170 lines
19 KiB
XML
170 lines
19 KiB
XML
<document id="7E4BD3AD95C020AFCB72E35F0807BC77" ID-DOI="10.1206/0003-0090(2000)255<0001:TSAGOT>2.0.CO;2" ID-ISSN="0003-0090" ID-Zenodo-Dep="12774565" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721325710038" checkinUser="felipe" docAuthor="ROIG-JUÑENT, SERGIO" docDate="2000" docId="5B1D87D7FF8BFFA61E9543B9FC80FD29" docLanguage="en" docName="BulAmeMusNatHis.2000.B255.1-83.pdf" docOrigin="Bulletin of the American Museum of Natural History 2000 (1)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0090(2000)255%3C0001%3ATSAGOT%3E2.0.CO%3B2" docStyle="DocumentStyle:25EFC9A7B3A675566A5530656FFE3DF4.1:BulAmeMusNatHis.2000-2010.chapters" docStyleId="25EFC9A7B3A675566A5530656FFE3DF4" docStyleName="BulAmeMusNatHis.2000-2010.chapters" docStyleVersion="1" docTitle="Nothobroscina ROIG-JUÑENT, 2000, NEW SUBTRIBE" docType="treatment" docVersion="2" lastPageNumber="32" masterDocId="A724FFAFFF94FFB91E1B4106FFCBFF80" masterDocTitle="The Subtribes And Genera Of The Tribe Broscini (Coleoptera: Carabidae): Cladistic Analysis, Taxonomic Treatment, And Biogeographical Considerations" masterLastPageNumber="90" masterPageNumber="1" pageNumber="32" updateTime="1721326802793" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="BB5469AF17602CC07B17014B8EA3917D">
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<mods:title id="3AAFB14383884A55C63CD69C988ECB35">The Subtribes And Genera Of The Tribe Broscini (Coleoptera: Carabidae): Cladistic Analysis, Taxonomic Treatment, And Biogeographical Considerations</mods:title>
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<mods:namePart id="C9A2E0221623E9FB353157753F05C43C">ROIG-JUÑENT, SERGIO</mods:namePart>
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<mods:typeOfResource id="4289F12FB2D03AC39AE6F6941FAD472A">text</mods:typeOfResource>
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<mods:title id="2ED4E7888286A89DDD37AACEE161E84A">Bulletin of the American Museum of Natural History</mods:title>
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<mods:part id="311F2A3DD5312D79433DEB7D0A40D44C">
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<mods:date id="227AE58ACAE4B93B9EA80C061AFE0A9B">2000</mods:date>
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<mods:number id="42A79B27071D419BC184BCD7DDE1BB57">2000-12-31</mods:number>
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<mods:number id="E060C8074D5DCAA141DB45D36EA9CDBE">2000</mods:number>
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<mods:classification id="FB17B951D1AFE0E9ACD13CBFF18BD360">journal article</mods:classification>
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<mods:identifier id="70DD54B42F1DE0E5A6B73421C239B667" type="DOI">10.1206/0003-0090(2000)255<0001:TSAGOT>2.0.CO;2</mods:identifier>
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<mods:identifier id="D7FF746059D93D04B74F037E369B554E" type="ISSN">0003-0090</mods:identifier>
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<treatment id="5B1D87D7FF8BFFA61E9543B9FC80FD29" ID-DOI="http://doi.org/10.5281/zenodo.12774611" ID-Zenodo-Dep="12774611" LSID="urn:lsid:plazi:treatment:5B1D87D7FF8BFFA61E9543B9FC80FD29" httpUri="http://treatment.plazi.org/id/5B1D87D7FF8BFFA61E9543B9FC80FD29" lastPageNumber="32" pageId="31" pageNumber="32">
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<subSubSection id="9BAE654AFF8BFFA61E9543B9FDBAFD56" box="[142,625,703,726]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="D30B36C1FF8BFFA61E9543B9FDBAFD56" blockId="31.[119,648,703,1737]" box="[142,625,703,726]" pageId="31" pageNumber="32">
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<heading id="884381ADFF8BFFA61E9543B9FDBAFD56" box="[142,625,703,726]" centered="true" fontSize="10" level="2" pageId="31" pageNumber="32" reason="4">
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<taxonomicName id="14B44D42FF8BFFA61E9543B9FE41FD56" ID-CoL="8LTC4" authorityName="ROIG-JUÑENT" authorityYear="2000" box="[142,394,703,726]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subTribe" status="new subtribe" subTribe="Nothobroscina">NOTHOBROSCINA</taxonomicName>
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,
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<taxonomicNameLabel id="FAF357A8FF8BFFA61F8343B9FDBAFD56" box="[408,625,703,726]" pageId="31" pageNumber="32" rank="subTribe">NEW SUBTRIBE</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9BAE654AFF8BFFA61E8F43ECFC80FD29" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="D30B36C1FF8BFFA61E8F43ECFEE5FC9E" blockId="31.[119,648,703,1737]" pageId="31" pageNumber="32">
