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<document id="8C5ECC8869F2C8BC6E1CF5A9BC7AE052" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FF0DFF7EFDF2BEADA7EEFA4E" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Cheiloneurus vitara Noyes 2023, sp.nov." docType="treatment" docUuid="D7EBB2A1-321C-40CD-9430-BC06C9F8DE0D" docUuidSource="ZooBank" docVersion="2" lastPageNumber="216" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="215" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="6CD1B1DBA7D7860F21096BD60179A22F">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="ABFF9DD839DED633AA3C617BCEDC7268">Noyes, John Stuart</mods:namePart>
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<mods:date id="63CDA26C5C14007642716E8E13E63D56">2023</mods:date>
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<paragraph id="8BAC36B1FF0DFF7FFDF2BEADA522FD64" blockId="216.[528,818,649,700]" box="[528,818,649,673]" pageId="216" pageNumber="215">
<heading id="D0E481DDFF0DFF7FFDF2BEADA522FD64" bold="true" box="[528,818,649,673]" centered="true" fontSize="10" level="2" pageId="216" pageNumber="215" reason="2">
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<taxonomicName id="4C134D32FF0DFF7FFDF2BEADA4F1FD64" authority="Noyes, 2023" authorityName="Noyes" authorityYear="2023" box="[528,737,649,673]" class="Insecta" family="Encyrtidae" genus="Cheiloneurus" kingdom="Animalia" order="Hymenoptera" pageId="216" pageNumber="215" phylum="Arthropoda" rank="species" species="vitara" status="sp.nov.">
<emphasis id="B967EAA3FF0DFF7FFDF2BEADA4F1FD64" bold="true" box="[528,737,649,673]" italics="true" pageId="216" pageNumber="215">Cheiloneurus vitara</emphasis>
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<subSubSection id="C309653AFF0DFF7FFDB7BE80A4FCFD79" box="[597,748,676,700]" pageId="216" pageNumber="215" type="description">
<paragraph id="8BAC36B1FF0DFF7FFDB7BE80A4FCFD79" blockId="216.[528,818,649,700]" box="[597,748,676,700]" pageId="216" pageNumber="215">
(
<figureCitation id="13282A34FF0DFF7FFDBEBE80A4F5FD79" box="[604,741,676,700]" captionStart="Figs 498-509" captionStartId="772.[181,226,1526,1550]" captionTargetBox="[181,1163,225,1507]" captionTargetId="figure-9@772.[181,1163,223,1508]" captionTargetPageId="772" captionText="Figs 498-509. Cheiloneurus magnolia (498) head, facial view (modified); (499) fore wing; (500) apex of fore wing venation; (501) mesosoma, dorsal view; (502) antenna; (503) ovipositor; (504) gastral tergites, showing “glands” on Gt1 and Gt5; (505) hypopygium (posterior margin partly folded inwards). Cheiloneurus vitara, HT (506) antenna; (507) mesosoma, dorsal view; (508) apex of fore wing venation; (509) fore wing." pageId="216" pageNumber="215">Figs 506-509</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF0DFF7EFF57BEFDA771FB72" lastPageId="217" lastPageNumber="216" pageId="216" pageNumber="215" type="diagnosis">
<paragraph id="8BAC36B1FF0DFF7FFF57BEFDA40CFBD3" blockId="216.[181,1164,729,1046]" pageId="216" pageNumber="215">
DIAGNOSIS.
<emphasis id="B967EAA3FF0DFF7FFEAEBEFDA788FD34" box="[332,408,729,753]" italics="true" pageId="216" pageNumber="215">Female</emphasis>
(overall length about
<quantity id="4CEB9B54FF0DFF7FFD99BEFDA4D7FD34" box="[635,711,729,753]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="216" pageNumber="215" unit="mm" value="0.8">0.8mm</quantity>
): body dark brown with a weak to moderate metallic sheen; posterior part of mesoscutum with only a few translucent, silvery setae that contrast with dark brown anterior setae; fore wing hyaline; fore coxa white; head about 3X as wide as frontovertex; antenna (
<figureCitation id="13282A34FF0DFF7FFE1DBF0DA44AFC84" box="[511,602,809,833]" captionStart="Figs 498-509" captionStartId="772.[181,226,1526,1550]" captionTargetBox="[181,1163,225,1507]" captionTargetId="figure-9@772.[181,1163,223,1508]" captionTargetPageId="772" captionText="Figs 498-509. Cheiloneurus magnolia (498) head, facial view (modified); (499) fore wing; (500) apex of fore wing venation; (501) mesosoma, dorsal view; (502) antenna; (503) ovipositor; (504) gastral tergites, showing “glands” on Gt1 and Gt5; (505) hypopygium (posterior margin partly folded inwards). Cheiloneurus vitara, HT (506) antenna; (507) mesosoma, dorsal view; (508) apex of fore wing venation; (509) fore wing." pageId="216" pageNumber="215">Fig. 506</figureCitation>
) with pedicel nearly as long as F1-F4 combined; all funicle segments transverse; linear sensilla on F6 only; clava about as long as pedicel and funicle combined, apex rounded; mandible with 3 teeth, upper tooth truncate; mesoscutum (
<figureCitation id="13282A34FF0DFF7FFBC9BF7AA295FCB3" box="[1067,1157,862,886]" captionStart="Figs 498-509" captionStartId="772.[181,226,1526,1550]" captionTargetBox="[181,1163,225,1507]" captionTargetId="figure-9@772.[181,1163,223,1508]" captionTargetPageId="772" captionText="Figs 498-509. Cheiloneurus magnolia (498) head, facial view (modified); (499) fore wing; (500) apex of fore wing venation; (501) mesosoma, dorsal view; (502) antenna; (503) ovipositor; (504) gastral tergites, showing “glands” on Gt1 and Gt5; (505) hypopygium (posterior margin partly folded inwards). Cheiloneurus vitara, HT (506) antenna; (507) mesosoma, dorsal view; (508) apex of fore wing venation; (509) fore wing." pageId="216" pageNumber="215">Fig. 507</figureCitation>
) with aciculate, longitudinally elongate, lineolate sculpture, some polygonally reticulate sculpture medially near posterior margin; scutellum with longitudinally elongate, lineolate sculpture; scutellum without a tuft of setae; fore wing (
<figureCitation id="13282A34FF0DFF7FFD90BF8AA4D7FC03" box="[626,711,942,966]" captionStart="Figs 498-509" captionStartId="772.[181,226,1526,1550]" captionTargetBox="[181,1163,225,1507]" captionTargetId="figure-9@772.[181,1163,223,1508]" captionTargetPageId="772" captionText="Figs 498-509. Cheiloneurus magnolia (498) head, facial view (modified); (499) fore wing; (500) apex of fore wing venation; (501) mesosoma, dorsal view; (502) antenna; (503) ovipositor; (504) gastral tergites, showing “glands” on Gt1 and Gt5; (505) hypopygium (posterior margin partly folded inwards). Cheiloneurus vitara, HT (506) antenna; (507) mesosoma, dorsal view; (508) apex of fore wing venation; (509) fore wing." pageId="216" pageNumber="215">Fig. 509</figureCitation>
) about 2.4X as long as broad; parastigma not downcurved; stigmal vein with 3 campaniform sensilla at apex; gaster with “glands” on Gt1 and Gt5, those on Gt1 separated by about 1.1X their own lengths, those on Gt5 touching; syntergum broader than long.
<emphasis id="B967EAA3FF0DFF7FFE97BFDAA7B9FBD3" box="[373,425,1022,1046]" italics="true" pageId="216" pageNumber="215">Male</emphasis>
: unknown.
</paragraph>
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<materialsCitation id="3B7B3CECFF0DFF7FFF57B810A26EFB89" box="[181,1150,1076,1100]" collectionCode="CPD" pageId="216" pageNumber="215" specimenCount="1" typeStatus="holotype">
Female
<emphasis id="B967EAA3FF0DFF7FFEE4B810A71EFB89" bold="true" box="[262,270,1076,1100]" pageId="216" pageNumber="215">(</emphasis>
<typeStatus id="54A88813FF0DFF7FFEEDB810A778FB89" box="[271,360,1076,1100]" pageId="216" pageNumber="215">holotype</typeStatus>
<emphasis id="B967EAA3FF0DFF7FFE85B810A77FFB89" bold="true" box="[359,367,1076,1100]" pageId="216" pageNumber="215">)</emphasis>
: length, including ovipositor,
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; excluding ovipositor,
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(
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.
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<paragraph id="8BAC36B1FF0DFF7FFF39B86AA29CF952" blockId="216.[181,1165,1076,1687]" pageId="216" pageNumber="215">Head dark brown to black with a weak metallic lustre; frontovertex coppery purple behind posterior ocelli, area around anterior ocellus dull, metallic dark blue mixed coppery purple, area above scrobes purple; temple and gena with a dull dark blue and coppery purple sheen, area immediately below lowest part of eye relatively shiny green; scrobal area coppery purple mixed metallic dark blue; a line of inconspicuous, brown setae along inner eye margin continuing below eye to gena and along position of malar sulcus; radicle brown; scape with proximal 0.6X dark brown, apex pale yellow; pedicel brown with venter and apex narrowly dusky pale yellow; F1-F5 dusky pale orange, F6 mostly very pale orange, mixed slightly dusky externally; clava brown; thorax dark brown to black; pronotum with a weak, coppery purple and brassy sheen; mesoscutum virtually matt, but with a very weak coppery purple and brassy sheen, mostly clothed with brown setae, but with 3 or 4 silvery setae on each side in posterior one-third; tegula dark brown; axilla similarly coloured to mesoscutum; scutellum matt black; mesopleuron with a weak, metallic blue, purple and brassy sheen; fore coxa white; fore femur white with a fairly broad, incomplete, brown, subapical band, tibia pale yellow with a broad, subbasal brown band, tarsus dusky pale yellow, apical tarsomere pale brown; mid coxa dark brown; mid femur white with a narrow, subapical brown band in distal half, tibia white with a narrow, subbasal brown band in basal half, spur and tarsus white; hind coxa dark brown; hind femur dark brown, narrowly white basally, tibia mostly dark brown with apices white, tarsus white with apical tarsomere brown; wings hyaline, venation pale yellow; propodeum dark brown with a weak brassy and purple sheen, side with a dull, metallic green sheen mixed coppery and brassy, outside spiracle with 2 or 3 inconspicuous brown setae; gaster dark purple-brown, dorsally with a coppery purple sheen, Gt1 mixed with a slight green sheen, side and venter with coppery, blue, brassy and green reflections; gonostylus</paragraph>
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<pageTitle id="CB8CEED6FF0CFF7EFEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="217" pageNumber="216">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FF0CFF7EFD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="217" pageNumber="216" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
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</paragraph>
<paragraph id="8BAC36B1FF0CFF7EFF57BCFAA7B8FF33" blockId="217.[181,1165,222,1153]" box="[181,424,222,246]" pageId="217" pageNumber="216">completely dark brown.</paragraph>
<paragraph id="8BAC36B1FF0CFF7EFF39BCDDA23FFD12" blockId="217.[181,1165,222,1153]" pageId="217" pageNumber="216">
Head about 3.0X as wide as frontovertex, in profile about 1.3X as high as deep, tangent to frontovertex above scrobes forming an angle of about 75° with face below top of scrobes; a very small, oval depression adjacent to eye very near occipital margin; ocelli forming an angle of about 75°; frontovertex narrowest a little behind posterior ocelli, relatively dull, with relatively deep, polygonally reticulate to imbricate-reticulate sculpture, in ocellar area of mesh size a little larger than eye facet, subequal elsewhere; temple and gena with sculpture of similar depth to anterior part of frontovertex, but more longitudinally elongate imbricate-reticulate to polygonally reticulate; area immediately below lowest part of eye completely smooth; scrobes with relatively shallow, irregular, elongate sculpture; interantennal prominence and mouth margin with shallow, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, narrowly ∩-shaped; interantennal prominence dorsally indistinctly acute; antenna as in
<figureCitation id="13282A34FF0CFF7EFC6FBE20A5FBFDD9" box="[909,1003,516,540]" captionStart="Figs 498-509" captionStartId="772.[181,226,1526,1550]" captionTargetBox="[181,1163,225,1507]" captionTargetId="figure-9@772.[181,1163,223,1508]" captionTargetPageId="772" captionText="Figs 498-509. Cheiloneurus magnolia (498) head, facial view (modified); (499) fore wing; (500) apex of fore wing venation; (501) mesosoma, dorsal view; (502) antenna; (503) ovipositor; (504) gastral tergites, showing “glands” on Gt1 and Gt5; (505) hypopygium (posterior margin partly folded inwards). Cheiloneurus vitara, HT (506) antenna; (507) mesosoma, dorsal view; (508) apex of fore wing venation; (509) fore wing." pageId="217" pageNumber="216">Fig. 506</figureCitation>
; scape slightly broadened and flattened, about 3.3X as long as broad; all funicle segments transverse, increasing in width distad, F1-F5 anelliform, F6 subquadrate, linear sensilla present only on F6; clava about as long as pedicel and funicle combined, sutures parallel, sensory area small, at apex only giving it a pointed or slightly rounded appearance; eye not reaching occipital margin, separated by at least about 2X diameter of facet, naked; malar sulcus partially present, weakly indicated medially; mandible tridentate, with 2 teeth and a truncate upper tooth. Relative measurements: HW 40,
<collectingRegion id="49D7F853FF0CFF7EFB8ABE80A29DFD7E" box="[1128,1165,676,699]" country="Swaziland" name="Hhohho" pageId="217" pageNumber="216">HH</collectingRegion>
30, FV 13.5, POL 5.5, OOL 1.5, OCL 5, AOL 5, EL 18, EW 19, MS 15, SL 16, SW 4.8.
</paragraph>
<paragraph id="8BAC36B1FF0CFF7EFF39BEFDA228FC24" blockId="217.[181,1165,222,1153]" pageId="217" pageNumber="216">
Thorax (
<figureCitation id="13282A34FF0CFF7EFED6BEFDA79CFD34" box="[308,396,729,753]" captionStart="Figs 498-509" captionStartId="772.[181,226,1526,1550]" captionTargetBox="[181,1163,225,1507]" captionTargetId="figure-9@772.[181,1163,223,1508]" captionTargetPageId="772" captionText="Figs 498-509. Cheiloneurus magnolia (498) head, facial view (modified); (499) fore wing; (500) apex of fore wing venation; (501) mesosoma, dorsal view; (502) antenna; (503) ovipositor; (504) gastral tergites, showing “glands” on Gt1 and Gt5; (505) hypopygium (posterior margin partly folded inwards). Cheiloneurus vitara, HT (506) antenna; (507) mesosoma, dorsal view; (508) apex of fore wing venation; (509) fore wing." pageId="217" pageNumber="216">Fig. 507</figureCitation>
) with imbricate-reticulate to polygonally reticulate sculpture on pronotum that is about as deep as that in ocellar area; mesoscutum with very fine, aciculate, longitudinally elongate, polygonally reticulate to striate sculpture of silky appearance that is about as deep as sculpture on pronotum, posterior part with more rounded, polygonally reticulate cells; scutellum with slightly deeper, polygonally reticulate sculpture composed of rounded or very slightly longitudinally elongate cells, extreme side and apex completely smooth; setae on anterior part of mesoscutum about as dense as those on posterior part; scutellum without a distinct, subapical tuft of longer setae; visible part of mesoscutum about 1.9X as broad as long; scutellum about 1.2X as broad as long; fore wing with venation and setation as in
<figureCitation id="13282A34FF0CFF7EFCFABF8BA5B6FC03" box="[792,934,942,967]" captionStart="Figs 498-509" captionStartId="772.[181,226,1526,1550]" captionTargetBox="[181,1163,225,1507]" captionTargetId="figure-9@772.[181,1163,223,1508]" captionTargetPageId="772" captionText="Figs 498-509. Cheiloneurus magnolia (498) head, facial view (modified); (499) fore wing; (500) apex of fore wing venation; (501) mesosoma, dorsal view; (502) antenna; (503) ovipositor; (504) gastral tergites, showing “glands” on Gt1 and Gt5; (505) hypopygium (posterior margin partly folded inwards). Cheiloneurus vitara, HT (506) antenna; (507) mesosoma, dorsal view; (508) apex of fore wing venation; (509) fore wing." pageId="217" pageNumber="216">Figs 508, 509</figureCitation>
; costal cell with 4 or 5 dorsal setae at apex. Relative measurements: FWL 113, FWW 47; HWL 79.5, HWW 20.
</paragraph>
<paragraph id="8BAC36B1FF0CFF7EFF39BFC0A782FBA2" blockId="217.[181,1165,222,1153]" pageId="217" pageNumber="216">Gaster with hypopygium reaching about 0.6X towards apex; last tergite about as long as mid tibia, with apex virtually truncate; “glands” present on Gt1 and Gt5, those on Gt1 separated by about 1.1X their own lengths, those on Gt5 touching; ovipositor slightly exserted, the exserted part about 0.6X length of mid tibial spur or about 0.25X mid tibia. Relative measurements: OL 58.5, GL 13 [MT 64].</paragraph>
<paragraph id="8BAC36B1FF0CFF7EFF39B84DA457FB44" blockId="217.[181,1165,222,1153]" box="[219,583,1129,1153]" pageId="217" pageNumber="216">
Variation. Only
<typeStatus id="54A88813FF0CFF7EFE9CB84DA7C7FB44" box="[382,471,1129,1153]" pageId="217" pageNumber="216">holotype</typeStatus>
examined.
</paragraph>
<paragraph id="8BAC36B1FF0CFF7EFF57B8BBA771FB72" blockId="217.[181,353,1183,1207]" box="[181,353,1183,1207]" pageId="217" pageNumber="216">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF0CFF7EFF57B8F0A7F9FB29" box="[181,489,1236,1260]" pageId="217" pageNumber="216" type="distribution">
<paragraph id="8BAC36B1FF0CFF7EFF57B8F0A7F9FB29" blockId="217.[181,489,1236,1260]" box="[181,489,1236,1260]" pageId="217" pageNumber="216">
DISTRIBUTION.
<collectingCountry id="F3047621FF0CFF7EFE91B8F0A7F5FB29" box="[371,485,1236,1260]" name="Costa Rica" pageId="217" pageNumber="216">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFF0CFF7EFF57B92DA76EFAE4" box="[181,382,1289,1313]" pageId="217" pageNumber="216" type="biology_ecology">
<paragraph id="8BAC36B1FF0CFF7EFF57B92DA76EFAE4" blockId="217.[181,382,1289,1313]" box="[181,382,1289,1313]" pageId="217" pageNumber="216">HOSTS. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C309653AFF0CFF7EFF57B964A7EEFA4E" pageId="217" pageNumber="216" type="materials_examined">
<paragraph id="8BAC36B1FF0CFF7EFF57B964A7BDFA90" blockId="217.[181,1164,1344,1419]" box="[181,429,1344,1365]" pageId="217" pageNumber="216">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FF0CFF7EFF39B97FA7EEFA4E" blockId="217.[181,1164,1344,1419]" pageId="217" pageNumber="216">
Type material.
<materialsCitation id="3B7B3CECFF0CFF7EFE87B97FA737FA4E" collectingDate="2004-02" collectorName="RB Hitoy Cerere &amp; J. S. Noyes" country="Costa Rica" elevation="100" latitude="9.666667" location="Limon" longLatPrecision="1299" longitude="-83.03333" pageId="217" pageNumber="216" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
<typeStatus id="54A88813FF0CFF7EFE87B97FA7A9FAB4" box="[357,441,1371,1393]" pageId="217" pageNumber="216">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FF0CFF7EFE3FB97FA478FAB4" bold="true" box="[477,616,1371,1393]" pageId="217" pageNumber="216">
<collectingCountry id="F3047621FF0CFF7EFE3FB97FA478FAB4" box="[477,616,1371,1393]" name="Costa Rica" pageId="217" pageNumber="216">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FF0CFF7EFD96B97EA4A4FAB5" bold="true" box="[628,692,1370,1392]" pageId="217" pageNumber="216">
<collectingRegion id="49D7F853FF0CFF7EFD96B97EA4A4FAB5" box="[628,692,1370,1392]" country="Costa Rica" name="Limon" pageId="217" pageNumber="216">Limón</collectingRegion>
</emphasis>
,
<collectorName id="26E65367FF0CFF7EFD22B97FA54EFAB5" box="[704,862,1371,1393]" pageId="217" pageNumber="216">RB Hitoy Cerere</collectorName>
,
<geoCoordinate id="EE275076FF0CFF7EFC85B97FA5BAFAB5" box="[871,938,1371,1392]" degrees="9" direction="north" minutes="40" orientation="latitude" pageId="217" pageNumber="216" precision="925" value="9.666667">9°40N</geoCoordinate>
<geoCoordinate id="EE275076FF0CFF7EFC52B97FA213FAB5" box="[944,1027,1371,1392]" degrees="83" direction="west" minutes="02" orientation="longitude" pageId="217" pageNumber="216" precision="925" value="-83.03333">83°02W</geoCoordinate>
,
<quantity id="4CEB9B54FF0CFF7EFBEFB97FA253FAB5" box="[1037,1091,1371,1392]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="217" pageNumber="216" unit="m" value="100.0">
<elevation id="003ED182FF0CFF7EFBEFB97FA253FAB5" box="[1037,1091,1371,1392]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="217" pageNumber="216" unit="m" value="100.0">100m</elevation>
</quantity>
,
<date id="FFAD1071FF0CFF7EFBAFB97FA29CFAB4" box="[1101,1164,1371,1393]" pageId="217" pageNumber="216" value="2004-02">
<collectingDate id="EFE9E999FF0CFF7EFBAFB97FA29CFAB4" box="[1101,1164,1371,1393]" pageId="217" pageNumber="216" value="2004-02">ii.2004</collectingDate>
</date>
(
<collectorName id="26E65367FF0CFF7EFF5FB952A731FA4E" box="[189,289,1398,1419]" pageId="217" pageNumber="216">J.S. Noyes</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFF0CFF7EFED2B951A7EBFA4E" box="[304,507,1397,1419]" collectionCode="NHMUK" country="Costa Rica" pageId="217" pageNumber="216" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
<typeStatus id="54A88813FF0CFF7EFED2B951A794FA4E" box="[304,388,1397,1419]" pageId="217" pageNumber="216">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FF0CFF7EFE40B952A7EBFA4E" box="[418,507,1398,1419]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="217" pageNumber="216" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>