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<document id="EC73066BF52263A23500434CA944950D" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF60FF12FE30BD30A29DFCE2" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf.imf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.2:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="2" docTitle="Ludesia Noyes 2023" docType="key" docVersion="1" lastPageNumber="180" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="5239" masterPageNumber="5228" pageNumber="180" updateTime="1699893637884" updateUser="felipe" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="1EEE3A56B38A2D53F86B1D78CC83EF4A">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="DB05F7EC9EEACE5638ECFBC16D40D9B2">Noyes, John Stuart</mods:namePart>
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<mods:title id="9041202B448641A3F469FBBBBC87E3FC">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
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<mods:date id="57840E732FB9C963EEF21FC9FCFCE917">2023</mods:date>
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<paragraph id="8BAC36B1FF60FF12FE30BD30A578FEEE" blockId="181.[466,872,275,326]" box="[466,872,275,300]" pageId="181" pageNumber="180">
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<emphasis id="B967EAA3FF60FF12FE30BD30A578FEEE" bold="true" box="[466,872,275,300]" pageId="181" pageNumber="180">
Key to
<collectingRegion id="49D7F853FF60FF12FDFDBD37A44DFEEE" box="[543,605,275,299]" country="Tanzania" name="Pwani" pageId="181" pageNumber="180">Costa</collectingRegion>
Rican species of
<taxonomicName id="4C134D32FF60FF12FCF7BD37A578FEEE" authorityName="Noyes" authorityYear="2023" box="[789,872,275,299]" class="Liliopsida" family="Poaceae" genus="Ludesia" kingdom="Plantae" order="Poales" pageId="181" pageNumber="180" phylum="Tracheophyta" rank="genus">
<emphasis id="B967EAA3FF60FF12FCF7BD37A578FEEE" bold="true" box="[789,872,275,299]" italics="true" pageId="181" pageNumber="180">Ludesia</emphasis>
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<paragraph id="8BAC36B1FF60FF12FDDABD0AA51AFE83" blockId="181.[466,872,275,326]" box="[568,778,302,326]" pageId="181" pageNumber="180">(Females and males)</paragraph>
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<keyStep id="30E72E14FF60FF12FF57BD40A29CFE74" pageId="181" pageNumber="180">
<paragraph id="8BAC36B1FF60FF12FF57BD40A29CFE52" blockId="181.[181,1164,356,434]" pageId="181" pageNumber="180">
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1. Wings reduced (
<figureCitation id="13282A34FF60FF12FE57BD40A455FEB9" box="[437,581,356,380]" captionStart="Figs 453-471" captionStartId="769.[181,226,1526,1550]" captionTargetBox="[181,1162,224,1508]" captionTargetId="figure-10@769.[181,1163,223,1508]" captionTargetPageId="769" captionText="Figs 453-471. Oodia japetus, HT (453) ovipositor. Ludesia hemera (454) ovipositor; (455) mesosoma, dorsal view; (456) mesosoma, dorsal view, G; (457) fore wing; (458) antenna; (459) antenna, G; (460) hypopygium; (461) genitalia, G; (462) fore wing, G. Ludesia amalthea (463) mesosoma, dorsal view; (464) fore wing; (465) antenna; (466) antenna,G; (467) hypopygium; (468) head, facial view, G; (469) ovipositor; (470) mesosoma, dorsal view, G. Angebra praxadike (471) head, facial view." pageId="181" pageNumber="180">Figs 457, 464</figureCitation>
), fore wing not reaching half way along gaster (females). ..............................................................................................................................2
</keyLead>
</paragraph>
<paragraph id="8BAC36B1FF60FF12FF57BDBEA29CFE74" blockId="181.[181,1164,356,434]" box="[181,1164,409,434]" pageId="181" pageNumber="180">
<keyLead id="30E29584FF60FF12FF57BDBEA29CFE74" box="[181,1164,409,434]" pageId="181" pageNumber="180">
- Wings fully developed (