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<smallCapsWord id="D5EDA01DFF8BFFA61E8F43ECFF05FC80" baselines="763,763" box="[148,206,746,769]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="31" pageNumber="32">
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<typeStatus id="0C0F8863FF8BFFA61E8F43ECFF05FC80" box="[148,206,746,769]" pageId="31" pageNumber="32">TYPE</typeStatus>
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</smallCapsWord>
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<smallCapsWord id="D5EDA01DFF8BFFA61EFB43ECFEE6FC80" baselines="763,763" box="[224,301,746,769]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Genus" pageId="31" pageNumber="32">GENUS</smallCapsWord>
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:
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<taxonomicName id="14B44D42FF8BFFA61F5C43ECFEE0FC9E" authority="Roig-Junent and Ball, 1995" authorityName="Roig-Junent and Ball" authorityYear="1995" class="Insecta" family="Carabidae" genus="Nothobroscus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="E1C0EAD3FF8BFFA61F5C43ECFE23FC81" box="[327,488,746,769]" italics="true" pageId="31" pageNumber="32">Nothobroscus</emphasis>
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Roig-Juñent and Ball, 1995
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="D30B36C1FF8BFFA61E8E4223FF6FFBE1" blockId="31.[119,648,703,1737]" pageId="31" pageNumber="32">
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<smallCapsWord id="D5EDA01DFF8BFFA61E8E4223FF13FCBB" baselines="822,822" box="[149,216,805,828]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Taxa" pageId="31" pageNumber="32">TAXA</smallCapsWord>
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<smallCapsWord id="D5EDA01DFF8BFFA61EF34223FE94FCBB" baselines="822,822" box="[232,351,805,828]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Included" pageId="31" pageNumber="32">INCLUDED</smallCapsWord>
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: The present cladistic analysis agrees with Balls (1956) postulate that the austral genera that have sclerites X and Y form a monophyletic group, but this assemblage is more closely related to the austral
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<taxonomicName id="14B44D42FF8BFFA61ECD42B1FEF2FC4E" authorityName="Sloane" authorityYear="1920" box="[214,313,951,974]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="tribe" tribe="Broscini">Broscini</taxonomicName>
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(
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<taxonomicName id="14B44D42FF8BFFA61F4942B1FE0BFC4E" box="[338,448,951,974]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="52" phylum="Arthropoda" rank="subTribe" subTribe="Barypina">Barypina</taxonomicName>
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and
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<taxonomicName id="14B44D42FF8BFFA61C1742B1FD4AFC4E" authorityName="Jeannel" authorityYear="1941" box="[524,641,951,974]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subTribe" subTribe="Creobiina">Creobiina</taxonomicName>
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) than to the Holarctic genera (
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<taxonomicName id="14B44D42FF8BFFA61FFC42D3FD9BFC6C" box="[487,592,981,1004]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subTribe" subTribe="Broscina">Broscina</taxonomicName>
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and
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<taxonomicName id="14B44D42FF8BFFA61E6C42F4FF24FB89" authorityName="ROIG-JUÑENT" authorityYear="2000" box="[119,239,1010,1033]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subTribe" subTribe="Axonyina">Axonyina</taxonomicName>
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). For this reason, I erect the subtribe
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<taxonomicName id="14B44D42FF8BFFA61EA44509FEA6FBA6" authorityName="ROIG-JUÑENT" authorityYear="2000" box="[191,365,1039,1062]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subTribe" subTribe="Nothobroscina">Nothobroscina</taxonomicName>
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, including the austral genera with sclerites X and Y on the internal sac.
