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<document id="7574ADC0E4F6CE81E3D75A19F4A0D1B0" ID-DOI="10.5281/zenodo.8074943" ID-ISSN="2754-9844" ID-Zenodo-Dep="8074943" ID-ZooBank="BCAD06E8-0AFE-46ED-B7FA-930983CD44C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1699388683108" checkinUser="felipe" docAuthor="Noyes, John Stuart" docDate="2023" docId="03BA87A7FDB5FDC6FE3AB9AAA578FA04" docLanguage="en" docName="TaxMonogNeotropHymenopt.2.1-921.pdf" docOrigin="Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (1)" docSource="http://dx.doi.org/10.5281/zenodo.8074943" docStyle="DocumentStyle:DAA2ADC4E839520FEEF1E0A10B782A41.1:TaxMonogNeotropHymenopt.2022-.journal_article" docStyleId="DAA2ADC4E839520FEEF1E0A10B782A41" docStyleName="TaxMonogNeotropHymenopt.2022-.journal_article" docStyleVersion="1" docTitle="Psyllaephagus gyces Noyes &amp; Hanson 1996" docType="treatment" docVersion="2" lastPageNumber="608" masterDocId="FF83FFDFFFD5FFA7FFE2BC24A610FFC5" masterDocTitle="ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)" masterLastPageNumber="921" masterPageNumber="1" pageNumber="607" updateTime="1699637169064" updateUser="admin" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="A6651D5B6B5B6B0D2135045AE50DA7F2">ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera)</mods:title>
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<mods:namePart id="8FD76438717B9E4C63CFE30BB9A42B87">Noyes, John Stuart</mods:namePart>
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<mods:title id="02FBF84FCEC0A284BA06FA7F13E048FF">Taxonomic Monographs on Neotropical Hymenoptera</mods:title>
</mods:titleInfo>
<mods:part id="C4188E1DF6F9D9ECCB67C724BEAA8DE5">
<mods:date id="86502C8AD14AE49A9807137999C66109">2023</mods:date>
<mods:detail id="A6EDA177BD4495F4A169AB029B53A834" type="series">
<mods:title id="334D1D35659DEA74F0806FFE262E4192">Oxford, England</mods:title>
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<mods:number id="351A4EAF9738FAF159BC2087684F7D24">2023-06-30</mods:number>
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<mods:number id="5BC711C275E02F13096F7668E4307FF7">2</mods:number>
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<mods:url id="B51A386221B827560A85C785409BA212">http://dx.doi.org/10.5281/zenodo.8074943</mods:url>
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<mods:classification id="3F41ACFC36B7D706C5C66A1D8215FFB3">journal article</mods:classification>
<mods:identifier id="E73255B9748CBE33A46001E44E849D7C" type="DOI">10.5281/zenodo.8074943</mods:identifier>
<mods:identifier id="B04678E659DBC4059776722F33048320" type="ISSN">2754-9844</mods:identifier>
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<treatment id="03BA87A7FDB5FDC6FE3AB9AAA578FA04" LSID="urn:lsid:plazi:treatment:03BA87A7FDB5FDC6FE3AB9AAA578FA04" httpUri="http://treatment.plazi.org/id/03BA87A7FDB5FDC6FE3AB9AAA578FA04" lastPageId="609" lastPageNumber="608" pageId="608" pageNumber="607">
<subSubSection id="C309653AFDB5FDC7FE3AB9AAA52BFA04" pageId="608" pageNumber="607" type="nomenclature">
<paragraph id="8BAC36B1FDB5FDC7FE3AB9AAA579FA62" blockId="608.[472,873,1422,1473]" box="[472,873,1422,1447]" pageId="608" pageNumber="607">
<heading id="D0E481DDFDB5FDC7FE3AB9AAA579FA62" bold="true" box="[472,873,1422,1447]" centered="true" fontSize="10" level="2" pageId="608" pageNumber="607" reason="2">
<taxonomicName id="4C134D32FDB5FDC7FE3AB9AAA579FA62" ID-CoL="4PS38" authority="Noyes &amp; Hanson" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[472,873,1422,1447]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="608" pageNumber="607" phylum="Arthropoda" rank="species" species="gyces">
<emphasis id="B967EAA3FDB5FDC7FE3AB9AAA579FA62" bold="true" box="[472,873,1422,1447]" pageId="608" pageNumber="607">
<emphasis id="B967EAA3FDB5FDC7FE3AB9AAA4BDFA63" bold="true" box="[472,685,1422,1446]" italics="true" pageId="608" pageNumber="607">Psyllaephagus gyces</emphasis>
Noyes &amp; Hanson
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAC36B1FDB5FDC7FDE4B98DA52BFA04" blockId="608.[472,873,1422,1473]" box="[518,827,1449,1473]" pageId="608" pageNumber="607">(Figs 1620-1625; Hab. E 241)</paragraph>
</subSubSection>
<subSubSection id="C309653AFDB5FDC7FF57B9FAA777F9D4" pageId="608" pageNumber="607" type="reference_group">
<paragraph id="8BAC36B1FDB5FDC7FF57B9FAA777F9D4" blockId="608.[181,1164,1502,1553]" pageId="608" pageNumber="607">
<taxonomicName id="4C134D32FDB5FDC7FF57B9FAA499FA33" authority="Noyes &amp; Hanson, 1996" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[181,649,1502,1527]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="608" pageNumber="607" phylum="Arthropoda" rank="species" species="gyces">
<emphasis id="B967EAA3FDB5FDC7FF57B9FAA79AFA33" box="[181,394,1502,1526]" italics="true" pageId="608" pageNumber="607">Psyllaephagus gyces</emphasis>
<materialsCitation id="3B7B3CECFDB5FDC7FE70B9FBA499FA33" box="[402,649,1502,1527]" collectionCode="NHMUK" country="Costa Rica" location="Costa Rica" pageId="608" pageNumber="607" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EF824B40FDB5FDC7FE70B9FBA499FA33" author="Noyes, J. S. &amp; Hanson, P." box="[402,649,1502,1527]" pageId="608" pageNumber="607" pagination="105 - 164" refId="ref654282" refString="Noyes, J. S. &amp; Hanson, P. 1996. Encyrtidae (Hymenoptera: Chalcidoidea) of Costa Rica: the genera and species associated with jumping plant-lice (Homoptera: Psylloidea). Bulletin of the Natural History Museum, London (Entomology) 65 (2):`105 - 164." type="journal article" year="1996">Noyes &amp; Hanson, 1996</bibRefCitation>
</materialsCitation>
</taxonomicName>
:126,131-132.
<typeStatus id="54A88813FDB5FDC7FCFFB9FAA56BFA33" box="[797,891,1502,1526]" pageId="608" pageNumber="607">Holotype</typeStatus>
E,
<collectingCountry id="F3047621FDB5FDC7FC45B9FBA20AFA33" box="[935,1050,1502,1527]" name="Costa Rica" pageId="608" pageNumber="607">Costa Rica</collectingCountry>
, NHMUK, examined.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFDB5FDC6FF57BA0AA418FD64" lastPageId="609" lastPageNumber="608" pageId="608" pageNumber="607" type="description">
<paragraph id="8BAC36B1FDB5FDC7FF57BA0AA29CF953" blockId="608.[181,1165,1582,1687]" pageId="608" pageNumber="607">
<emphasis id="B967EAA3FDB5FDC7FF57BA0AA711F983" box="[181,257,1582,1606]" italics="true" pageId="608" pageNumber="607">Female</emphasis>
(length about
<quantity id="4CEB9B54FDB5FDC7FE7EBA0AA400F983" box="[412,528,1582,1606]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="608" pageNumber="607" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">1.2-1.6mm</quantity>
): head and mesoscutum generally dull metallic green or blue with some violet reflections, scutellum generally dull metallic green with apex slightly brighter; antenna (Fig. 1624) with scape mostly dark brown; flagellum brown; tegula dark brown; femora dark brown; tibiae mostly yellow with bases brownish; head (Fig. 1623) about 2.4X frontovertex
</paragraph>
<paragraph id="8BAC36B1FDB4FDC6FF57BCA6A6F3FF5A" blockId="609.[181,227,130,159]" box="[181,227,130,159]" pageId="609" pageNumber="608">
<pageNumber id="BBB83D67FDB4FDC6FF57BCA6A6F3FF5A" ambiguity="4" box="[181,227,130,159]" fuzzyness="0" pageId="609" pageNumber="608" score="12" value="608">
<pageTitle id="CB8CEED6FDB4FDC6FF57BCA6A6F3FF5A" box="[181,227,130,159]" pageId="609" pageNumber="608">608</pageTitle>
</pageNumber>
</paragraph>
<paragraph id="8BAC36B1FDB4FDC6FEB5BCA4A5FBFF5D" blockId="609.[343,1003,128,152]" box="[343,1003,128,152]" pageId="609" pageNumber="608">
<pageTitle id="CB8CEED6FDB4FDC6FEB5BCA4A5FBFF5D" box="[343,1003,128,152]" pageId="609" pageNumber="608">
Taxonomic Monographs on Neotropical
<taxonomicName id="4C134D32FDB4FDC6FD11BCA4A5C7FF5D" authority=", Volume" authorityName="Volume" box="[755,983,128,152]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="order">Hymenoptera, Volume</taxonomicName>
2
</pageTitle>
</paragraph>
<paragraph id="8BAC36B1FDB4FDC6FF57BCFAA418FD64" blockId="609.[181,1165,222,673]" pageId="609" pageNumber="608">
width and 2.6X as wide as mouth opening; frontovertex with piliferous punctures deep and conspicuous, largest clearly smaller than ocellus, but all much larger than eye facet; ocelli forming an angle of about 100°; posterior ocellus equidistant from eye and occipital margin; scrobes fairly deep, but separated from anterior ocellus by at least about 2X length of torulus; antenna (Fig. 1624) with scape about 3.7X as long as broad and slightly shorter than width of frontovertex; pedicel slightly longer than F1; funicle segments longer than broad except for F6 which is quadrate, linear sensilla on all segments; clava with segments becoming gradually shorter towards apex which is narrowly rounded; clypeal margin slightly convex, protuberant; mandible with two acute teeth and a truncation, upper tooth short; mesoscutum generally with a dull metallic violet and blue sheen; scutellum with a dull dark coppery purple and metallic blue sheen; mesoscutum and scutellum with similar polygonally reticulate sculpture, but that on scutellum clearly deeper, more sharply defined and more elongate; scutellum convex; fore wing (Fig. 1622) about 2.4X as long as broad, marginal vein (Fig. 1621) about 1.5X as long as broad; costal cell with only a single line of setae dorsally; linea clava more or less open; gaster without a distinct, strongly reticulate anteromedian area on Gt3; ovipositor (Fig. 1620) about 1.7X as long as mid tibia, slightly exserted, the exserted part about 0.5X as long as mid tibial spur to about as long; second valvifer with about 9 or 10 subapical setae.
<emphasis id="B967EAA3FDB4FDC6FE83BEADA785FD64" box="[353,405,649,673]" italics="true" pageId="609" pageNumber="608">Male</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFDB4FDC6FF57BE9BA229FD13" box="[181,1081,702,727]" pageId="609" pageNumber="608" type="distribution">
<paragraph id="8BAC36B1FDB4FDC6FF57BE9BA229FD13" blockId="609.[181,1081,702,727]" box="[181,1081,702,727]" pageId="609" pageNumber="608">
DISTRIBUTION.
<collectingCountry id="F3047621FDB4FDC6FE91BE9AA7D0FD13" box="[371,448,702,726]" name="Mexico" pageId="609" pageNumber="608">Mexico</collectingCountry>
(Trjapitzin et al., 2008),
<collectingCountry id="F3047621FDB4FDC6FD5DBE9BA53DFD13" box="[703,813,702,727]" name="Costa Rica" pageId="609" pageNumber="608">Costa Rica</collectingCountry>
(
<bibRefCitation id="EF824B40FDB4FDC6FCD8BE9BA23EFD13" author="Noyes, J. S. &amp; Hanson, P." box="[826,1070,702,727]" pageId="609" pageNumber="608" pagination="105 - 164" refId="ref654282" refString="Noyes, J. S. &amp; Hanson, P. 1996. Encyrtidae (Hymenoptera: Chalcidoidea) of Costa Rica: the genera and species associated with jumping plant-lice (Homoptera: Psylloidea). Bulletin of the Natural History Museum, London (Entomology) 65 (2):`105 - 164." type="journal article" year="1996">Noyes &amp; Hanson, 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C309653AFDB4FDC6FF57BED0A53BFCE2" pageId="609" pageNumber="608" type="biology_ecology">
<paragraph id="8BAC36B1FDB4FDC6FF57BED0A53BFCE2" blockId="609.[181,1164,756,807]" pageId="609" pageNumber="608">
HOSTS. Reared from
<taxonomicName id="4C134D32FDB4FDC6FE41BED0A4D8FCC9" authority="Enderlein" authorityName="Enderlein" box="[419,712,756,780]" class="Insecta" family="Triozidae" genus="Triozoida" kingdom="Animalia" order="Hemiptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="limbata">
<emphasis id="B967EAA3FDB4FDC6FE41BED0A44BFCC9" box="[419,603,756,780]" italics="true" pageId="609" pageNumber="608">Triozoida limbata</emphasis>
Enderlein
</taxonomicName>
(
<taxonomicName id="4C134D32FDB4FDC6FD3BBED0A55AFCC9" box="[729,842,756,780]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="order">Hemiptera</taxonomicName>
:
<taxonomicName id="4C134D32FDB4FDC6FCB4BED0A5A8FCC9" box="[854,952,756,780]" class="Insecta" family="Triozidae" kingdom="Animalia" order="Hemiptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="family">Triozidae</taxonomicName>
) on guava,
<taxonomicName id="4C134D32FDB4FDC6FBDBBED0A717FCE3" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" kingdom="Plantae" order="Myrtales" pageId="609" pageNumber="608" phylum="Tracheophyta" rank="species" species="guajava">
<emphasis id="B967EAA3FDB4FDC6FBDBBED0A717FCE3" italics="true" pageId="609" pageNumber="608">Psidium guajava</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C134D32FDB4FDC6FED3BF2AA780FCE3" box="[305,400,782,806]" class="Magnoliopsida" kingdom="Plantae" order="Myrtales" pageId="609" pageNumber="608" phylum="Tracheophyta" rank="order">Myrtales</taxonomicName>
:
<taxonomicName id="4C134D32FDB4FDC6FE7BBF2BA416FCE2" box="[409,518,783,807]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="609" pageNumber="608" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) (see Trjapitzin et al., 2008).
</paragraph>
</subSubSection>
<subSubSection id="C309653AFDB4FDC6FF57BF62A29DFBD0" pageId="609" pageNumber="608" type="materials_examined">
<paragraph id="8BAC36B1FDB4FDC6FF57BF62A7BDFC9E" blockId="609.[181,1165,838,1046]" box="[181,429,838,859]" pageId="609" pageNumber="608">MATERIAL EXAMINED.</paragraph>
<paragraph id="8BAC36B1FDB4FDC6FF39BF45A4ADFC6E" blockId="609.[181,1165,838,1046]" pageId="609" pageNumber="608">
Type material.
<materialsCitation id="3B7B3CECFDB4FDC6FE85BF44A233FCB3" box="[359,1059,864,886]" collectingDate="1990-03-20" collectorName="J. S. Noyes" country="Costa Rica" location="Golfito" pageId="609" pageNumber="608" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54A88813FDB4FDC6FE85BF44A7ABFCB3" box="[359,443,864,886]" pageId="609" pageNumber="608">Holotype</typeStatus>
E:
<emphasis id="B967EAA3FDB4FDC6FE01BF44A47FFCB3" bold="true" box="[483,623,864,886]" pageId="609" pageNumber="608">
<collectingCountry id="F3047621FDB4FDC6FE01BF44A47FFCB3" box="[483,623,864,886]" name="Costa Rica" pageId="609" pageNumber="608">COSTA RICA</collectingCountry>
</emphasis>
,
<emphasis id="B967EAA3FDB4FDC6FD9EBF44A4FAFCB0" bold="true" box="[636,746,864,885]" pageId="609" pageNumber="608">
<collectingRegion id="49D7F853FDB4FDC6FD9EBF44A4FAFCB0" box="[636,746,864,885]" country="Costa Rica" name="Puntarenas" pageId="609" pageNumber="608">Puntarenas</collectingRegion>
</emphasis>
,
<location id="8ECC606AFDB4FDC6FD1ABF44A52BFCB3" LSID="urn:lsid:plazi:treatment:03BA87A7FDB5FDC6FE3AB9AAA578FA04:8ECC606AFDB4FDC6FD1ABF44A52BFCB3" box="[760,827,864,886]" country="Costa Rica" name="Golfito" pageId="609" pageNumber="608" stateProvince="Puntarenas">Golfito</location>
,
<date id="FFAD1071FDB4FDC6FCA7BF44A5B8FCB3" box="[837,936,864,886]" pageId="609" pageNumber="608" value="1990-03-20">
<collectingDate id="EFE9E999FDB4FDC6FCA7BF44A5B8FCB3" box="[837,936,864,886]" pageId="609" pageNumber="608" value="1990-03-20">20.iii.1990</collectingDate>
</date>
(
<collectorName id="26E65367FDB4FDC6FC55BF44A20DFCB3" box="[951,1053,864,886]" pageId="609" pageNumber="608">J.S. Noyes</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFDB4FDC6FBCCBF45A299FC55" collectingDate="viii. 1991 and xi. 1991" collectorName="P. Hanson" country="Costa Rica" elevation="10" latitude="8.533334" location="Puerto Jimenez" longLatPrecision="1301" longitude="-83.316666" pageId="609" pageNumber="608" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<typeStatus id="54A88813FDB4FDC6FBCCBF45A29AFCB3" box="[1070,1162,865,886]" pageId="609" pageNumber="608">Paratypes</typeStatus>
:
<emphasis id="B967EAA3FDB4FDC6FF57BF5FA750FC54" bold="true" box="[181,320,891,913]" pageId="609" pageNumber="608">
<collectingCountry id="F3047621FDB4FDC6FF57BF5FA750FC54" box="[181,320,891,913]" name="Costa Rica" pageId="609" pageNumber="608">COSTA RICA</collectingCountry>
</emphasis>
, 2E,
<emphasis id="B967EAA3FDB4FDC6FE91BF5FA7F1FC55" bold="true" box="[371,481,891,912]" pageId="609" pageNumber="608">
<collectingRegion id="49D7F853FDB4FDC6FE91BF5FA7F1FC55" box="[371,481,891,912]" country="Costa Rica" name="Puntarenas" pageId="609" pageNumber="608">Puntarenas</collectingRegion>
</emphasis>
,
<location id="8ECC606AFDB4FDC6FE0EBF5FA46AFC54" LSID="urn:lsid:plazi:treatment:03BA87A7FDB5FDC6FE3AB9AAA578FA04:8ECC606AFDB4FDC6FE0EBF5FA46AFC54" box="[492,634,891,913]" country="Costa Rica" latitude="8.533334" longLatPrecision="1301" longitude="-83.316666" name="Puerto Jimenez" pageId="609" pageNumber="608" stateProvince="Puntarenas">Puerto Jimenez</location>
,
<geoCoordinate id="EE275076FDB4FDC6FD60BF5FA4D1FC55" box="[642,705,891,912]" degrees="8" direction="north" minutes="32" orientation="latitude" pageId="609" pageNumber="608" precision="925" value="8.533334">8 32’N</geoCoordinate>
<geoCoordinate id="EE275076FDB4FDC6FD25BF5FA50AFC55" box="[711,794,891,912]" degrees="83" direction="west" minutes="19" orientation="longitude" pageId="609" pageNumber="608" precision="925" value="-83.316666">83°19’W</geoCoordinate>
,
<quantity id="4CEB9B54FDB4FDC6FCC1BF5FA55EFC55" box="[803,846,891,912]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="609" pageNumber="608" unit="m" value="10.0">
<elevation id="003ED182FDB4FDC6FCC1BF5FA55EFC55" box="[803,846,891,912]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="609" pageNumber="608" unit="m" value="10.0">10m</elevation>
</quantity>
,
<collectingDate id="EFE9E999FDB4FDC6FCB4BF5FA208FC54" box="[854,1048,891,913]" isEnumeration="true" pageId="609" pageNumber="608">
<date id="FFAD1071FDB4FDC6FCB4BF5FA5B8FC54" box="[854,936,891,913]" pageId="609" pageNumber="608" value="1991-08">viii.1991</date>
and
<date id="FFAD1071FDB4FDC6FC31BF5FA208FC54" box="[979,1048,891,913]" pageId="609" pageNumber="608" value="1991-11">xi.1991</date>
</collectingDate>
(
<collectorName id="26E65367FDB4FDC6FBC6BF5FA293FC55" box="[1060,1155,891,912]" pageId="609" pageNumber="608">P. Hanson</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B7B3CECFDB4FDC6FF57BFB1A4A9FC6E" box="[181,697,917,939]" collectionCode="NHMUK, INBio" country="Costa Rica" pageId="609" pageNumber="608" specimenCount="2" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54A88813FDB4FDC6FF57BFB1A719FC6E" box="[181,265,917,939]" pageId="609" pageNumber="608">Holotype</typeStatus>
in
<collectionCode id="ED02AE74FDB4FDC6FEC5BFB2A76FFC6E" box="[295,383,918,939]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="609" pageNumber="608" type="Museum">NHMUK</collectionCode>
,
<typeStatus id="54A88813FDB4FDC6FE6ABFB2A7CFFC6E" box="[392,479,918,939]" pageId="609" pageNumber="608">paratypes</typeStatus>
in
<collectionCode id="ED02AE74FDB4FDC6FE1EBFB1A425FC6E" box="[508,565,917,939]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="609" pageNumber="608">INBio</collectionCode>
and
<collectionCode id="ED02AE74FDB4FDC6FD82BFB2A4A9FC6E" box="[608,697,918,939]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="609" pageNumber="608" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC36B1FDB4FDC6FF39BF95A29DFBD0" blockId="609.[181,1165,838,1046]" pageId="609" pageNumber="608">
<materialsCitation id="3B7B3CECFDB4FDC6FF39BF95A446FC3E" collectingDate="1990-03" collectingDateMax="2003-07-28" collectingDateMin="1990-03" collectionCode="LN" collectorName="La Chiles &amp; P. Hanson &amp; Alajuela, PN &amp; Est. El Pilon &amp; A. Azofeifa &amp; Alajuela &amp; Arenal &amp; Sendero Pilon" country="Costa Rica" county="Non" elevation="750" location="La Chiles" municipality="Aguas Zarcas" pageId="609" pageNumber="608" specimenCode="LN 269100" specimenCount="1" stateProvince="Alajuela">
<collectingCounty id="62CD4E3DFDB4FDC6FF39BF95A712FC03" box="[219,258,945,966]" pageId="609" pageNumber="608">Non</collectingCounty>
<typeStatus id="54A88813FDB4FDC6FEEBBF95A720FC03" box="[265,304,945,966]" pageId="609" pageNumber="608">type</typeStatus>
material.
<emphasis id="B967EAA3FDB4FDC6FE6CBF94A40AFC03" bold="true" box="[398,538,944,966]" pageId="609" pageNumber="608">
<collectingCountry id="F3047621FDB4FDC6FE6CBF94A40AFC03" box="[398,538,944,966]" name="Costa Rica" pageId="609" pageNumber="608">COSTA RICA</collectingCountry>
</emphasis>
, 1E,
<emphasis id="B967EAA3FDB4FDC6FDB2BF94A4B2FC03" bold="true" box="[592,674,944,966]" pageId="609" pageNumber="608">
<collectingRegion id="49D7F853FDB4FDC6FDB2BF94A4B2FC03" box="[592,674,944,966]" country="Costa Rica" name="Alajuela" pageId="609" pageNumber="608">Alajuela</collectingRegion>
</emphasis>
,
<collectingMunicipality id="6BC8ACCBFDB4FDC6FD4FBF94A53DFC03" box="[685,813,944,966]" pageId="609" pageNumber="608">Aguas Zarcas</collectingMunicipality>
,
<collectorName id="26E65367FDB4FDC6FCD5BF95A581FC03" box="[823,913,944,966]" pageId="609" pageNumber="608">
<location id="8ECC606AFDB4FDC6FCD5BF95A581FC03" LSID="urn:lsid:plazi:treatment:03BA87A7FDB5FDC6FE3AB9AAA578FA04:8ECC606AFDB4FDC6FCD5BF95A581FC03" box="[823,913,944,966]" country="Costa Rica" county="Non" municipality="Aguas Zarcas" name="La Chiles" pageId="609" pageNumber="608" stateProvince="Alajuela">La Chiles</location>
</collectorName>
, café,
<date id="FFAD1071FDB4FDC6FC2CBF94A204FC03" box="[974,1044,944,966]" pageId="609" pageNumber="608" value="1990-03">
<collectingDate id="EFE9E999FDB4FDC6FC2CBF94A204FC03" box="[974,1044,944,966]" pageId="609" pageNumber="608" value="1990-03">iii.1990</collectingDate>
</date>
(
<collectorName id="26E65367FDB4FDC6FBC3BF94A292FC03" box="[1057,1154,944,966]" pageId="609" pageNumber="608">P. Hanson</collectorName>
); 1E,
<collectorName id="26E65367FDB4FDC6FF3EBFEFA741FC25" box="[220,337,971,993]" pageId="609" pageNumber="608">Alajuela, PN</collectorName>
Volcán Tenorio,
<collectorName id="26E65367FDB4FDC6FE0EBFEFA470FC24" box="[492,608,971,993]" pageId="609" pageNumber="608">Est. El Pilón</collectorName>
,
<quantity id="4CEB9B54FDB4FDC6FD8BBFEFA4D9FC25" box="[617,713,971,992]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="609" pageNumber="608" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">
<elevation id="003ED182FDB4FDC6FD8BBFEFA4D9FC25" box="[617,713,971,992]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="609" pageNumber="608" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">700-800m</elevation>
</quantity>
, #74622,
<date id="FFAD1071FDB4FDC6FCC3BFEFA599FC24" box="[801,905,971,993]" pageId="609" pageNumber="608" value="2003-07-28">
<collectingDate id="EFE9E999FDB4FDC6FCC3BFEFA599FC24" box="[801,905,971,993]" pageId="609" pageNumber="608" value="2003-07-28">28.vii.2003</collectingDate>
</date>
(
<collectorName id="26E65367FDB4FDC6FC77BFEFA214FC24" box="[917,1028,971,993]" pageId="609" pageNumber="608">A. Azofeifa</collectorName>
); 2E,
<collectorName id="26E65367FDB4FDC6FBD8BFEFA299FC24" box="[1082,1161,971,993]" pageId="609" pageNumber="608">Alajuela</collectorName>
, RF
<collectorName id="26E65367FDB4FDC6FF34BFC1A706FC3E" box="[214,278,997,1019]" pageId="609" pageNumber="608">Arenal</collectorName>
,
<collectorName id="26E65367FDB4FDC6FEC2BFC1A7B3FC3E" box="[288,419,997,1019]" pageId="609" pageNumber="608">Sendero Pilón</collectorName>
,
<quantity id="4CEB9B54FDB4FDC6FE4FBFC1A7F3FC3E" box="[429,483,997,1019]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="609" pageNumber="608" unit="m" value="600.0">
<elevation id="003ED182FDB4FDC6FE4FBFC1A7F3FC3E" box="[429,483,997,1019]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="609" pageNumber="608" unit="m" value="600.0">600m</elevation>
</quantity>
,
<specimenCode id="DBB59ECAFDB4FDC6FE0FBFC2A446FC3E" box="[493,598,997,1019]" collectionCode="LN" pageId="609" pageNumber="608">LN 269100</specimenCode>
</materialsCitation>
<materialsCitation id="3B7B3CECFDB4FDC6FDBEBFC1A29AFBD0" collectingDate="14. x- 3. xii. 1998 and 17 - 18. v. 1999" collectionCode="NHMUK, MZUCR" collectorName="G. Carballo &amp; P. Hanson" country="Costa Rica" elevation="10" latitude="8.533334" location="Puerto Jimenez" longLatPrecision="1301" longitude="-83.35" pageId="609" pageNumber="608" specimenCount="1" stateProvince="Puntarenas">
457900,
<collectingDate id="EFE9E999FDB4FDC6FD4EBFC1A5CDFC3E" box="[684,989,997,1019]" isEnumeration="true" pageId="609" pageNumber="608">
<date id="FFAD1071FDB4FDC6FD4EBFC1A527FC3E" box="[684,823,997,1019]" pageId="609" pageNumber="608" value="1998-10-14" valueMax="1998-12-03" valueMin="1998-10-14">14.x-3.xii.1998</date>
and
<date id="FFAD1071FDB4FDC6FC87BFC2A5CDFC3E" box="[869,989,998,1019]" pageId="609" pageNumber="608" value="1999-05-17" valueMax="1999-05-18" valueMin="1999-05-17">17-18.v.1999</date>
</collectingDate>
(
<collectorName id="26E65367FDB4FDC6FC08BFC2A249FC3E" box="[1002,1113,997,1019]" pageId="609" pageNumber="608">G. Carballo</collectorName>
); 1E,
<emphasis id="B967EAA3FDB4FDC6FF57B824A733FBD0" bold="true" box="[181,291,1024,1045]" pageId="609" pageNumber="608">
<collectingRegion id="49D7F853FDB4FDC6FF57B824A733FBD0" box="[181,291,1024,1045]" country="Russia" name="Puntarenas" pageId="609" pageNumber="608">Puntarenas</collectingRegion>
</emphasis>
,
<location id="8ECC606AFDB4FDC6FECDB825A7ACFBD3" LSID="urn:lsid:plazi:treatment:03BA87A7FDB5FDC6FE3AB9AAA578FA04:8ECC606AFDB4FDC6FECDB825A7ACFBD3" box="[303,444,1024,1046]" country="Costa Rica" latitude="8.533334" longLatPrecision="1301" longitude="-83.35" name="Puerto Jimenez" pageId="609" pageNumber="608" stateProvince="Puntarenas">Puerto Jimenez</location>
,
<geoCoordinate id="EE275076FDB4FDC6FE26B824A417FBD0" box="[452,519,1024,1045]" degrees="8" direction="north" minutes="32" orientation="latitude" pageId="609" pageNumber="608" precision="925" value="8.533334">8°32’N</geoCoordinate>
<geoCoordinate id="EE275076FDB4FDC6FDEEB824A470FBD0" box="[524,608,1024,1045]" degrees="83" direction="west" minutes="21" orientation="longitude" pageId="609" pageNumber="608" precision="925" value="-83.35">83°21’W</geoCoordinate>
,
<quantity id="4CEB9B54FDB4FDC6FD8AB824A483FBD0" box="[616,659,1024,1045]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="609" pageNumber="608" unit="m" value="10.0">
<elevation id="003ED182FDB4FDC6FD8AB824A483FBD0" box="[616,659,1024,1045]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="609" pageNumber="608" unit="m" value="10.0">10m</elevation>
</quantity>
, x-xi.1991 (
<collectorName id="26E65367FDB4FDC6FD1DB824A54EFBD3" box="[767,862,1024,1046]" pageId="609" pageNumber="608">P. Hanson</collectorName>
). Material in
<collectionCode id="ED02AE74FDB4FDC6FC37B825A23DFBD3" box="[981,1069,1025,1046]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="609" pageNumber="608" type="Museum">NHMUK</collectionCode>
,
<collectionCode id="ED02AE74FDB4FDC6FBD4B824A29AFBD0" box="[1078,1162,1024,1045]" pageId="609" pageNumber="608">MZUCR</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C309653AFDB4FDC6FF57B810A578FA04" pageId="609" pageNumber="608" type="discussion">
<paragraph id="8BAC36B1FDB4FDC6FF57B810A578FA04" blockId="609.[181,1165,1076,1473]" pageId="609" pageNumber="608">
COMMENTS. Females of
<taxonomicName id="4C134D32FDB4FDC6FE33B810A4B7FB8E" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[465,679,1076,1100]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="gyces">
<emphasis id="B967EAA3FDB4FDC6FE33B810A4B7FB8E" box="[465,679,1076,1100]" italics="true" pageId="609" pageNumber="608">Psyllaephagus gyces</emphasis>
</taxonomicName>
are very similar to
<taxonomicName id="4C134D32FDB4FDC6FC9FB810A5D0FB8E" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[893,960,1076,1099]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="creusa">
<emphasis id="B967EAA3FDB4FDC6FC9FB810A5D0FB8E" box="[893,960,1076,1099]" italics="true" pageId="609" pageNumber="608">creusa</emphasis>
</taxonomicName>
, both having deep, conspicuous piliferous punctures on the frontovertex and very similar colouration. Apart from the more extensive brown marking on the hind tibia of
<taxonomicName id="4C134D32FDB4FDC6FD2DB84EA502FB44" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[719,786,1130,1153]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="creusa">
<emphasis id="B967EAA3FDB4FDC6FD2DB84EA502FB44" box="[719,786,1130,1153]" italics="true" pageId="609" pageNumber="608">creusa</emphasis>
</taxonomicName>
the two species can be separated on body size, relative size of punctures on the frontovertex, relative position of the ocelli, scutellum sculpture and the ovipositor structure. Females of
<taxonomicName id="4C134D32FDB4FDC6FD22B8BBA4E8FB73" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[704,760,1183,1206]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="gyces">
<emphasis id="B967EAA3FDB4FDC6FD22B8BBA4E8FB73" box="[704,760,1183,1206]" italics="true" pageId="609" pageNumber="608">gyces</emphasis>
</taxonomicName>
are generally not more than
<quantity id="4CEB9B54FDB4FDC6FBCBB8BAA260FB73" box="[1065,1136,1182,1206]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="609" pageNumber="608" unit="mm" value="1.6">1.6mm</quantity>
in length, the piliferous punctures on the frontovertex are much smaller than the diameter of an ocellus, the posterior ocellus is equidistant from the occipital margin and eye, the sculpture of the scutellum is clearly deeper than that on the mesoscutum, the darker, coppery purple colouration of the scutellum is limited to the ridges of the cells and the ovipositor is longer than the mid tibia and also slightly exserted. In
<taxonomicName id="4C134D32FDB4FDC6FE0EB901A43FFAF9" authorityName="Noyes &amp; Hanson" authorityYear="1996" box="[492,559,1317,1340]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="creusa">
<emphasis id="B967EAA3FDB4FDC6FE0EB901A43FFAF9" box="[492,559,1317,1340]" italics="true" pageId="609" pageNumber="608">creusa</emphasis>
</taxonomicName>
the females are mostly longer than
<quantity id="4CEB9B54FDB4FDC6FC4EB900A5E6FAF9" box="[940,1014,1316,1340]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="609" pageNumber="608" unit="mm" value="1.6">1.6mm</quantity>
, the piliferous punctures on the frontovertex are mostly subequal in diameter to an ocellus, the posterior ocellus is nearer the occipital margin than the eye, the sculpture of the scutellum is hardly deeper than that on the mesoscutum and the darker coppery purple colouration is not limited to the ridges of the cells and the ovipositor is shorter than the mid tibia and hidden. See also comments under
<taxonomicName id="4C134D32FDB4FDC6FF57B98DA703FA04" authorityName="Noyes" authorityYear="2023" box="[181,275,1449,1473]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="kallichoa">
<emphasis id="B967EAA3FDB4FDC6FF57B98DA703FA04" box="[181,275,1449,1473]" italics="true" pageId="609" pageNumber="608">kallichoa</emphasis>
</taxonomicName>
(p. 592),
<taxonomicName id="4C134D32FDB4FDC6FE95B98EA7B7FA04" authorityName="Noyes" authorityYear="2023" box="[375,423,1450,1473]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="xepo">
<emphasis id="B967EAA3FDB4FDC6FE95B98EA7B7FA04" box="[375,423,1450,1473]" italics="true" pageId="609" pageNumber="608">xepo</emphasis>
</taxonomicName>
(p. 597),
<taxonomicName id="4C134D32FDB4FDC6FDE8B98DA452FA04" authorityName="Noyes" authorityYear="2023" box="[522,578,1449,1473]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="korus">
<emphasis id="B967EAA3FDB4FDC6FDE8B98DA452FA04" box="[522,578,1449,1473]" italics="true" pageId="609" pageNumber="608">korus</emphasis>
</taxonomicName>
(p. 602) and
<taxonomicName id="4C134D32FDB4FDC6FD28B98DA51BFA04" box="[714,779,1449,1473]" class="Insecta" family="Encyrtidae" genus="Psyllaephagus" kingdom="Animalia" order="Hymenoptera" pageId="609" pageNumber="608" phylum="Arthropoda" rank="species" species="phelys">
<emphasis id="B967EAA3FDB4FDC6FD28B98DA51BFA04" box="[714,779,1449,1473]" italics="true" pageId="609" pageNumber="608">phelys</emphasis>
</taxonomicName>
(p. 607).
</paragraph>
</subSubSection>
</treatment>
</document>
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