<figureCitation id="13282A34FF60FF12FDE6BDBDA44DFE74" box="[516,605,409,433]" captionStart="Figs 453-471" captionStartId="769.[181,226,1526,1550]" captionTargetBox="[181,1162,224,1508]" captionTargetId="figure-10@769.[181,1163,223,1508]" captionTargetPageId="769" captionText="Figs 453-471. Oodia japetus, HT (453) ovipositor. Ludesia hemera (454) ovipositor; (455) mesosoma, dorsal view; (456) mesosoma, dorsal view, G; (457) fore wing; (458) antenna; (459) antenna, G; (460) hypopygium; (461) genitalia, G; (462) fore wing, G. Ludesia amalthea (463) mesosoma, dorsal view; (464) fore wing; (465) antenna; (466) antenna,G; (467) hypopygium; (468) head, facial view, G; (469) ovipositor; (470) mesosoma, dorsal view, G. Angebra praxadike (471) head, facial view." pageId="181" pageNumber="180">Fig. 462</figureCitation>
), fore wing exceeding apex of gaster (males) .............3
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<keyStep id="30E72E14FF60FF12FF57BDEBA29DFD42" pageId="181" pageNumber="180">
<paragraph id="8BAC36B1FF60FF12FF57BDEBA29CFDD9" blockId="181.[181,1165,462,647]" pageId="181" pageNumber="180">
<keyLead id="30E29584FF60FF12FF57BDEBA29CFDD9" pageId="181" pageNumber="180">
2 (1) Head and thorax largely black or very dark brown with a weak metallic sheen; frontovertex and posterior part of mesoscutum (
<figureCitation id="13282A34FF60FF12FD57BDCDA51CFDC4" box="[693,780,489,513]" captionStart="Figs 453-471" captionStartId="769.[181,226,1526,1550]" captionTargetBox="[181,1162,224,1508]" captionTargetId="figure-10@769.[181,1163,223,1508]" captionTargetPageId="769" captionText="Figs 453-471. Oodia japetus, HT (453) ovipositor. Ludesia hemera (454) ovipositor; (455) mesosoma, dorsal view; (456) mesosoma, dorsal view, G; (457) fore wing; (458) antenna; (459) antenna, G; (460) hypopygium; (461) genitalia, G; (462) fore wing, G. Ludesia amalthea (463) mesosoma, dorsal view; (464) fore wing; (465) antenna; (466) antenna,G; (467) hypopygium; (468) head, facial view, G; (469) ovipositor; (470) mesosoma, dorsal view, G. Angebra praxadike (471) head, facial view." pageId="181" pageNumber="180">Fig. 455</figureCitation>
) completely smooth and shiny .......... ..........................................................................................
<taxonomicName id="4C134D32FF60FF12FC7BBE20A5F5FDD9" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[921,997,516,540]" class="Liliopsida" family="Poaceae" genus="Ludesia" kingdom="Plantae" order="Poales" pageId="181" pageNumber="180" phylum="Tracheophyta" rank="species" species="hemera" status="sp. nov.">
<emphasis id="B967EAA3FF60FF12FC7BBE20A5F5FDD9" box="[921,997,516,540]" italics="true" pageId="181" pageNumber="180">hemera</emphasis>
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<emphasis id="B967EAA3FF60FF12FC09BE20A226FDD9" bold="true" box="[1003,1078,516,540]" pageId="181" pageNumber="180">
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</emphasis>
(p. 180)
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<paragraph id="8BAC36B1FF60FF12FF57BE3BA29DFD42" blockId="181.[181,1165,462,647]" pageId="181" pageNumber="180">
<keyLead id="30E29584FF60FF12FF57BE3BA29DFD42" pageId="181" pageNumber="180">
- Head and thorax largely orange or orange-brown; frontovertex with distinct punctate-reticulate sculpture and virtually matt; mesoscutum (
<figureCitation id="13282A34FF60FF12FC77BE1DA5E2FD94" box="[917,1010,569,593]" captionStart="Figs 453-471" captionStartId="769.[181,226,1526,1550]" captionTargetBox="[181,1162,224,1508]" captionTargetId="figure-10@769.[181,1163,223,1508]" captionTargetPageId="769" captionText="Figs 453-471. Oodia japetus, HT (453) ovipositor. Ludesia hemera (454) ovipositor; (455) mesosoma, dorsal view; (456) mesosoma, dorsal view, G; (457) fore wing; (458) antenna; (459) antenna, G; (460) hypopygium; (461) genitalia, G; (462) fore wing, G. Ludesia amalthea (463) mesosoma, dorsal view; (464) fore wing; (465) antenna; (466) antenna,G; (467) hypopygium; (468) head, facial view, G; (469) ovipositor; (470) mesosoma, dorsal view, G. Angebra praxadike (471) head, facial view." pageId="181" pageNumber="180">Fig. 463</figureCitation>
) with distinct polygonally reticulate sculpture in posterior half or so .......................................... .....................................................................................