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</paragraph>
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<paragraph id="D30B36C1FF8BFFA61E8F4561FC1DFEE6" blockId="31.[119,648,703,1737]" lastBlockId="31.[695,1224,189,681]" pageId="31" pageNumber="32">
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<figureCitation id="4B8F2A44FF8BFFA61E8F4561FECCFBFE" box="[148,263,1127,1150]" captionStart="Fig" captionStartId="12.[141,179,1636,1657]" captionTargetBox="[218,1116,184,1610]" captionTargetId="figure-14@12.[217,1119,182,1614]" captionTargetPageId="12" captionText="Fig. 15. Strict consensus tree from the 15 cladograms obtained by successive weighting procedure." figureDoi="http://doi.org/10.5281/zenodo.12774571" httpUri="https://zenodo.org/record/12774571/files/figure.png" pageId="31" pageNumber="32">Figure 15</figureCitation>
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shows five synapomorphies defining
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<taxonomicName id="14B44D42FF8BFFA61EDE4583FEBEFB1C" authorityName="ROIG-JUÑENT" authorityYear="2000" box="[197,373,1157,1180]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subTribe" subTribe="Nothobroscina">Nothobroscina</taxonomicName>
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: tooth of mentum bifid at apex (
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<superScript id="24C19B89FF8BFFA61EC445A4FECDFB2F" attach="none" box="[223,262,1185,1209]" fontSize="6" pageId="31" pageNumber="32">103</superScript>
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) and mentum with two rounded foveae (
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<superScript id="24C19B89FF8BFFA61EC045B9FEC9FB4C" attach="none" box="[219,258,1214,1238]" fontSize="6" pageId="31" pageNumber="32">121</superScript>
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), sclerotized ring of sternum 9 complete and wide (
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<superScript id="24C19B89FF8BFFA61F6245DBFE6BFB6A" attach="none" box="[377,416,1244,1268]" fontSize="6" pageId="31" pageNumber="32">500</superScript>
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), sclerite X broad with central constriction and without lateroapical projections (
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<superScript id="24C19B89FF8BFFA61F734411FE44FAA4" attach="none" box="[360,399,1302,1326]" fontSize="6" pageId="31" pageNumber="32">592</superScript>
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), accessory gland of bursa copulatrix far off the spermatheca (
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<superScript id="24C19B89FF8BFFA61E9B4454FF6CFADF" attach="none" box="[128,167,1361,1385]" fontSize="6" pageId="31" pageNumber="32">652</superScript>
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), and helminthoid sclerite broad and short (
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<superScript id="24C19B89FF8BFFA61EDD4469FF26FAFC" attach="none" box="[198,237,1390,1414]" fontSize="6" pageId="31" pageNumber="32">732</superScript>
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). Some of these characters revert within
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<taxonomicName id="14B44D42FF8BFFA61EC8448BFE4AFA24" authorityName="ROIG-JUÑENT" authorityYear="2000" box="[211,385,1421,1444]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subTribe" subTribe="Nothobroscina">Nothobroscina</taxonomicName>
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, except the apex of mentum tooth, form of internal sac sclerite X of male, and the shape of sclerotized ring of sternum 9. Within
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<taxonomicName id="14B44D42FF8BFFA61F6B44E3FDD5FA7C" authorityName="ROIG-JUÑENT" authorityYear="2000" box="[368,542,1509,1532]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subTribe" subTribe="Nothobroscina">Nothobroscina</taxonomicName>
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,
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<taxonomicName id="14B44D42FF8BFFA61C3344E3FF50F999" authorityName="Sharp" authorityYear="1886" class="Insecta" family="Carabidae" genus="Diglymma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="E1C0EAD3FF8BFFA61C3344E3FF50F999" italics="true" pageId="31" pageNumber="32">Diglymma</emphasis>
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</taxonomicName>
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appears as the most plesiomorphic genus, as pointed out by Ball (1956). The sister group of
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<taxonomicName id="14B44D42FF8BFFA61EFE473BFE95F9D4" authorityName="Sharp" authorityYear="1886" box="[229,350,1597,1620]" class="Insecta" family="Carabidae" genus="Diglymma" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="E1C0EAD3FF8BFFA61EFE473BFE95F9D4" box="[229,350,1597,1620]" italics="true" pageId="31" pageNumber="32">Diglymma</emphasis>
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</taxonomicName>
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is formed by two monophyletic groups, one including
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<taxonomicName id="14B44D42FF8BFFA61FEC475CFD43F9F1" authorityName="Bates" authorityYear="1891" box="[503,648,1626,1649]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="E1C0EAD3FF8BFFA61FEC475CFD43F9F1" box="[503,648,1626,1649]" italics="true" pageId="31" pageNumber="32">Eurylychnus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="14B44D42FF8BFFA61EB2477EFE81F90F" box="[169,330,1656,1679]" class="Insecta" family="Carabidae" genus="Nothobroscus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="E1C0EAD3FF8BFFA61EB2477EFE81F90F" box="[169,330,1656,1679]" italics="true" pageId="31" pageNumber="32">Nothobroscus</emphasis>
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</taxonomicName>
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, characterized by the presence of narrow and curved frontal sulcus (
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<superScript id="24C19B89FF8BFFA61E9B47B4FF53F93F" attach="none" box="[128,152,1713,1737]" fontSize="6" pageId="31" pageNumber="32">72</superScript>
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), and the other including seven genera characterized by the absence of scrobal seta (
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<superScript id="24C19B89FF8BFFA61CDA41DCFD13FF67" attach="none" box="[705,728,217,241]" fontSize="6" pageId="31" pageNumber="32">81</superScript>
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), metafemur setae not aligned (
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<superScript id="24C19B89FF8BFFA61A4041DCFB49FF67" attach="none" box="[1115,1154,217,241]" fontSize="6" pageId="31" pageNumber="32">461</superScript>
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), and accessory gland of bursa copulatrix near spermatheca (
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<superScript id="24C19B89FF8BFFA61D414013FC4AFEA2" attach="none" box="[858,897,276,300]" fontSize="6" pageId="31" pageNumber="32">651</superScript>
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). This latter group includes the genus
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<taxonomicName id="14B44D42FF8BFFA61D274034FC77FEC9" authorityName="Blanchard" authorityYear="1853" box="[828,956,306,329]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="E1C0EAD3FF8BFFA61D274034FC77FEC9" box="[828,956,306,329]" italics="true" pageId="31" pageNumber="32">Mecodema</emphasis>
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</taxonomicName>
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(the most diversified genus, with 61 species).
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</paragraph>
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<paragraph id="D30B36C1FF8BFFA61CCF406BFC80FD29" blockId="31.[695,1224,189,681]" pageId="31" pageNumber="32">
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This subtribe comprises 10 genera (only two monospecific) with 92 species that occur in
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<collectingCountry id="ABA37651FF8BFFA61CC140A1FC8EFE3E" box="[730,837,423,446]" name="Australia" pageId="31" pageNumber="32">Australia</collectingCountry>
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(plus
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<collectingRegion id="1170F823FF8BFFA61D8D40A1FBC1FE3E" box="[918,1034,423,446]" country="Australia" name="Tasmania" pageId="31" pageNumber="32">Tasmania</collectingRegion>
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),
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<collectingCountry id="ABA37651FF8BFFA61A3A40A1FB0FFE3E" box="[1057,1220,423,446]" name="New Zealand" pageId="31" pageNumber="32">New Zealand</collectingCountry>
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,
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<collectingCountry id="ABA37651FF8BFFA61CAC40C3FCB9FE5C" box="[695,882,453,476]" name="New Caledonia" pageId="31" pageNumber="32">New Caledonia</collectingCountry>
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, and southern South America. The nothobroscine species live in different habitats. The species of
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<taxonomicName id="14B44D42FF8BFFA61DE54306FBB5FD97" authorityName="Blanchard" authorityYear="1853" box="[1022,1150,512,535]" class="Insecta" family="Carabidae" genus="Mecodema" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="E1C0EAD3FF8BFFA61DE54306FBB5FD97" box="[1022,1150,512,535]" italics="true" pageId="31" pageNumber="32">Mecodema</emphasis>
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</taxonomicName>
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mostly inhabit alpine regions in
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<collectingCountry id="ABA37651FF8BFFA61A06431BFB0FFDB4" box="[1053,1220,541,564]" name="New Zealand" pageId="31" pageNumber="32">New Zealand</collectingCountry>
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,
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<taxonomicName id="14B44D42FF8BFFA61CAC433CFCC5FDD1" authorityName="Castelnau" authorityYear="1867" box="[695,782,570,593]" class="Insecta" family="Carabidae" genus="Brullea" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="E1C0EAD3FF8BFFA61CAC433CFCC5FDD1" box="[695,782,570,593]" italics="true" pageId="31" pageNumber="32">Brullea</emphasis>
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</taxonomicName>
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has a psammophilus habitat, and
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<taxonomicName id="14B44D42FF8BFFA61A87433CFCFCFDEF" class="Insecta" family="Carabidae" genus="Nothobroscus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E1C0EAD3FF8BFFA61A87433CFCFCFDEF" italics="true" pageId="31" pageNumber="32">Nothobroscus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="14B44D42FF8BFFA61D60435EFBC7FDEF" authorityName="Bates" authorityYear="1891" box="[891,1036,600,623]" class="Insecta" family="Carabidae" genus="Eurylychnus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="E1C0EAD3FF8BFFA61D60435EFBC7FDEF" box="[891,1036,600,623]" italics="true" pageId="31" pageNumber="32">Eurylychnus</emphasis>
|
|
</taxonomicName>
|
|
inhabit
|
|
<taxonomicName id="14B44D42FF8BFFA61A6C435EFD3DFD0C" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="E1C0EAD3FF8BFFA61A6C435EFD3DFD0C" italics="true" pageId="31" pageNumber="32">Nothofagus</emphasis>
|
|
</taxonomicName>
|
|
forest in South America and
|
|
<collectingCountry id="ABA37651FF8BFFA61A4D4373FB0FFD0C" box="[1110,1220,629,652]" name="Australia" pageId="31" pageNumber="32">Australia</collectingCountry>
|
|
, respectively.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |