168 lines
23 KiB
XML
168 lines
23 KiB
XML
<document id="1ED88990E29F235A382198B0C98CA453" ID-DOI="10.5281/zenodo.4665621" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617712844033" checkinUser="felipe" docAuthor="Venczel, Márton & Stiuc, Emanoil" docDate="2008" docId="03D08790FFD9FFF45615A66BFCB45555" docLanguage="en" docName="Geodiversitas.30.4.731-763.pdf" docOrigin="Geodiversitas 30 (4)" docStyle="DocumentStyle:9DCE1246056655C2F9CBE9822E831E4E.3:Geodiversitas.2000-2008.journal_article.type3" docStyleId="9DCE1246056655C2F9CBE9822E831E4E" docStyleName="Geodiversitas.2000-2008.journal_article.type3" docStyleVersion="3" docTitle="Natrix rudabanyaensis Szyndlar 2005" docType="treatment" docVersion="3" lastPageNumber="759" masterDocId="FFE9FFE8FFC5FFE85703A736FFA55105" masterDocTitle="Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania" masterLastPageNumber="763" masterPageNumber="731" pageNumber="759" updateTime="1698901057390" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="2102434AAB10BA962475D47701BEB6D7">
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<mods:title id="0D6CF306AE318A33FD93D62BC0CE71FE">Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania</mods:title>
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</mods:titleInfo>
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<mods:name id="498F8354CD57039C8DE029F0BC52E021" type="personal">
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<mods:role id="50D37BBBCA168D899BC86A6444B76266">
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<mods:roleTerm id="2DC376C14324EDAC8E61B83B0B71C719">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="798FC2E67D515521ADA00A71BDD83060">Venczel, Márton</mods:namePart>
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<mods:affiliation id="DD78A52ECB8441FD62A4E05F1AFB2E82">Tării ‚ Crişurilor Museum, B-dul Dacia 1 - 3, RO- 410464 Oradea (Romania) mvenczel @ rdslink. ro</mods:affiliation>
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<mods:nameIdentifier id="B73FCA7C06EEBA07DFAE351BAD248100" type="email">mvenczel@rdslink.ro</mods:nameIdentifier>
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<mods:name id="D8CE1B41D1C812E922C65DC4ED0C65F1" type="personal">
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<mods:roleTerm id="201D754C1BC481D3849C9D047F1139D8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FAB671B7390F9E84C87D76F2B43BDEE4">Stiuc, Emanoil</mods:namePart>
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<mods:affiliation id="53101B1BBC5C14D7EC65B222A4762B8B">Speleological Institute “ Emil Racovit, ă ”, Calea 13 Septembrie nr. 13, RO- 050711 Bucureşti (Romania) stiucaemil @ yahoo. com.</mods:affiliation>
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<mods:nameIdentifier id="DAD0D525205EFB0F39AE084ACAF97FED" type="email">stiucaemil@yahoo.com</mods:nameIdentifier>
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<mods:title id="E3ED78634EC2B2874C89C620FF2B9B4E">Geodiversitas</mods:title>
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<mods:part id="EF63B8321978CF4F515511D1354B56F3">
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<mods:date id="8A57E30ACBC0CD53FDA53C975B96ECA4">2008</mods:date>
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<mods:number id="D4C2E1432DEF961589BAD3D0BB9AAFCA">30</mods:number>
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<mods:number id="A26DFB84B786EF046D37291844DECD0B">4</mods:number>
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<mods:identifier id="BC46DD5C14471F94246FF3F02D2C2E10" type="DOI">10.5281/zenodo.4665621</mods:identifier>
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<mods:identifier id="28053F28F6A1DE29814BC6DAAD7F13E6" type="ISSN">1638-9395</mods:identifier>
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<subSubSection id="C363650DFFD9FFF45615A66BFE405092" pageId="28" pageNumber="759" type="nomenclature">
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<paragraph id="8BC63686FFD9FFF45615A66BFE405092" blockId="28.[278,558,348,407]" pageId="28" pageNumber="759">
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<heading id="D08E81EAFFD9FFF45615A66BFE405092" centered="true" fontSize="11" level="2" pageId="28" pageNumber="759" reason="2">
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<taxonomicName id="4C794D05FFD9FFF45615A66BFD8B5072" authorityName="Szyndlar" authorityYear="2005" box="[278,558,348,376]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="rudabanyaensis">
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<emphasis id="B90DEA94FFD9FFF45615A66BFEFB5072" bold="true" box="[278,350,349,375]" italics="true" pageId="28" pageNumber="759">Natrix</emphasis>
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cf.
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<emphasis id="B90DEA94FFD9FFF45685A66AFD8B5072" bold="true" box="[390,558,348,375]" italics="true" pageId="28" pageNumber="759">rudabanyaensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13422A03FFD9FFF45665A64AFE155092" box="[358,432,380,407]" captionStart="FIG" captionStartId="27.[151,162,1127,1144]" captionTargetBox="[198,1208,257,1093]" captionTargetId="figure-15@27.[155,1245,234,1128]" captionTargetPageId="27" captionText="FIG. 16. — A-F’, Natrix cf. rudabanyaensis; A, quadrate (ISER Tt-0502); B, B’, presacral vertebra (ISER Tt-0503/1); C, same (ISER Tt-0503/2); D, same (ISER Tt-0503/3); E, same (ISER Tt-0503/4); F, F’, posterior presacral vertebra (ISER Tt-0503/5); G, Macrovipera sp., presacral vertebra (ISER Tt-0504); A, B, D, F, dorsal views; B’, G, right lateral views;C, E, ventral views; F’, left lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665665" httpUri="https://zenodo.org/record/4665665/files/figure.png" pageId="28" pageNumber="759">Fig. 16</figureCitation>
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A-F)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C363650DFFD9FFF45794A680FDEC50ED" pageId="28" pageNumber="759" type="materials_examined">
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<paragraph id="8BC63686FFD9FFF45794A680FDEC50ED" blockId="28.[149,684,437,488]" pageId="28" pageNumber="759">
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MATERIAL EXAMINED. — One quadrate (
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<collectionCode id="ED68AE43FFD9FFF45517A683FDEC50C8" box="[532,585,437,461]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="28" pageNumber="759">ISER</collectionCode>
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Tt-0502), 30 presacral vertebrae (
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<collectionCode id="ED68AE43FFD9FFF45676A6E6FE0E50ED" box="[373,427,464,488]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="28" pageNumber="759">ISER</collectionCode>
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Tt-0503/1-30).
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</paragraph>
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</subSubSection>
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<subSubSection id="C363650DFFD9FFF45794A52DFC6D505D" pageId="28" pageNumber="759" type="description">
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<paragraph id="8BC63686FFD9FFF45794A52DFE8D5336" blockId="28.[149,686,539,883]" box="[151,296,539,565]" pageId="28" pageNumber="759">DESCRIPTION</paragraph>
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<paragraph id="8BC63686FFD9FFF45794A50CFF5F5351" blockId="28.[149,686,539,883]" box="[151,250,570,596]" pageId="28" pageNumber="759">
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<emphasis id="B90DEA94FFD9FFF45794A50CFF5F5351" box="[151,250,570,596]" italics="true" pageId="28" pageNumber="759">Quadrate</emphasis>
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</paragraph>
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<paragraph id="8BC63686FFD9FFF45794A56CFEE65276" blockId="28.[149,686,539,883]" pageId="28" pageNumber="759">
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ISER Tt-0502 is slightly damaged at its anterodorsal margin; it might have belonged to a rather small individual (
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<figureCitation id="13422A03FFD9FFF4561FA5AFFE2153B1" box="[284,388,665,692]" captionStart="FIG" captionStartId="27.[151,162,1127,1144]" captionTargetBox="[198,1208,257,1093]" captionTargetId="figure-15@27.[155,1245,234,1128]" captionTargetPageId="27" captionText="FIG. 16. — A-F’, Natrix cf. rudabanyaensis; A, quadrate (ISER Tt-0502); B, B’, presacral vertebra (ISER Tt-0503/1); C, same (ISER Tt-0503/2); D, same (ISER Tt-0503/3); E, same (ISER Tt-0503/4); F, F’, posterior presacral vertebra (ISER Tt-0503/5); G, Macrovipera sp., presacral vertebra (ISER Tt-0504); A, B, D, F, dorsal views; B’, G, right lateral views;C, E, ventral views; F’, left lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665665" httpUri="https://zenodo.org/record/4665665/files/figure.png" pageId="28" pageNumber="759">Fig. 16A</figureCitation>
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). In lateral view, the bone widens distally while the quadrate crest is better defined above the
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<emphasis id="B90DEA94FFD9FFF45661A5EFFDB353F6" box="[354,534,729,755]" italics="true" pageId="28" pageNumber="759">trochlea quadrati</emphasis>
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. Dorsally the height of the crest is abruptly reduced merging into the anterior margin of the bone. In posterolateral view the bone is straight, somewhat thinner in the middle portion.
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</paragraph>
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<paragraph id="8BC63686FFD9FFF45794A4A1FF5252B4" blockId="28.[149,686,919,1706]" box="[151,247,919,945]" pageId="28" pageNumber="759">
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<emphasis id="B90DEA94FFD9FFF45794A4A1FF5252B4" box="[151,247,919,945]" italics="true" pageId="28" pageNumber="759">Vertebrae</emphasis>
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</paragraph>
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<paragraph id="8BC63686FFD9FFF45794A481FD0957AF" blockId="28.[149,686,919,1706]" pageId="28" pageNumber="759">
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The vertebral centrum is elongated and provided with a sigmoid hypapophysis. The distal portion of the hypapophysis strongly projects beyond the condyle and displays a rounded tip (
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<figureCitation id="13422A03FFD9FFF45519A320FDDF5534" box="[538,634,1046,1073]" captionStart="FIG" captionStartId="27.[151,162,1127,1144]" captionTargetBox="[198,1208,257,1093]" captionTargetId="figure-15@27.[155,1245,234,1128]" captionTargetPageId="27" captionText="FIG. 16. — A-F’, Natrix cf. rudabanyaensis; A, quadrate (ISER Tt-0502); B, B’, presacral vertebra (ISER Tt-0503/1); C, same (ISER Tt-0503/2); D, same (ISER Tt-0503/3); E, same (ISER Tt-0503/4); F, F’, posterior presacral vertebra (ISER Tt-0503/5); G, Macrovipera sp., presacral vertebra (ISER Tt-0504); A, B, D, F, dorsal views; B’, G, right lateral views;C, E, ventral views; F’, left lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665665" httpUri="https://zenodo.org/record/4665665/files/figure.png" pageId="28" pageNumber="759">Fig. 16B</figureCitation>
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, B’). The subcentral ridge is rather prominent posteriorly to parapophyses (
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<figureCitation id="13422A03FFD9FFF45651A363FE705575" box="[338,469,1109,1136]" captionStart="FIG" captionStartId="27.[151,162,1127,1144]" captionTargetBox="[198,1208,257,1093]" captionTargetId="figure-15@27.[155,1245,234,1128]" captionTargetPageId="27" captionText="FIG. 16. — A-F’, Natrix cf. rudabanyaensis; A, quadrate (ISER Tt-0502); B, B’, presacral vertebra (ISER Tt-0503/1); C, same (ISER Tt-0503/2); D, same (ISER Tt-0503/3); E, same (ISER Tt-0503/4); F, F’, posterior presacral vertebra (ISER Tt-0503/5); G, Macrovipera sp., presacral vertebra (ISER Tt-0504); A, B, D, F, dorsal views; B’, G, right lateral views;C, E, ventral views; F’, left lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665665" httpUri="https://zenodo.org/record/4665665/files/figure.png" pageId="28" pageNumber="759">Fig. 16C, E</figureCitation>
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). The neural arch is moderately depressed, provided with large and posterolaterally projecting epizygapohyseal spines (
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<figureCitation id="13422A03FFD9FFF4579CA382FEB855CA" box="[159,285,1204,1231]" captionStart="FIG" captionStartId="27.[151,162,1127,1144]" captionTargetBox="[198,1208,257,1093]" captionTargetId="figure-15@27.[155,1245,234,1128]" captionTargetPageId="27" captionText="FIG. 16. — A-F’, Natrix cf. rudabanyaensis; A, quadrate (ISER Tt-0502); B, B’, presacral vertebra (ISER Tt-0503/1); C, same (ISER Tt-0503/2); D, same (ISER Tt-0503/3); E, same (ISER Tt-0503/4); F, F’, posterior presacral vertebra (ISER Tt-0503/5); G, Macrovipera sp., presacral vertebra (ISER Tt-0504); A, B, D, F, dorsal views; B’, G, right lateral views;C, E, ventral views; F’, left lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665665" httpUri="https://zenodo.org/record/4665665/files/figure.png" pageId="28" pageNumber="759">Fig. 16B, D</figureCitation>
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). In most specimens the neural spine is strongly damaged. However, small remnants of this structure indicate that the neural spine was longer than high. The anterior margin of the zygosphene is crenate with three distinct lobes. The prezygapophyseal articular facets are of oval shape, while the prezygapophyseal processes are moderately long, flattened dorsoventrally and obtuse distally.The paradiapophyses are well differentiated into diapophyseal and parapophyseal portions. The parapophyses are provided with a rather long parapophyseal process which has usually an obtuse anterior margin. In 12 measured presacral vertebrae the centrum length ranges between 2.84 and
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<quantity id="4C819B63FFD9FFF456FCA166FDC2576E" box="[511,615,1616,1643]" metricMagnitude="-3" metricUnit="m" metricValue="4.86" pageId="28" pageNumber="759" unit="mm" value="4.86">4.86 mm</quantity>
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, while the centrum width is between 1.72 and
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<quantity id="4C819B63FFD9FFF4554DA146FD0C578E" box="[590,681,1648,1675]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="28" pageNumber="759" unit="mm" value="3.3">3.3 mm</quantity>
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. The centrum length/centrum width ratio range
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</paragraph>
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<paragraph id="8BC63686FFD9FFF4579AA199FD0E57CF" blockId="28.[153,683,1711,1738]" box="[153,683,1711,1738]" pageId="28" pageNumber="759">is 1.34-1.65 (mean = 1.5). The vertebraefrom</paragraph>
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<paragraph id="8BC63686FFD9FFF455E4A742FB6D5180" blockId="28.[743,1224,115,136]" box="[743,1224,115,136]" pageId="28" pageNumber="759">
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||
Late middle Miocene amphibians and reptiles from Taut,,
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||
<collectingCountry id="F36E7616FFD9FFF45384A745FB6D5180" box="[1159,1224,115,133]" name="Romania" pageId="28" pageNumber="759">Romania</collectingCountry>
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</paragraph>
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<paragraph id="8BC63686FFD9FFF455CFA7C8FC6D505D" blockId="28.[714,1249,254,344]" pageId="28" pageNumber="759">
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the posterior presacral region are more depressed dorsoventrally and provided with a low and long neural spine (
|
||
<figureCitation id="13422A03FFD9FFF4545FA60BFC19505D" box="[860,956,317,344]" captionStart="FIG" captionStartId="27.[151,162,1127,1144]" captionTargetBox="[198,1208,257,1093]" captionTargetId="figure-15@27.[155,1245,234,1128]" captionTargetPageId="27" captionText="FIG. 16. — A-F’, Natrix cf. rudabanyaensis; A, quadrate (ISER Tt-0502); B, B’, presacral vertebra (ISER Tt-0503/1); C, same (ISER Tt-0503/2); D, same (ISER Tt-0503/3); E, same (ISER Tt-0503/4); F, F’, posterior presacral vertebra (ISER Tt-0503/5); G, Macrovipera sp., presacral vertebra (ISER Tt-0504); A, B, D, F, dorsal views; B’, G, right lateral views;C, E, ventral views; F’, left lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665665" httpUri="https://zenodo.org/record/4665665/files/figure.png" pageId="28" pageNumber="759">Fig. 16F</figureCitation>
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).
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C363650DFFD9FFF455C9A64BFCB45555" pageId="28" pageNumber="759" type="discussion">
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||
<paragraph id="8BC63686FFD9FFF455C9A64BFC8B5090" blockId="28.[712,1249,381,1104]" box="[714,814,381,407]" pageId="28" pageNumber="759">REMARKS</paragraph>
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<paragraph id="8BC63686FFD9FFF455C9A6AAFCB45555" blockId="28.[712,1249,381,1104]" pageId="28" pageNumber="759">
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The fossils from Tauţ closely resemble
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<taxonomicName id="4C794D05FFD9FFF45358A6ABFC7A50D2" authority="Szyndlar, 2005" authorityName="Szyndlar" authorityYear="2005" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="rudabanyaensis">
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<emphasis id="B90DEA94FFD9FFF45358A6ABFC9350D3" italics="true" pageId="28" pageNumber="759">Natrix rudabanyaensis</emphasis>
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||
<bibRefCitation id="EFE84B77FFD9FFF4543DA68AFC7A50D2" author="SZYNDLAR Z." box="[830,991,444,471]" pageId="28" pageNumber="759" pagination="31 - 52" refId="ref18137" refString="SZYNDLAR Z. 2005. - Snake fauna from the late Miocene of Rudabanya. Palaeontographia Italica 90: 31 - 52." type="journal article" year="2005">Szyndlar, 2005</bibRefCitation>
|
||
</taxonomicName>
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||
by their relatively small size, moderately developed subcentral ridges, round- ed distal hypapophyseal tip and similar centrum length/centrum width ratio (
|
||
<bibRefCitation id="EFE84B77FFD9FFF4530EA52DFB115333" author="SZYNDLAR Z." box="[1037,1204,539,566]" pageId="28" pageNumber="759" pagination="31 - 52" refId="ref18137" refString="SZYNDLAR Z. 2005. - Snake fauna from the late Miocene of Rudabanya. Palaeontographia Italica 90: 31 - 52." type="journal article" year="2005">Szyndlar 2005</bibRefCitation>
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||
). In
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<taxonomicName id="4C794D05FFD9FFF455CFA50DFC5A5350" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[716,1023,570,597]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="natrix">
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<emphasis id="B90DEA94FFD9FFF455CFA50DFC915351" box="[716,820,571,596]" italics="true" pageId="28" pageNumber="759">N. natrix</emphasis>
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(Linnaeus, 1758)
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</taxonomicName>
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and
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<taxonomicName id="4C794D05FFD9FFF45338A50DFCF75370" authority="Ivanov, 2002" authorityName="Ivanov" authorityYear="2002" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="merkurensis">
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<emphasis id="B90DEA94FFD9FFF45338A50DFB7A5351" box="[1083,1247,570,596]" italics="true" pageId="28" pageNumber="759">N. merkurensis</emphasis>
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Ivanov, 2002
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</taxonomicName>
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, in both of which the hypapophysis has a rounded tip, the centrum length/centrum width ratio is distinctly higher. In several other members of the genus the hypapophysis is provided with a pointed distal tip (e.g.,
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<taxonomicName id="4C794D05FFD9FFF454BDA5ECFCA15211" authority="Szyndlar, 1984" authorityName="Szyndlar" authorityYear="1984" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="longivertebrata">
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<emphasis id="B90DEA94FFD9FFF454BDA5ECFBDD53F6" box="[958,1144,729,755]" italics="true" pageId="28" pageNumber="759">N. longivertebrata</emphasis>
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Szyndlar, 1984