<taxonomicName id="4C134D32FF60FF12FC62BE4AA5CEFD43" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[896,990,622,646]" class="Liliopsida" family="Poaceae" genus="Ludesia" kingdom="Plantae" order="Poales" pageId="181" pageNumber="180" phylum="Tracheophyta" rank="species" species="amalthea" status="sp. nov.">
<emphasis id="B967EAA3FF60FF12FC62BE4AA5CEFD43" box="[896,990,622,646]" italics="true" pageId="181" pageNumber="180">amalthea</emphasis>
</taxonomicName>
<emphasis id="B967EAA3FF60FF12FC07BE4BA225FD42" bold="true" box="[997,1077,623,647]" pageId="181" pageNumber="180">
<taxonomicNameLabel id="A25457D8FF60FF12FC07BE4BA225FD42" box="[997,1077,623,647]" pageId="181" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(p. 183)
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<keyStep id="30E72E14FF60FF12FF57BE80A29DFCE2" pageId="181" pageNumber="180">
<paragraph id="8BAC36B1FF60FF12FF57BE80A29CFD34" blockId="181.[181,1165,676,807]" pageId="181" pageNumber="180">
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3 (1) Frontovertex with relatively shallow imbricate-reticulate to polygonally reticulate sculpture that is about as deep as that on mesoscutum (
<figureCitation id="13282A34FF60FF12FC9DBE9AA5C5FD13" box="[895,981,702,726]" captionStart="Figs 453-471" captionStartId="769.[181,226,1526,1550]" captionTargetBox="[181,1162,224,1508]" captionTargetId="figure-10@769.[181,1163,223,1508]" captionTargetPageId="769" captionText="Figs 453-471. Oodia japetus, HT (453) ovipositor. Ludesia hemera (454) ovipositor; (455) mesosoma, dorsal view; (456) mesosoma, dorsal view, G; (457) fore wing; (458) antenna; (459) antenna, G; (460) hypopygium; (461) genitalia, G; (462) fore wing, G. Ludesia amalthea (463) mesosoma, dorsal view; (464) fore wing; (465) antenna; (466) antenna,G; (467) hypopygium; (468) head, facial view, G; (469) ovipositor; (470) mesosoma, dorsal view, G. Angebra praxadike (471) head, facial view." pageId="181" pageNumber="180">Fig. 456</figureCitation>
) .......................... .........................................................................................
