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<document id="053692C67E193404275CEE31EF984EF1" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF23FF5FFDF2BE3AA232FDAF" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Cheiloneurus miklo Noyes 2023, sp.nov." docType="treatment" docUuid="09D55524-C84A-42A2-8D7F-E898829B9F1E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="247" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="245" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="D4DB6B153E88117578223D0B51DEC739">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="D60D891E96023475399217A8417131A0">Noyes, John Stuart</mods:namePart>
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<subSubSection id="C309653AFF23FF51FDF2BE3AA521FDF2" box="[528,817,542,567]" pageId="246" pageNumber="245" type="nomenclature">
<paragraph id="8BAC36B1FF23FF51FDF2BE3AA521FDF2" blockId="246.[528,817,542,593]" box="[528,817,542,567]" pageId="246" pageNumber="245">
<heading id="D0E481DDFF23FF51FDF2BE3AA521FDF2" bold="true" box="[528,817,542,567]" centered="true" fontSize="10" level="2" pageId="246" pageNumber="245" reason="2">
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<taxonomicName id="4C134D32FF23FF51FDF2BE3AA4F0FDF3" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[528,736,542,566]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="246" pageNumber="245" phylum="Arthropoda" rank="species" species="miklo" status="sp.nov.">
<emphasis id="B967EAA3FF23FF51FDF2BE3AA4F0FDF3" bold="true" box="[528,736,542,566]" italics="true" pageId="246" pageNumber="245">Cheiloneurus miklo</emphasis>
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<subSubSection id="C309653AFF23FF51FDFFBE1DA534FD94" box="[541,804,569,593]" pageId="246" pageNumber="245" type="description">
<paragraph id="8BAC36B1FF23FF51FDFFBE1DA534FD94" blockId="246.[528,817,542,593]" box="[541,804,569,593]" pageId="246" pageNumber="245">
(
<figureCitation id="13282A34FF23FF51FDC6BE1DA4BEFD94" box="[548,686,569,593]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="246" pageNumber="245">Figs 608-611</figureCitation>
; Hab.E 93)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF23FF50FF57BE4BA771F9D4" lastPageId="247" lastPageNumber="246" pageId="246" pageNumber="245" type="diagnosis">
<paragraph id="8BAC36B1FF23FF51FF57BE4BA237FAC2" blockId="246.[181,1165,622,1287]" pageId="246" pageNumber="245">
DIAGNOSIS.
<emphasis id="B967EAA3FF23FF51FEA9BE4AA787FD43" box="[331,407,622,646]" italics="true" pageId="246" pageNumber="245">Female</emphasis>
(length about
<quantity id="4CEB9B54FF23FF51FDC8BE4BA467FD42" box="[554,631,623,647]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="246" pageNumber="245" unit="mm" value="2.4">2.4mm</quantity>
): body generally dark brown to black with a weak to moderate metallic sheen; antenna (
<figureCitation id="13282A34FF23FF51FDFCBEADA466FD64" box="[542,630,649,673]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="246" pageNumber="245">Fig. 611</figureCitation>
) with scape pale orange, dorsally margined brown; pedicel mostly pale orange, dorsally dark brown proximally; funicle pale orange; clava dark brown; mesoscutum with uniform pale brown setae that appear silvery in some lights; coxae white; mid and hind tibiae each with a narrow, dark brown, subbasal band, legs otherwise yellowish with some dark brown areas; fore wing (
<figureCitation id="13282A34FF23FF51FDFBBED0A47FFCC9" box="[537,623,756,780]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="246" pageNumber="245">Fig. 608</figureCitation>
