</taxonomicName>
in
<taxonomicName box="[1132,1266,1459,1485]" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="840" phylum="Mollusca" rank="subFamily" subFamily="Sonniniinae">Sonniniinae</taxonomicName>
. However, as also shown above,
<taxonomicName authorityName="Buckman" authorityYear="1899" box="[963,1091,1491,1517]" class="Cephalopoda" family="Sonniniidae" genus="Asthenoceras" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[963,1091,1491,1517]" italics="true" pageId="23" pageNumber="822">Asthenoceras</emphasis>
</taxonomicName>
and presumably also
<emphasis italics="true" pageId="23" pageNumber="822">
<taxonomicName authorityName="Buckman" authorityYear="1902" box="[1322,1450,1492,1517]" class="Cephalopoda" family="Sonniniidae" genus="Fontannesia" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">Fontannesia</taxonomicName>
,
<taxonomicName authorityName="Hall" authorityYear="1989" box="[813,1028,1523,1548]" class="Cephalopoda" family="Sonniniidae" genus="Newmarracarroceras" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">Newmarracarroceras</taxonomicName>
,
</emphasis>
and
<emphasis box="[1080,1224,1523,1549]" italics="true" pageId="23" pageNumber="822">Latiwitchellia</emphasis>
should be transferred to
<taxonomicName box="[838,1033,1554,1580]" higherTaxonomySource="Manual Input" pageId="23" pageNumber="807" rank="subFamily" subFamily="Grammoceratinae">Grammoceratinae</taxonomicName>
, as they coincide in coiling, ribbing, and
<typeStatus box="[811,857,1587,1613]" pageId="23" pageNumber="822">type</typeStatus>
of keel as well as presenting an extremely simple septal suture (see
<bibRefCitation author="SANDOVAL J. &amp; HENRIQUES M. H. &amp; CHANDLER R. B. &amp; URETA M." box="[942,1168,1618,1644]" pageId="23" pageNumber="822" pagination="109 - 219" refId="ref46889" refString="SANDOVAL J., HENRIQUES M. H., CHANDLER R. B. &amp; URETA M. 2012. - Latest Toarcian-earliest Bajocian (Jurassic) Grammoceratinae (Hildoceratidae, Ammonitina) of the western Tethys: their palaeobiogeographic and phylogenetic significance. Geobios 45: 109 - 219. https: // doi. org / 10.1016 / j. geobios. 2011.11.001" type="journal article" year="2012">
Sandoval
<emphasis box="[1048,1103,1618,1644]" italics="true" pageId="23" pageNumber="822">et al.</emphasis>
2012
</bibRefCitation>
). Nevertheless, the genus
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[815,935,1651,1676]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[815,935,1651,1676]" italics="true" pageId="23" pageNumber="822">Dorsetensia</emphasis>
</taxonomicName>
, which succeeds
<taxonomicName authorityName="Buckman" authorityYear="1889" box="[1134,1244,1650,1676]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[1134,1244,1650,1676]" italics="true" pageId="23" pageNumber="822">Witchellia</emphasis>
</taxonomicName>
in time, and from which it possibly descended, is included here in
<taxonomicName box="[1308,1454,1682,1708]" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="subFamily" subFamily="Witchelliinae">Witchelliinae</taxonomicName>
.
<taxonomicName authorityName="Buckman" authorityYear="1889" box="[811,917,1714,1740]" class="Cephalopoda" family="Sonniniidae" genus="Witchellia" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[811,917,1714,1740]" italics="true" pageId="23" pageNumber="822">Witchellia</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[970,1088,1715,1740]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[970,1088,1715,1740]" italics="true" pageId="23" pageNumber="822">Dorsetensia</emphasis>
</taxonomicName>
have a similar section, similar coiling and ornamentation, and also a comparable dimorphism. The slightly more complex septal suture of
<taxonomicName authorityName="Buckman" authorityYear="1892" box="[1273,1391,1779,1804]" class="Cephalopoda" family="Sonniniidae" genus="Dorsetensia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonoidea" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">
<emphasis box="[1273,1391,1779,1804]" italics="true" pageId="23" pageNumber="822">Dorsetensia</emphasis>
</taxonomicName>
is the only remarkable difference.
</paragraph>
</subSubSection>
<subSubSection lastPageId="24" lastPageNumber="823" pageId="23" pageNumber="822" type="distribution">
<paragraph blockId="23.[810,1456,1874,2028]" lastBlockId="24.[131,777,215,433]" lastPageId="24" lastPageNumber="823" pageId="23" pageNumber="822">
DISTRIBUTION Middle Jurassic (Lower Bajocian, Ovale Zone-Humphriesianum Zone): Europe (
<collectingCountry box="[989,1085,1937,1963]" name="United Kingdom" pageId="23" pageNumber="822">England</collectingCountry>
,
<collectingCountry box="[1098,1193,1937,1963]" name="Belgium" pageId="23" pageNumber="822">Belgium</collectingCountry>
,
<collectingCountry box="[1206,1300,1937,1963]" name="Bulgaria" pageId="23" pageNumber="822">Bulgaria</collectingCountry>
,
<collectingCountry box="[1313,1389,1938,1964]" name="France" pageId="23" pageNumber="822">France</collectingCountry>
,
<collectingCountry name="Germany" pageId="23" pageNumber="822">Germany</collectingCountry>
,
<collectingCountry box="[882,930,1969,1995]" name="Italy" pageId="23" pageNumber="822">Italy</collectingCountry>
,
<collectingCountry box="[940,1081,1969,1995]" name="Luxembourg" pageId="23" pageNumber="822">Luxembourg</collectingCountry>
,
<collectingCountry box="[1090,1167,1969,1995]" name="Poland" pageId="23" pageNumber="822">Poland</collectingCountry>
,
<collectingCountry box="[1176,1269,1969,1995]" name="Portugal" pageId="23" pageNumber="822">Portugal</collectingCountry>
,
<collectingCountry box="[1279,1380,1969,1995]" name="Romania" pageId="23" pageNumber="822">Romania</collectingCountry>
,
<collectingCountry box="[1388,1451,1969,1995]" name="Spain" pageId="23" pageNumber="822">Spain</collectingCountry>
, and
<collectingCountry box="[858,988,2001,2028]" name="Switzerland" pageId="23" pageNumber="822">Switzerland</collectingCountry>
), North Africa (
<taxonomicName authority="Ranges" authorityName="Ranges" box="[1159,1303,2001,2028]" genus="Atlas" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="822" phylum="Mollusca" rank="genus">Atlas Ranges</taxonomicName>
),
<collectingCountry box="[1321,1449,2001,2027]" name="Madagascar" pageId="23" pageNumber="822">Madagascar</collectingCountry>
, Caucasus,
<collectingCountry box="[254,303,216,242]" name="Iran" pageId="24" pageNumber="823">Iran</collectingCountry>
,
<collectingCountry box="[317,387,215,241]" name="China" pageId="24" pageNumber="823">China</collectingCountry>
(
<collectingRegion box="[404,471,215,242]" country="China" name="Xizang" pageId="24" pageNumber="823">Tibet</collectingRegion>
),
<collectingCountry box="[493,559,216,242]" name="Japan" pageId="24" pageNumber="823">Japan</collectingCountry>
, western Australia,
<collectingCountry box="[134,216,247,273]" name="Canada" pageId="24" pageNumber="823">Canada</collectingCountry>
(
<collectingRegion box="[229,422,247,274]" country="Canada" name="British Columbia" pageId="24" pageNumber="823">British Columbia</collectingRegion>
),
<collectingCountry box="[439,582,247,273]" name="United States of America" pageId="24" pageNumber="823">United States</collectingCountry>
(
<collectingRegion box="[594,669,247,273]" country="United States of America" name="Alaska" pageId="24" pageNumber="823">Alaska</collectingRegion>
,
<collectingRegion box="[678,765,247,273]" country="United States of America" name="Oregon" pageId="24" pageNumber="823">Oregon</collectingRegion>
), and
<collectingCountry box="[177,288,279,305]" name="Argentina" pageId="24" pageNumber="823">Argentina</collectingCountry>
. In the Subbetic domain,
<taxonomicName box="[564,708,279,305]" class="Cephalopoda" family="Sonniniidae" kingdom="Animalia" order="Ammonoidea" pageId="24" pageNumber="822" phylum="Mollusca" rank="subFamily" subFamily="Witchelliinae">Witchelliinae</taxonomicName>
spans the Lower Bajocian (Ovale, Laeviuscula Propinquans and Humphriesianum zones), being especially abundant in the Laeviuscula Zone of the central sector of the Median Subbetic, Sierra de Alta Coloma area (Jaén Province).
</paragraph>
</subSubSection>
</treatment>
</document>