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<mods:title id="34FC10F30E95A4CE7D68474C100C3F05">Taxonomy of the genus Chlamisus Rafinesque (Coleoptera: Chrysomelidae) from China with description of three new species</mods:title>
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<treatment id="0394878DFF0DE3146EB09A45FBE0F811" ID-DOI="http://doi.org/10.5281/zenodo.5686251" ID-GBIF-Taxon="127697293" ID-Zenodo-Dep="5686251" LSID="urn:lsid:plazi:treatment:0394878DFF0DE3146EB09A45FBE0F811" httpUri="http://treatment.plazi.org/id/0394878DFF0DE3146EB09A45FBE0F811" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="C3276510FF0DE3146EB09A45FE78FC21" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B82369BFF0DE3146EB09A45FD96FCA1" blockId="9.[151,588,797,823]" box="[151,588,797,823]" pageId="9" pageNumber="10">
<heading id="D0CA81F7FF0DE3146EB09A45FD96FCA1" bold="true" box="[151,588,797,823]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B949EA89FF0DE3146EB09A45FD96FCA1" bold="true" box="[151,588,797,823]" pageId="9" pageNumber="10">
Genus
<taxonomicName id="4C3D4D18FF0DE3146EC99A45FD96FCA1" ID-CoL="3NFJ" authority="Rafinesque, 1815" authorityName="Rafinesque" authorityYear="1815" box="[238,588,797,823]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146EC99A45FEB5FCA1" bold="true" box="[238,367,797,823]" italics="true" pageId="9" pageNumber="10">Chlamisus</emphasis>
Rafinesque, 1815
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B82369BFF0DE3146EB09A3CFE78FC21" blockId="9.[151,1436,868,1135]" pageId="9" pageNumber="10">
Rafinesque, 1815: 16;
<bibRefCitation id="EFAC4B6AFF0DE3146F5C9A3CFDD1FCEC" author="Gressitt" box="[379,523,868,890]" pageId="9" pageNumber="10" refString="Gressitt J. L. (1946) Chinese Chrysomelid Beetles of the subfamily Chlamisinae. Annals of the Entomological Society of America, 39, 84 - 100." type="journal article" year="1946">Gressitt, 1946</bibRefCitation>
: 84;
<bibRefCitation id="EFAC4B6AFF0DE3146C1C9A3CFD16FCEC" author="Monros" box="[571,716,868,890]" pageId="9" pageNumber="10" refString="Monros, F. (1949) Descripcion de seis nuevas Chlamisinae neotropicales (Col., Chrysomolidae. Revista de Entomologia Rio de Janeiro, 20, 617 - 629." type="journal article" year="1949">Monrós, 1949</bibRefCitation>
: 617; 1950: 412413; 1952: 540, 547549;
<bibRefCitation id="EFAC4B6AFF0DE3146AA39A3CFADEFCEC" author="Chujo" box="[1156,1284,868,890]" pageId="9" pageNumber="10" refString="Chujo, M. (1956) A taxonomic study on the Chrysomelidae (Insecta: Coleoptera) from Formosa IX. Subfamily Chlamisinae (1), (2). Technical Bulletin Kagawa Agricultural College, 7, 51 - 59." type="journal article" year="1956">Chûjô, 1956</bibRefCitation>
: 54;
<bibRefCitation id="EFAC4B6AFF0DE3146B139A3CFE80FC0E" author="Gressitt" pageId="9" pageNumber="10" refString="Gressitt, J. L. &amp; Kimoto, S. (1961) The Chrysomelidae (Coleopt.) of China and Korea. Part 1. Pacific Insects Monograph, 1 A, 1 - 299" type="journal article" year="1961">Gressitt &amp; Kimoto, 1961</bibRefCitation>
: 176189;
<bibRefCitation id="EFAC4B6AFF0DE3146FF59ADAFD36FC0E" author="Kimoto" box="[466,748,898,920]" pageId="9" pageNumber="10" refString="Kimoto, S. &amp; Gressitt, J. L. (1981) Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. 2. Clytrinae, Cryptocephalinae, Chlamisinae, Lamprosomatinae and Chrysomelinae. Pacific Insects, 23 (4), 286 - 391." type="journal article" year="1981">Kimoto and Gressitt, 1981</bibRefCitation>
: 387;
<bibRefCitation id="EFAC4B6AFF0DE3146D169ADAFB05FC0E" author="Chamorro-Lacayo" box="[817,1247,898,920]" pageId="9" pageNumber="10" refString="Chamorro-Lacayo, M. L. &amp; Konstantinov, A. S. (2009) Synopsis of warty leaf beetle genera of the World (Coleoptera, Chrysomelidae, Cryptocephalinae, Chlamisini). Zookeys, 8, 63 - 88." type="journal article" year="2009">Chamorro-Lacayo &amp; Konstantinov, 2009</bibRefCitation>
: 71 (difference in generic type species).
</paragraph>
</subSubSection>
<subSubSection id="C3276510FF0DE3146EB09A98FC23FB85" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8B82369BFF0DE3146EB09A98FC6CFC40" blockId="9.[151,1436,868,1135]" box="[151,950,960,982]" pageId="9" pageNumber="10">
<emphasis id="B949EA89FF0DE3146EB09A98FEC6FC40" bold="true" box="[151,284,960,982]" pageId="9" pageNumber="10">Type species</emphasis>
:
<taxonomicName id="4C3D4D18FF0DE3146F0D9A98FD59FC40" authority="Fabricius, 1777" authorityName="Fabricius" authorityYear="1777" box="[298,643,960,982]" class="Insecta" family="Chrysomelidae" genus="Bruchus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B949EA89FF0DE3146F0D9A98FE05FC40" box="[298,479,960,982]" italics="true" pageId="9" pageNumber="10">Bruchus gibbosus</emphasis>
Fabricius, 1777
</taxonomicName>
(=
<taxonomicName id="4C3D4D18FF0DE3146C819A98FC71FC40" authority="Knock" authorityName="Knock" box="[678,939,960,982]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinoida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="tuberosa">
<emphasis id="B949EA89FF0DE3146C819A98FC87FC40" box="[678,861,960,982]" italics="true" pageId="9" pageNumber="10">Chlamys tuberosa</emphasis>
Knock
</taxonomicName>
).
</paragraph>
<paragraph id="8B82369BFF0DE3146EB09A86FC23FB85" blockId="9.[151,1436,868,1135]" pageId="9" pageNumber="10">
<emphasis id="B949EA89FF0DE3146EB09A86FF14FC62" bold="true" box="[151,206,990,1012]" pageId="9" pageNumber="10">Syn.:</emphasis>
<taxonomicName id="4C3D4D18FF0DE3146EF29A86FE34FC62" authority="Knoch, 1801: 122" authorityName="Knoch" authorityPageNumber="122" authorityYear="1801" box="[213,494,990,1012]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinoida" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B949EA89FF0DE3146EF29A86FEF4FC62" box="[213,302,990,1012]" italics="true" pageId="9" pageNumber="10">Chlamys</emphasis>
<bibRefCitation id="EFAC4B6AFF0DE3146F139A86FE66FC62" author="Knoch" box="[308,444,990,1012]" pageId="9" pageNumber="10" refString="Knoch, A. W. (1801) Neue Beytrage zur Insectenkunde. Fol. 1. Schwinkert, Leipzig, 208 pp." type="book" year="1801">Knoch, 1801</bibRefCitation>
: 122
</taxonomicName>
; (nec Röding IN Bolten, 1798);
<bibRefCitation id="EFAC4B6AFF0DE3146D109A86FC1AFC62" author="Jacoby" box="[823,960,990,1012]" pageId="9" pageNumber="10" refString="Jacoby, M. (1908) The fauna of British India, including Ceylon and Burma. Coleoptera. Chrysomelidae. Fol. 1. Taylor and Francis, London, 534 pp." type="book" year="1908">Jacoby, 1908</bibRefCitation>
: 271 (type species);
<bibRefCitation id="EFAC4B6AFF0DE3146AB79A86FAD3FC62" author="Chen" box="[1168,1289,990,1012]" pageId="9" pageNumber="10" refString="Chen, S. H. (1940) On the Coleoptera Chlamydinae of China. Sinensia Nanking, 11, 189 - 206." type="journal article" year="1940">Chen, 1940</bibRefCitation>
: 192;
<bibRefCitation id="EFAC4B6AFF0DE3146B609A86FF27FB85" author="Gressitt" pageId="9" pageNumber="10" refString="Gressitt, J L. (1942) Plant-beetles from south and west China. I. Sagrinae, Donaciinae, Orsodacninae and Megalopodinae, II, Criocerinae (Chrysomelidae). III. Clytrinae, Cryptocephalinae and Chlamydinae (Coleoptera). Lingnan Science Journal, 20, 271 - 293." type="journal article" year="1942">Gressitt, 1942</bibRefCitation>
: 354. Type species:
<taxonomicName id="4C3D4D18FF0DE3146FEF9AA5FD3FFB85" authority="Fabricius" authorityName="Fabricius" box="[456,741,1021,1043]" class="Insecta" family="Chrysomelidae" genus="Bruchus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B949EA89FF0DE3146FEF9AA5FDA7FB85" box="[456,637,1021,1043]" italics="true" pageId="9" pageNumber="10">Bruchus gibbosus</emphasis>
Fabricius
</taxonomicName>
, designed by
<bibRefCitation id="EFAC4B6AFF0DE3146D499AA5FC2FFB85" author="Jacoby" box="[878,1013,1021,1043]" pageId="9" pageNumber="10" refString="Jacoby, M. (1908) The fauna of British India, including Ceylon and Burma. Coleoptera. Chrysomelidae. Fol. 1. Taylor and Francis, London, 534 pp." type="book" year="1908">Jacoby, 1908</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276510FF0DE3146EB09D44FC91FBF9" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8B82369BFF0DE3146EB09D44FB4EFBA4" blockId="9.[151,1436,868,1135]" box="[151,1172,1052,1074]" pageId="9" pageNumber="10">
<taxonomicName id="4C3D4D18FF0DE3146EB09D44FE03FBA4" authority="Ihering, 1905: 642" authorityName="Ihering" authorityPageNumber="642" authorityYear="1905" box="[151,473,1052,1074]" class="Insecta" family="Chrysomelidae" genus="Myochlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146EB09D44FECDFBA4" box="[151,279,1052,1074]" italics="true" pageId="9" pageNumber="10">Myochlamys</emphasis>
Ihering, 1905: 642
</taxonomicName>
(nec Fairmaire, 1876; replacement name for
<taxonomicName id="4C3D4D18FF0DE3146D849D44FB52FBA4" authority="Knoch, 1801" authorityName="Knoch" authorityYear="1801" box="[931,1160,1052,1074]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinoida" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B949EA89FF0DE3146D849D44FC26FBA4" box="[931,1020,1052,1074]" italics="true" pageId="9" pageNumber="10">Chlamys</emphasis>
<bibRefCitation id="EFAC4B6AFF0DE3146A259D44FB52FBA4" author="Knoch" box="[1026,1160,1052,1074]" pageId="9" pageNumber="10" refString="Knoch, A. W. (1801) Neue Beytrage zur Insectenkunde. Fol. 1. Schwinkert, Leipzig, 208 pp." type="book" year="1801">Knoch, 1801</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8B82369BFF0DE3146EB09D62FC24FBC6" blockId="9.[151,1436,868,1135]" box="[151,1022,1082,1104]" pageId="9" pageNumber="10">
<taxonomicName id="4C3D4D18FF0DE3146EB09D62FE34FBC6" authority="Ihering, 1907: 251" authorityName="Ihering" authorityPageNumber="251" authorityYear="1907" box="[151,494,1082,1104]" class="Insecta" family="Chrysomelidae" genus="Arthrochlamys" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146EB09D62FEF7FBC6" box="[151,301,1082,1104]" italics="true" pageId="9" pageNumber="10">Arthrochlamys</emphasis>
Ihering, 1907: 251
</taxonomicName>
(replacement name for
<taxonomicName id="4C3D4D18FF0DE3146CC79D62FC28FBC6" authority="Ihering, 1905" authorityName="Ihering" authorityYear="1905" box="[736,1010,1082,1104]" class="Insecta" family="Chrysomelidae" genus="Myochlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146CC79D62FCBAFBC6" box="[736,864,1082,1104]" italics="true" pageId="9" pageNumber="10">Myochlamys</emphasis>
Ihering, 1905
</taxonomicName>
).
</paragraph>
<paragraph id="8B82369BFF0DE3146EB09D01FC91FBF9" blockId="9.[151,1436,868,1135]" box="[151,843,1113,1135]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB2D11B5FF0DE3146EB09D01FC91FBF9" box="[151,843,1113,1135]" pageId="9" pageNumber="10">
<taxonomicName id="4C3D4D18FF0DE3146EB09D01FE3EFBF9" authority="Jacobson, 1924: 239" authorityName="Jacobson" authorityPageNumber="239" authorityYear="1924" box="[151,484,1113,1135]" class="Insecta" family="Chrysomelidae" genus="Boloschesis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146EB09D01FED5FBF9" box="[151,271,1113,1135]" italics="true" pageId="9" pageNumber="10">Boloschesis</emphasis>
<treatmentCitation id="0A9C108AFF0DE3146F339D01FE3EFBF9" author="Jacobson" box="[276,484,1113,1135]" page="239" pageId="9" pageNumber="10" year="1924">
<bibRefCitation id="EFAC4B6AFF0DE3146F339D01FE6EFBF9" author="Jacobson" box="[276,436,1113,1135]" pageId="9" pageNumber="10" refString="Jacobson, G. (1924) Annotationes synonymicae et systematicae de Coleopteris. Revue Russe d'Entomologie, 18, 237 - 244." type="journal article" year="1924">Jacobson, 1924</bibRefCitation>
: 239
</treatmentCitation>
</taxonomicName>
(name for
<taxonomicName id="4C3D4D18FF0DE3146C7D9D01FCE5FBF9" authority="Knoch, 1801" authorityName="Knoch" authorityYear="1801" box="[602,831,1113,1135]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinoida" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B949EA89FF0DE3146C7D9D01FD69FBF9" box="[602,691,1113,1135]" italics="true" pageId="9" pageNumber="10">Chlamys</emphasis>
<bibRefCitation id="EFAC4B6AFF0DE3146C9E9D01FCE5FBF9" author="Knoch" box="[697,831,1113,1135]" pageId="9" pageNumber="10" refString="Knoch, A. W. (1801) Neue Beytrage zur Insectenkunde. Fol. 1. Schwinkert, Leipzig, 208 pp." type="book" year="1801">Knoch, 1801</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3276510FF0DE3146EB09DC4FBE0F811" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8B82369BFF0DE3146EB09DC4FBF2F91D" blockId="9.[151,1437,1180,1927]" pageId="9" pageNumber="10">
<emphasis id="B949EA89FF0DE3146EB09DC4FECBFB23" bold="true" box="[151,273,1180,1205]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Body subpentagonal or subcylindrical, mostly black or dark, some red to yellow. Body surface with tubercles and ridges, very similar to feces of moth larva (Lepidoptera). Head embed on prothorax and head surface flat; eyes large, inner margin deeply concaved; frons and clypeus without distinct boarder; antennae concealed into the cavity between prosternum and propleuron, short and narrow, composed of 11 segments: scape club like, narrowed at base, pedicel usually globular or triangular, 3rd and 4th segment usually short and slender, apical six to eight segments serrated. Pronotum with disc strongly raised, usually with some ridges and tubercles at the surface; posterior declivity extending back, with a triangular notch in the middle, which conjunct with the triangular projection of base of scutellum. Prosternum broad anteriorly, prosternal process usually narrowly extended. Scutellum rectangular with posterior lateral angle produced. Elytra slightly wider or as wide as pronotum, narrower on apex, humeri swelled and arched; usually with four rows of tubercles and/or ridges with their names, from suture to outside respectively: sutural row, median row, humeral row, lateral row; a large longitudinal depression below humeri; sutural margin of each elytron with a row of teeth, which is complement for each other when elytra closed. Pygidium exposed, usually with one to three longitudinal carina, sometimes with a median transverse carina. Legs short and flat, fits into the groove on bottom of thorax and abdomen.
</paragraph>
<paragraph id="8B82369BFF0DE3146EE09FCEFBE0F811" blockId="9.[151,1437,1180,1927]" pageId="9" pageNumber="10">
This genus is closely related to the genus
<taxonomicName id="4C3D4D18FF0DE3146CBD9FC0FC76F939" authority="Lacordaire, 1848" authorityName="Lacordaire" authorityYear="1848" box="[666,940,1686,1711]" class="Insecta" family="Chrysomelidae" genus="Exema" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146CBD9FC0FD3CF939" box="[666,742,1688,1711]" italics="true" pageId="9" pageNumber="10">Exema</emphasis>
<bibRefCitation id="EFAC4B6AFF0DE3146CCA9FCEFC76F939" author="Lacordaire" box="[749,940,1686,1711]" pageId="9" pageNumber="10" refString="Lacordaire, M. T. (1848) Monographie des Coleopteres subpentameres de la famille des Phytophages. Tome Second. Memoires de la Societe Royale des Sciences de Liege, 5, i - vi + 1 - 890." type="journal article" year="1848">Lacordaire, 1848</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C3D4D18FF0DE3146DC49FCFFAC7F939" authority="Karren, 1972" authorityName="Karren" authorityYear="1972" box="[995,1309,1687,1711]" class="Insecta" family="Chrysomelidae" genus="Neochlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146DC49FCFFB5AF939" box="[995,1152,1687,1711]" italics="true" pageId="9" pageNumber="10">Neochlamisus</emphasis>
<bibRefCitation id="EFAC4B6AFF0DE3146AA09FC0FAC7F939" author="Karren" box="[1159,1309,1687,1711]" pageId="9" pageNumber="10" refString="Karren, J. B. (1972) A revision of the subfamily Chlamisinae of America north of Mexico (Coleoptera: Chrysomelidae). The University of Kansas Science Bulletin, 49, 875 - 988." type="journal article" year="1972">Karren, 1972</bibRefCitation>
</taxonomicName>
, but can be distinguished by the following characteristics: body mostly subpentagonal, usually not metallic, canthus of eyes usually without yellow spots, prosternum posteriorly much narrower than anterior margin, elytra without velvety spots, males without spines or spinulae on first ventrite, male ejaculatory guide symmetrical, without sheath, apex of spermathecal duct as wide as rest of pump. The genus
<taxonomicName id="4C3D4D18FF0DE3146D019E70FBE1F8A9" authority="Lacordaire, 1848" authorityName="Lacordaire" authorityYear="1848" box="[806,1083,1830,1855]" class="Insecta" family="Chrysomelidae" genus="Exema" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146D019E70FCA9F8A9" box="[806,883,1832,1855]" italics="true" pageId="9" pageNumber="10">Exema</emphasis>
<bibRefCitation id="EFAC4B6AFF0DE3146D5D9E7EFBE1F8A9" author="Lacordaire" box="[890,1083,1830,1855]" pageId="9" pageNumber="10" refString="Lacordaire, M. T. (1848) Monographie des Coleopteres subpentameres de la famille des Phytophages. Tome Second. Memoires de la Societe Royale des Sciences de Liege, 5, i - vi + 1 - 890." type="journal article" year="1848">Lacordaire, 1848</bibRefCitation>
</taxonomicName>
was treated as synonym of the genus
<taxonomicName id="4C3D4D18FF0DE3146EC79E13FDFEF8F5" authority="Rafinesque, 1815" authorityName="Rafinesque" authorityYear="1815" box="[224,548,1866,1891]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B949EA89FF0DE3146EC79E13FE8CF8F5" box="[224,342,1867,1891]" italics="true" pageId="9" pageNumber="10">Chlamisus</emphasis>
Rafinesque, 1815
</taxonomicName>
by most taxonomists (see references given for the genus), but was considered as valid by
<bibRefCitation id="EFAC4B6AFF0DE3146F309E37FDF8F811" author="Seeno" box="[279,546,1902,1927]" pageId="9" pageNumber="10" refString="Seeno, T. N. &amp; Wilcox, J. A. (1982) Leaf Beetle Genera (Coleoptera: Chrysomelidae). Entomography, 1, 1 - 222." type="journal article" year="1982">Seeno &amp; Wilcox (1982)</bibRefCitation>
and
<bibRefCitation id="EFAC4B6AFF0DE3146C7E9E36FBECF811" author="Chamorro-Lacayo" box="[601,1078,1902,1927]" pageId="9" pageNumber="10" refString="Chamorro-Lacayo, M. L. &amp; Konstantinov, A. S. (2009) Synopsis of warty leaf beetle genera of the World (Coleoptera, Chrysomelidae, Cryptocephalinae, Chlamisini). Zookeys, 8, 63 - 88." type="journal article" year="2009">Chamorro-Lacayo &amp; Konstantinov (2009)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>