) largely hyaline but weakly infuscate below marginal and stigmal veins and with a weak, infuscate median streak in apical half; head (
<figureCitation id="13282A34FF23FF51FC0CBF2AA255FCE3" box="[1006,1093,782,806]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="246" pageNumber="245">Fig. 610</figureCitation>
) about 4.4X as wide as frontovertex, in facial view very slightly broader than long, subcircular, genae converging, moderately and evenly curved; frontovertex naked medially below anterior ocellus; area between eye and scrobe about as wide as diameter of anterior ocellus, with distinct sculpture, scrobal margin rounded; interantennal prominence dorsally rounded with only a few long setae; mandible with two long, sharp lower teeth and a short, sharp, upper tooth; antenna (
<figureCitation id="13282A34FF23FF51FC1FBFB0A242FC69" box="[1021,1106,916,940]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="246" pageNumber="245">Fig. 611</figureCitation>
) with scape about 4.6X as long as broad; funicle with F1 slightly longer than broad, remaining segments slightly transverse; head width slightly greater than length of flagellum; linear sensilla on F4-F6; clava 3-segmented, about as long as F2-F6 combined, sensory area large and forming an oblique truncation, extending about 0.4X along ventral margin; mesoscutum with polygonally reticulate sculpture; scutellum with a distinct, but short apical tuft of setae; wings fully developed; fore wing (
<figureCitation id="13282A34FF23FF51FF1BB810A742FB89" box="[249,338,1076,1100]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="246" pageNumber="245">Fig. 608</figureCitation>
) about 2.5-2.6X as long as broad; parastigma very weakly curved downwards; costal cell dorsally with about 5 to 11 setae at apex and a single, complete line of setae ventrally; area below proximal part of parastigma with a group of about 35 setae; apices of postmarginal and stigmal veins (
<figureCitation id="13282A34FF23FF51FE96B8A0A7DAFB59" box="[372,458,1156,1180]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="246" pageNumber="245">Fig. 609</figureCitation>
) not connected by a naked hyaline area; apical bristle of postmarginal vein about 0.4X as long as marginal vein; mid tibial spur about as long as basitarsus; propodeum with about 35 setae around each spiracle, side naked; syntergum about 1.3X as long as mid tibia; ovipositor about 2.7X gonostylus or 2.1X mid tibia; gonostylus about 2.3X as long as mid tibial spur; exserted part of ovipositor about 0.8X as long as mid tibial spur.
<emphasis id="B967EAA3FF23FF51FC62B8CBA5A4FAC2" box="[896,948,1263,1287]" italics="true" pageId="246" pageNumber="245">Male</emphasis>
: unknown.
</paragraph>
<paragraph id="8BAC36B1FF23FF51FF57B900A293FAF9" blockId="246.[181,1165,1316,1687]" box="[181,1155,1316,1340]" pageId="246" pageNumber="245">
Female (
<typeStatus id="54A88813FF23FF51FEECB900A77DFAF9" box="[270,365,1316,1340]" pageId="246" pageNumber="245">holotype</typeStatus>
): length, including ovipositor,
<quantity id="4CEB9B54FF23FF51FD40B900A4E9FAF9" box="[674,761,1316,1340]" metricMagnitude="-3" metricUnit="m" metricValue="2.38" pageId="246" pageNumber="245" unit="mm" value="2.38">2.38mm</quantity>
; excluding ovipositor,
<quantity id="4CEB9B54FF23FF51FC3DB900A223FAF9" box="[991,1075,1316,1340]" metricMagnitude="-3" metricUnit="m" metricValue="2.22" pageId="246" pageNumber="245" unit="mm" value="2.22">2.22mm</quantity>
(CPD).
</paragraph>
<paragraph id="8BAC36B1FF23FF51FF39B91BA29CF952" blockId="246.[181,1165,1316,1687]" pageId="246" pageNumber="245">
Head black with a weak blue and purple sheen on frontovertex, more strongly purple anteriorly; occiput with a blue sheen; scrobal area mostly purple with some dark blue reflections, interantennal prominence with a coppery purple band connecting toruli; mouth margin metallic blue-green; gena and temple strongly metallic blue-green with some weak purple and coppery reflections; frontovertex with fairly conspicuous dark brown setae; antenna (
<figureCitation id="13282A34FF23FF51FC5AB98DA21CFA04" box="[952,1036,1449,1473]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="246" pageNumber="245">Fig. 611</figureCitation>
)with radicle dark brown; scape mostly pale orange with a narrow, proximal brown ring and dorsal margin brown except at extreme apex; pedicel mostly pale orange, basal half or so dark brown dorsally; funicle pale orange; clava dark brown; thorax mostly dark brown to black; pronotum mainly dull, dark green with a slight brassy sheen, posterior margin coppery; mesoscutum with a blue to blue-green sheen with purple reflections, posterior margin coppery purple, setae uniform and translucent pale brown, but silvery in some lights; tegula dark brown; axilla dull, dark purple-brown with coppery reflections; scutellum noticeably less shiny than mesoscutum, but with a distinct dark blue sheen, medially strongly coppery, clothed with dark brown setae, extreme sides
</paragraph>
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<pageTitle id="CB8CEED6FF22FF50FEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="247" pageNumber="246">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FF22FF50FD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="247" pageNumber="246" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
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</paragraph>
<paragraph id="8BAC36B1FF22FF50FF57BCFAA450FD97" blockId="247.[181,1165,222,1500]" pageId="247" pageNumber="246">
and apex shining green mixed coppery, especially at apex; metanotum with slight purple sheen; mesopleuron in posterior half with a coppery purple and brassy sheen, anterior half mixed blue, green, coppery and brassy, these narrowly separated by a coppery, dark blue and violet sheen; fore coxa white; fore femur white, pale orange distally, fore tibia pale orange with an inconspicuous pale brown, subbasal spot; mid coxa white; mid femur white, pale orange towards apex, tibia pale orange with a dark brown, subbasal spot; hind coxa white; hind femur white, pale orange towards apex, tibia mostly pale orange with an incomplete, brown, subbasal ring; fore tarsus pale orange-brown, mid and hind tarsi pale orange, pretarsi slightly dusky; fore wing (
<figureCitation id="13282A34FF22FF50FBCFBDBDA295FE74" box="[1069,1157,409,433]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="247" pageNumber="246">Fig. 608</figureCitation>
) almost completely hyaline but weakly infuscate immediately below marginal and stigmal veins, and with an extremely weakly infuscate median streak distad of pale brown venation; propodeum dark purple-brown; sides metallic green with a few conspicuous silvery setae in spiracular area; gaster dark purple-brown, Gt1 metallic green proximally, dorsum slightly coppery, mixed purple and brassy, sides and outer plate of ovipositor mixed metallic green, blue, coppery and brassy; gonostylus slightly dusky, pale orange.
</paragraph>
<paragraph id="8BAC36B1FF22FF50FF39BE70A6C5FC24" blockId="247.[181,1165,222,1500]" pageId="247" pageNumber="246">
Head (
<figureCitation id="13282A34FF22FF50FEC0BE70A76BFDA9" box="[290,379,596,620]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="247" pageNumber="246">Fig. 610</figureCitation>
) about 4.4X as wide as frontovertex, not subtriangular in profile, occipital margin carinate, occiput without a median groove above foramen; ocelli forming an angle of about 75°; frontovertex slightly shiny, with conspicuous reticulate sculpture of mesh size generally very slightly smaller than diameter of eye facet; scrobal area and mouth margin with shallower sculpture; area immediately below eye almost completely smooth; gena and temple with more elongate sculpture of larger mesh; scrobes shallow, meeting dorsally, dorsal and lateral margins weakly rounded; antenna as in
<figureCitation id="13282A34FF22FF50FE0FBED0A453FCC9" box="[493,579,756,780]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="247" pageNumber="246">Fig. 611</figureCitation>
; scape hardly broadened and flattened, about 4.6X as long as broad; F1 a little longer than broad, F2-F6 at least slightly transverse; clava hardly broadened, almost as long as F2-F6 combined; only outer suture noticeably oblique, sensory area enlarged, extending about two-fifths along clava ventrally and giving it an obliquely truncate appearance; eye reaching occipital margin, clothed with fairly conspicuous setae, each slightly longer than diameter of facet; inner eye margins not converging anteriorly; malar sulcus absent; mandible with two, relatively long, acute, lower teeth, and a short, upper tooth. Relative measurements: HW 87,
<collectingRegion id="49D7F853FF22FF50FF3EBF8BA711FC03" box="[220,257,943,966]" country="Swaziland" name="Hhohho" pageId="247" pageNumber="246">HH</collectingRegion>
85, FV 20, OD 8, POL 10, OOL 1, OCL 5.5, AOL 11, EL 56, EW 54, MS 32, SL 46, SW 10.
</paragraph>
<paragraph id="8BAC36B1FF22FF50FF39BFC0A754FAE7" blockId="247.[181,1165,222,1500]" pageId="247" pageNumber="246">
Thorax with shallow, irregular, polygonally reticulate sculpture on pronotum, generally of mesh size subequal to diameter of facet; sculpture on mesoscutum generally of similar depth to that on pronotum, but more regular and of smaller mesh; sculpture on axilla similar to that on mesoscutum, but slightly deeper; scutellum with conspicuously deeper, polygonally reticulate sculpture; apex and sides completely smooth and shiny; visible part of mesoscutum about 1.6X as broad as long; scutellum about as long as broad, with a short subapical tuft, setae about one-fifth to one-quarter as long as scutellum; hind femur about 4X as long as broad; fore wing with venation and setation as in
<figureCitation id="13282A34FF22FF50FE8DB8BBA7EEFB72" box="[367,510,1183,1207]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="247" pageNumber="246">Figs 609, 611</figureCitation>
; costal cell with about 5 setae dorsally at apex; no naked streak present between apices of postmarginal and stigmal veins (
<figureCitation id="13282A34FF22FF50FCE7B89DA54DFB14" box="[773,861,1209,1233]" captionStart="Figs 608-621" captionStartId="780.[181,226,1526,1550]" captionTargetBox="[184,1161,229,1508]" captionTargetId="figure-9@780.[181,1163,223,1508]" captionTargetPageId="780" captionText="Figs 608-621. Cheiloneurus miklo (608) fore wing; (609) apex of fore wing venation; (610) head, facial view; (611) antenna (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus lamus (612) fore wing, variant; (613) fore wing, variant; (614) apex of fore wing venation; (615) ovipositor; (616) hypopygium; (617) head, facial view; (618) antenna, variant (a, outer aspect of clava; b, inner aspect of clava); (619) antenna, variant (a, outer aspect of clava; b, inner aspect of clava). Cheiloneurus varablus, HT (620) head, facial view; (621) mesosoma, dorsal view." pageId="247" pageNumber="246">Fig. 609</figureCitation>
); propodeum with moderately dense setae in spiracular area, side naked, medially about 0.1X as long as scutellum and with some shallow polygonally reticulate sculpture. Relative measurements: FWL 109, FWW 42.5; HWL 85, HWW 23.
</paragraph>
<paragraph id="8BAC36B1FF22FF50FF39B900A75FFAB4" blockId="247.[181,1165,222,1500]" pageId="247" pageNumber="246">Gaster with hypopygium reaching about half-way to apex; syntergum about 1.3X as long as mid tibia, with apex acute; ovipositor slightly exserted, the exserted part about 0.75X as long as mid tibial spur.</paragraph>
<paragraph id="8BAC36B1FF22FF50FF39B951A536FA62" blockId="247.[181,1165,222,1500]" pageId="247" pageNumber="246">
<typeStatus id="54A88813FF22FF50FF39B951A727FA48" box="[219,311,1397,1421]" pageId="247" pageNumber="246">Paratype</typeStatus>
. Funicle with linear sensilla only on F4-F6; gaster without “gland-like” structures on Gt1 or Gt5. Relative measurements: OL 107, GL 40 [MT 51].
</paragraph>
<paragraph id="8BAC36B1FF22FF50FF39B98DA755FA19" blockId="247.[181,1165,222,1500]" pageId="247" pageNumber="246">Variation. Very little in material examined, but the fore wing varies from about 2.5-2.6X as long as broad.</paragraph>
<paragraph id="8BAC36B1FF22FF50FF57B9DDA771F9D4" blockId="247.[181,353,1529,1553]" box="[181,353,1529,1553]" pageId="247" pageNumber="246">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF22FF50FF57BA0BA7F9F982" box="[181,489,1583,1607]" pageId="247" pageNumber="246" type="distribution">
<paragraph id="8BAC36B1FF22FF50FF57BA0BA7F9F982" blockId="247.[181,489,1583,1607]" box="[181,489,1583,1607]" pageId="247" pageNumber="246">
DISTRIBUTION.
<collectingCountry id="F3047621FF22FF50FE91BA0BA7F5F982" box="[371,485,1583,1607]" name="Costa Rica" pageId="247" pageNumber="246">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF22FF5FFF57BA40A29CFF5A" box="[181,1164,130,1660]" lastPageId="248" lastPageNumber="247" pageId="247" pageNumber="246" type="biology_ecology">
<paragraph id="8BAC36B1FF22FF50FF57BA40A76EF9B9" blockId="247.[181,382,1636,1660]" box="[181,382,1636,1660]" pageId="247" pageNumber="246">HOSTS. Unknown.</paragraph>
<paragraph id="8BAC36B1FF2DFF5FFBBCBCA6A29CFF5A" blockId="248.[1118,1164,130,159]" box="[1118,1164,130,159]" pageId="248" pageNumber="247">247</paragraph>
</subSubSection>
<subSubSection id="C309653AFF2DFF5FFF57BCFBA739FE81" pageId="248" pageNumber="247" type="materials_examined">
<paragraph id="8BAC36B1FF2DFF5FFF57BCFBA7BDFF31" blockId="248.[181,1165,223,324]" box="[181,429,223,244]" pageId="248" pageNumber="247">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF2DFF5FFF39BCDDA739FE81" blockId="248.[181,1165,223,324]" pageId="248" pageNumber="247">
Type material.
<materialsCitation id="3B7B3CECFF2DFF5FFE87BCDDA299FECB" box="[357,1161,248,271]" collectingDate="1995-01" collectorName="de Moravia &amp; P. Hanson" country="Costa Rica" elevation="1600" location="San Jose" municipality="Zurqui de Moravia" pageId="248" pageNumber="247" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A88813FF2DFF5FFE87BCDDA7A9FECA" box="[357,441,249,271]" pageId="248" pageNumber="247">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF2DFF5FFE3DBCDDA47AFECA" bold="true" box="[479,618,249,271]" pageId="248" pageNumber="247">
<collectingCountry id="F3047621FF2DFF5FFE3DBCDDA47AFECA" box="[479,618,249,271]" name="Costa Rica" pageId="248" pageNumber="247">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF2DFF5FFD95BCDDA4DCFECB" bold="true" box="[631,716,248,271]" pageId="248" pageNumber="247">
<location id="8ECC606AFF2DFF5FFD95BCDDA4DCFECB" LSID="urn:lsid:plazi:treatment:03BA87A7FF23FF5FFDF2BE3AA232FDAF:8ECC606AFF2DFF5FFD95BCDDA4DCFECB" box="[631,716,248,271]" country="Costa Rica" municipality="Zurqui de Moravia" name="San Jose" pageId="248" pageNumber="247">San José</location>
</emphasis>
,
<collectingMunicipality id="6BC8ACCBFF2DFF5FFD35BCDDA597FECA" box="[727,903,249,271]" pageId="248" pageNumber="247">
Zurqui
<collectorName id="26E65367FF2DFF5FFCFEBCDDA597FECA" box="[796,903,249,271]" pageId="248" pageNumber="247">de Moravia</collectorName>
</collectingMunicipality>
,
<quantity id="4CEB9B54FF2DFF5FFC73BCDDA5C2FECA" box="[913,978,249,271]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="248" pageNumber="247" unit="m" value="1600.0">
<elevation id="003ED182FF2DFF5FFC73BCDDA5C2FECA" box="[913,978,249,271]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="248" pageNumber="247" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<date id="FFAD1071FF2DFF5FFC3EBCDDA205FECA" box="[988,1045,249,271]" pageId="248" pageNumber="247" value="1995-01">
<collectingDate id="EFE9E999FF2DFF5FFC3EBCDDA205FECA" box="[988,1045,249,271]" pageId="248" pageNumber="247" value="1995-01">i.1995</collectingDate>
</date>
(
<collectorName id="26E65367FF2DFF5FFBC0BCDDA293FECB" box="[1058,1155,249,270]" pageId="248" pageNumber="247">P. Hanson</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF2DFF5FFF57BD30A579FEEF" box="[181,873,275,298]" collectingDate="1992-07" collectorName="de Moravia &amp; P. Hanson" country="Costa Rica" elevation="1600" location="San Jose" municipality="Zurqui de Moravia" pageId="248" pageNumber="247" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
<typeStatus id="54A88813FF2DFF5FFF57BD30A718FEEC" box="[181,264,276,297]" pageId="248" pageNumber="247">Paratype</typeStatus>
:
<emphasis id="B967EAA3FF2DFF5FFEF2BD37A78BFEEC" bold="true" box="[272,411,275,297]" pageId="248" pageNumber="247">
<collectingCountry id="F3047621FF2DFF5FFEF2BD37A78BFEEC" box="[272,411,275,297]" name="Costa Rica" pageId="248" pageNumber="247">COSTA RICA</collectingCountry>
</emphasis>
, 1E,
<emphasis id="B967EAA3FF2DFF5FFE2CBD37A432FEEC" bold="true" box="[462,546,275,297]" pageId="248" pageNumber="247">
<collectingRegion id="49D7F853FF2DFF5FFE2CBD37A432FEEC" box="[462,546,275,297]" pageId="248" pageNumber="247">San José</collectingRegion>
</emphasis>
, same data as holotype but
<date id="FFAD1071FF2DFF5FFCF9BD30A579FEEF" box="[795,873,276,298]" pageId="248" pageNumber="247" value="1992-07">
<collectingDate id="EFE9E999FF2DFF5FFCF9BD30A579FEEF" box="[795,873,276,298]" pageId="248" pageNumber="247" value="1992-07">vii.1992</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF2DFF5FFC90BD30A736FE81" collectionCode="MZUCR, NHMUK" country="Costa Rica" pageId="248" pageNumber="247" specimenCount="2" stateProvince="San Jose" typeStatus="holotype">
<typeStatus id="54A88813FF2DFF5FFC90BD30A5D6FEEF" box="[882,966,276,298]" pageId="248" pageNumber="247">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FF2DFF5FFC00BD30A225FEEC" box="[994,1077,276,297]" pageId="248" pageNumber="247">MZUCR</collectionCode>
,
<typeStatus id="54A88813FF2DFF5FFBDCBD30A29CFEEC" box="[1086,1164,276,297]" pageId="248" pageNumber="247">paratype</typeStatus>
in
<collectionCode id="ED02AE74FF2DFF5FFF2FBD0BA736FE81" box="[205,294,303,324]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="248" pageNumber="247" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF2DFF5FFF57BD46A232FDAF" pageId="248" pageNumber="247" type="discussion">
<paragraph id="8BAC36B1FF2DFF5FFF57BD46A298FD8A" blockId="248.[181,1165,354,618]" pageId="248" pageNumber="247">
COMMENTS. As the key suggests,
<taxonomicName id="4C134D32FF2DFF5FFDD6BD46A4EEFEBF" authorityName="Noyes" authorityYear="2023" box="[564,766,354,378]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="miklo">
<emphasis id="B967EAA3FF2DFF5FFDD6BD46A4EEFEBF" box="[564,766,354,378]" italics="true" pageId="248" pageNumber="247">Cheiloneurus miklo</emphasis>
</taxonomicName>
is quite similar to
<taxonomicName id="4C134D32FF2DFF5FFC28BD46A202FEBF" authorityName="Noyes" authorityYear="2023" box="[970,1042,354,378]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="fidenas">
<emphasis id="B967EAA3FF2DFF5FFC28BD46A202FEBF" box="[970,1042,354,378]" italics="true" pageId="248" pageNumber="247">fidenas</emphasis>
</taxonomicName>
,
<taxonomicName id="4C134D32FF2DFF5FFBC0BD46A24FFEBF" authorityName="Noyes" authorityYear="2023" box="[1058,1119,354,378]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="lamus">
<emphasis id="B967EAA3FF2DFF5FFBC0BD46A24FFEBF" box="[1058,1119,354,378]" italics="true" pageId="248" pageNumber="247">lamus</emphasis>
</taxonomicName>
and
<emphasis id="B967EAA3FF2DFF5FFF57BD59A6E4FE51" box="[181,244,381,404]" italics="true" pageId="248" pageNumber="247">
<taxonomicName id="4C134D32FF2DFF5FFF57BD59A6E0FE51" authorityName="Noyes" authorityYear="2023" box="[181,240,381,404]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="rosea">rosea</taxonomicName>
.
</emphasis>
In addition to the characters given in the key it can be separated from both
<taxonomicName id="4C134D32FF2DFF5FFBF5BD59A24FFE50" authorityName="Noyes" authorityYear="2023" box="[1047,1119,381,405]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="fidenas">
<emphasis id="B967EAA3FF2DFF5FFBF5BD59A24FFE50" box="[1047,1119,381,405]" italics="true" pageId="248" pageNumber="247">fidenas</emphasis>
</taxonomicName>
and
<taxonomicName id="4C134D32FF2DFF5FFF57BDB3A6E2FE6A" authorityName="Noyes" authorityYear="2023" box="[181,242,407,431]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="lamus">
<emphasis id="B967EAA3FF2DFF5FFF57BDB3A6E2FE6A" box="[181,242,407,431]" italics="true" pageId="248" pageNumber="247">lamus</emphasis>
</taxonomicName>
by the lack of linear sensilla on F3. In both
<taxonomicName id="4C134D32FF2DFF5FFD27BDB3A51DFE6A" authorityName="Noyes" authorityYear="2023" box="[709,781,407,431]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="fidenas">
<emphasis id="B967EAA3FF2DFF5FFD27BDB3A51DFE6A" box="[709,781,407,431]" italics="true" pageId="248" pageNumber="247">fidenas</emphasis>
</taxonomicName>
and
<taxonomicName id="4C134D32FF2DFF5FFCA3BDB3A56EFE6A" authorityName="Noyes" authorityYear="2023" box="[833,894,407,431]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="lamus">
<emphasis id="B967EAA3FF2DFF5FFCA3BDB3A56EFE6A" box="[833,894,407,431]" italics="true" pageId="248" pageNumber="247">lamus</emphasis>
</taxonomicName>
linear sensilla are present on F3. Also, in
<taxonomicName id="4C134D32FF2DFF5FFEBABD96A780FE0F" authorityName="Noyes" authorityYear="2023" box="[344,400,434,458]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="miklo">
<emphasis id="B967EAA3FF2DFF5FFEBABD96A780FE0F" box="[344,400,434,458]" italics="true" pageId="248" pageNumber="247">miklo</emphasis>
</taxonomicName>
the sensory area of the clava extends about 0.4X along ventral surface of the clava whereas in
<taxonomicName id="4C134D32FF2DFF5FFE69BDE9A7C3FE20" authorityName="Noyes" authorityYear="2023" box="[395,467,461,485]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="fidenas">
<emphasis id="B967EAA3FF2DFF5FFE69BDE9A7C3FE20" box="[395,467,461,485]" italics="true" pageId="248" pageNumber="247">fidenas</emphasis>
</taxonomicName>
it extends only about 0.3X along ventral surface. It can be separated from
<taxonomicName id="4C134D32FF2DFF5FFF0DBDCCA736FE3A" authorityName="Noyes" authorityYear="2023" box="[239,294,488,511]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B967EAA3FF2DFF5FFF0DBDCCA736FE3A" box="[239,294,488,511]" italics="true" pageId="248" pageNumber="247">rosea</emphasis>
</taxonomicName>
on the colour of the hind femur and relative proportions of the ovipositor. In
<taxonomicName id="4C134D32FF2DFF5FFBB6BDC3A29CFE3A" authorityName="Noyes" authorityYear="2023" box="[1108,1164,487,511]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="miklo">
<emphasis id="B967EAA3FF2DFF5FFBB6BDC3A29CFE3A" box="[1108,1164,487,511]" italics="true" pageId="248" pageNumber="247">miklo</emphasis>
</taxonomicName>
the hind femur is completely yellow, the ovipositor is about 2X as long as the mid tibia and less than 3X as long as the gonostylus whereas in
<taxonomicName id="4C134D32FF2DFF5FFD64BE39A4ADFDF1" authorityName="Noyes" authorityYear="2023" box="[646,701,541,564]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="rosea">
<emphasis id="B967EAA3FF2DFF5FFD64BE39A4ADFDF1" box="[646,701,541,564]" italics="true" pageId="248" pageNumber="247">rosea</emphasis>
</taxonomicName>
the hind femur is apically marked brown and the ovipositor is about 1.5X as long as the mid tibia and more than 3X as long as the gonostylus.
</paragraph>
<paragraph id="8BAC36B1FF2DFF5FFF39BE76A232FDAF" blockId="248.[181,1165,354,618]" box="[219,1058,594,618]" pageId="248" pageNumber="247">
See also comments under
<taxonomicName id="4C134D32FF2DFF5FFE06BE76A45CFDAF" authorityName="Noyes" authorityYear="2023" box="[484,588,594,618]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="herndonia">
<emphasis id="B967EAA3FF2DFF5FFE06BE76A45CFDAF" box="[484,588,594,618]" italics="true" pageId="248" pageNumber="247">herndonia</emphasis>
</taxonomicName>
(p. 240),
<taxonomicName id="4C134D32FF2DFF5FFD4DBE77A4FFFDAF" authorityName="Noyes" authorityYear="2023" box="[687,751,595,618]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="exeres">
<emphasis id="B967EAA3FF2DFF5FFD4DBE77A4FFFDAF" box="[687,751,595,618]" italics="true" pageId="248" pageNumber="247">exeres</emphasis>
</taxonomicName>
(p. 243) and
<taxonomicName id="4C134D32FF2DFF5FFC95BE76A5D4FDAF" authorityName="Noyes" authorityYear="2023" box="[887,964,594,618]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="248" pageNumber="247" phylum="Arthropoda" rank="species" species="myndus">
<emphasis id="B967EAA3FF2DFF5FFC95BE76A5D4FDAF" box="[887,964,594,618]" italics="true" pageId="248" pageNumber="247">myndus</emphasis>
</taxonomicName>
(p. 253).
</paragraph>
</subSubSection>
</treatment>
</document>