<taxonomicName id="4C134D32FF60FF12FC7BBEFDA5F5FD34" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[921,997,729,753]" class="Liliopsida" family="Poaceae" genus="Ludesia" kingdom="Plantae" order="Poales" pageId="181" pageNumber="180" phylum="Tracheophyta" rank="species" species="hemera" status="sp. nov.">
<emphasis id="B967EAA3FF60FF12FC7BBEFDA5F5FD34" box="[921,997,729,753]" italics="true" pageId="181" pageNumber="180">hemera</emphasis>
</taxonomicName>
<emphasis id="B967EAA3FF60FF12FC09BEFDA226FD34" bold="true" box="[1003,1078,729,753]" pageId="181" pageNumber="180">
<taxonomicNameLabel id="A25457D8FF60FF12FC09BEFDA226FD34" box="[1003,1078,729,753]" pageId="181" pageNumber="180" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
(p. 180)
</keyLead>
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<paragraph id="8BAC36B1FF60FF12FF57BED0A29DFCE2" blockId="181.[181,1165,676,807]" pageId="181" pageNumber="180">
<keyLead id="30E29584FF60FF12FF57BED0A29DFCE2" pageId="181" pageNumber="180">
- Frontovertex (
<figureCitation id="13282A34FF60FF12FE47BED0A7EFFCC9" box="[421,511,756,780]" captionStart="Figs 453-471" captionStartId="769.[181,226,1526,1550]" captionTargetBox="[181,1162,224,1508]" captionTargetId="figure-10@769.[181,1163,223,1508]" captionTargetPageId="769" captionText="Figs 453-471. Oodia japetus, HT (453) ovipositor. Ludesia hemera (454) ovipositor; (455) mesosoma, dorsal view; (456) mesosoma, dorsal view, G; (457) fore wing; (458) antenna; (459) antenna, G; (460) hypopygium; (461) genitalia, G; (462) fore wing, G. Ludesia amalthea (463) mesosoma, dorsal view; (464) fore wing; (465) antenna; (466) antenna,G; (467) hypopygium; (468) head, facial view, G; (469) ovipositor; (470) mesosoma, dorsal view, G. Angebra praxadike (471) head, facial view." pageId="181" pageNumber="180">Fig. 468</figureCitation>
) with punctate-reticulate sculpture that is clearly deeper than sculpture on mesoscutum (
<figureCitation id="13282A34FF60FF12FD86BF2AA4ABFCE2" box="[612,699,782,807]" captionStart="Figs 453-471" captionStartId="769.[181,226,1526,1550]" captionTargetBox="[181,1162,224,1508]" captionTargetId="figure-10@769.[181,1163,223,1508]" captionTargetPageId="769" captionText="Figs 453-471. Oodia japetus, HT (453) ovipositor. Ludesia hemera (454) ovipositor; (455) mesosoma, dorsal view; (456) mesosoma, dorsal view, G; (457) fore wing; (458) antenna; (459) antenna, G; (460) hypopygium; (461) genitalia, G; (462) fore wing, G. Ludesia amalthea (463) mesosoma, dorsal view; (464) fore wing; (465) antenna; (466) antenna,G; (467) hypopygium; (468) head, facial view, G; (469) ovipositor; (470) mesosoma, dorsal view, G. Angebra praxadike (471) head, facial view." pageId="181" pageNumber="180">Fig. 470</figureCitation>
).............................
<taxonomicName id="4C134D32FF60FF12FC64BF2AA5F4FCE3" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[902,996,782,806]" class="Liliopsida" family="Poaceae" genus="Ludesia" kingdom="Plantae" order="Poales" pageId="181" pageNumber="180" phylum="Tracheophyta" rank="species" species="amalthea" status="sp. nov.">
<emphasis id="B967EAA3FF60FF12FC64BF2AA5F4FCE3" box="[902,996,782,806]" italics="true" pageId="181" pageNumber="180">amalthea</emphasis>
</taxonomicName>
<emphasis id="B967EAA3FF60FF12FC09BF2BA226FCE2" bold="true" box="[1003,1078,783,807]" pageId="181" pageNumber="180">
<taxonomicNameLabel id="A25457D8FF60FF12FC09BF2BA226FCE2" box="[1003,1078,783,807]" pageId="181" pageNumber="180" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
(p. 183)
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