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794D05FFD9FFF45411A5CFFBE45211" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[786,1089,761,788]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="maura">
|
||
<emphasis id="B90DEA94FFD9FFF45411A5CFFCDE5217" box="[786,891,761,786]" italics="true" pageId="28" pageNumber="759">N. maura</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794D05FFD9FFF4534CA5CFFC235236" authority="(Lartet, 1851)" baseAuthorityName="Lartet" baseAuthorityYear="1851" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="sansaniensis">
|
||
<emphasis id="B90DEA94FFD9FFF4534CA5CFFD455237" italics="true" pageId="28" pageNumber="759">N. sansaniensis</emphasis>
|
||
(Lartet, 1851)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794D05FFD9FFF45497A42FFB7E5236" authority="(Laurenti, 1768))" authorityName="" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[916,1243,792,819]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="tessellata">
|
||
<emphasis id="B90DEA94FFD9FFF45497A42FFBB45237" box="[916,1041,792,818]" italics="true" pageId="28" pageNumber="759">N. tessellata</emphasis>
|
||
(Laurenti, 1768))
|
||
</taxonomicName>
|
||
, and with variable centrum length/centrum width ratio (
|
||
<bibRefCitation id="EFE84B77FFD9FFF45413A46EFC195276" author="SZYNDLAR Z." box="[784,956,856,883]" pageId="28" pageNumber="759" pagination="31 - 52" refId="ref18137" refString="SZYNDLAR Z. 2005. - Snake fauna from the late Miocene of Rudabanya. Palaeontographia Italica 90: 31 - 52." type="journal article" year="2005">Szyndlar 2005</bibRefCitation>
|
||
). In
|
||
<taxonomicName id="4C794D05FFD9FFF454F7A46EFC3B5297" authority="Auge & Rage, 2000" authorityName="Auge & Rage" authorityYear="2000" class="Reptilia" family="Colubridae" genus="Neonatrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="species" species="natricoides">
|
||
<emphasis id="B90DEA94FFD9FFF454F7A46EFB7A5274" box="[1012,1247,855,881]" italics="true" pageId="28" pageNumber="759">Neonatrix natricoides</emphasis>
|
||
<bibRefCitation id="EFE84B77FFD9FFF455C9A441FC3B5297" author="AUGE M. & RAGE J. - C." box="[714,926,887,914]" pageId="28" pageNumber="759" pagination="263 - 313" refId="ref15831" refString="AUGE M. & RAGE J. - C. 2000. - Les Squamates (Reptilia) du Miocene moyen de Sansan, in GINSBURG L. (ed.), La faune miocene de Sansan et son environnement. Memoires du Museum national d'Histoire naturelle 183: 263 - 313." type="journal article" year="2000">Augé & Rage, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
, known from the middle Miocene (MN 6) of Sansan (sometimes regarded as a member of the genus
|
||
<taxonomicName id="4C794D05FFD9FFF454B3A481FC5052D5" authority="Szyndlar 2005" authorityName="Laurenti" authorityYear="1768" box="[944,1013,951,976]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="759" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DEA94FFD9FFF454B3A481FC5052D5" box="[944,1013,951,976]" italics="true" pageId="28" pageNumber="759">Natrix</emphasis>
|
||
</taxonomicName>
|
||
|
||
<bibRefCitation id="EFE84B77FFD9FFF4530BA481FB3852D7" author="SZYNDLAR Z." box="[1032,1181,951,978]" pageId="28" pageNumber="759" pagination="31 - 52" refId="ref18137" refString="SZYNDLAR Z. 2005. - Snake fauna from the late Miocene of Rudabanya. Palaeontographia Italica 90: 31 - 52." type="journal article" year="2005">Szyndlar 2005</bibRefCitation>
|
||
), the prezygapophyseal processes and the hypapophysis are extremely short, while the neural lamina usually lacks an epizygapophyseal spine (
|
||
<bibRefCitation id="EFE84B77FFD9FFF45353A320FCA35555" author="AUGE M. & RAGE J. - C." pageId="28" pageNumber="759" pagination="263 - 313" refId="ref15831" refString="AUGE M. & RAGE J. - C. 2000. - Les Squamates (Reptilia) du Miocene moyen de Sansan, in GINSBURG L. (ed.), La faune miocene de Sansan et son environnement. Memoires du Museum national d'Histoire naturelle 183: 263 - 313." type="journal article" year="2000">Augé & Rage 2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |