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<document id="B972C92E19EC2C0CA2E982DE29AF3E68" ID-DOI="10.5281/zenodo.5395859" ID-ISSN="1638-9387" ID-Zenodo-Dep="5395859" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630325111336" checkinUser="marcus" docAuthor="Pace, Roberto" docDate="2008" docId="03BA8792FFE8972572ADFA42FF58FA28" docLanguage="fr" docName="Zoosystema.2008.30.3.555-577.pdf" docOrigin="Zoosystema 30 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Placusa gomyi Pace 1984" docType="treatment" docVersion="3" lastPageNumber="572" masterDocId="FF83FFEAFFE79734725CFF88FF84FE23" masterDocTitle="Aleocharinae (Insecta, Coleoptera, Staphylinidae) de Madagascar" masterLastPageNumber="577" masterPageNumber="555" pageNumber="570" updateTime="1699227851486" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C85C91D3226AE3E73F2A6E5486F6197E">Aleocharinae (Insecta, Coleoptera, Staphylinidae) de Madagascar</mods:title>
</mods:titleInfo>
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<mods:namePart id="95C900DB68B07824D93C243DF0F0F7D4">Pace, Roberto</mods:namePart>
<mods:affiliation id="79B98E8421364DCBD67B3E41E890CF59">Via Vittorio Veneto, 13, I- 37032 Monteforte dAlpone, Verona (Italie) pace. ent @ tiscali. it</mods:affiliation>
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<mods:title id="570D48416D6B75E35D3337BA29566A37">Zoosystema</mods:title>
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<mods:part id="9E69CE2459198E8B4A9CAF421DF5025D">
<mods:date id="D0AF30A95B7877EAB0B30B1491C6BA23">2008</mods:date>
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<mods:number id="6A47BCCEE9EFB33C7B7ABE963416BF60">30</mods:number>
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<mods:number id="2871239C5A82A58AF952DE2378FAB902">3</mods:number>
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<mods:identifier id="81DE4D4CDEB4B7B0255856AA883DB444" type="DOI">10.5281/zenodo.5395859</mods:identifier>
<mods:identifier id="4F1E4E22806DC0886B0593CA0ADD2F57" type="ISSN">1638-9387</mods:identifier>
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<treatment id="03BA8792FFE8972572ADFA42FF58FA28" LSID="urn:lsid:plazi:treatment:03BA8792FFE8972572ADFA42FF58FA28" httpUri="http://treatment.plazi.org/id/03BA8792FFE8972572ADFA42FF58FA28" lastPageId="17" lastPageNumber="572" pageId="15" pageNumber="570">
<subSubSection id="C309650FFFE8973B72ADFA42FD83FBC5" box="[241,519,1482,1510]" pageId="15" pageNumber="570" type="nomenclature">
<paragraph id="8BAC3684FFE8973B72ADFA42FD83FBC5" blockId="15.[241,519,1482,1510]" box="[241,519,1482,1510]" pageId="15" pageNumber="570">
<heading id="D0E481E8FFE8973B72ADFA42FD83FBC5" box="[241,519,1482,1510]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="570" reason="2">
<taxonomicName id="4C134D07FFE8973B72ADFA42FD83FBC5" ID-CoL="77MY7" authority="Pace, 1984" authorityName="Pace" authorityYear="1984" box="[241,519,1482,1510]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="570" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="B967EA96FFE8973B72ADFA42FE0EFBC6" bold="true" box="[241,394,1482,1509]" italics="true" pageId="15" pageNumber="570">Placusa gomyi</emphasis>
Pace, 1984
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309650FFFE8973B722DF980FEDBF84C" pageId="15" pageNumber="570" type="reference_group">
<paragraph id="8BAC3684FFE8973B722DF980FD96F802" blockId="15.[113,530,1544,1569]" box="[113,530,1544,1569]" pageId="15" pageNumber="570">
<treatmentCitationGroup id="AB0311AAFFE8973B722DF980FD96F802" box="[113,530,1544,1569]" pageId="15" pageNumber="570">
<treatmentCitation id="0AB21095FFE8973B722DF980FE26F802" author="PACE R." box="[113,418,1544,1569]" page="526" pageId="15" pageNumber="570" year="1984">
<taxonomicName id="4C134D07FFE8973B722DF980FE26F802" ID-CoL="77MY7" authority="Pace, 1984 c: 526" authorityName="Pace" authorityPageNumber="526" authorityYear="1984" box="[113,418,1544,1569]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="570" phylum="Arthropoda" rank="species" species="gomyi">
<emphasis id="B967EA96FFE8973B722DF980FF71F803" box="[113,245,1544,1568]" italics="true" pageId="15" pageNumber="570">Placusa gomyi</emphasis>
<bibRefCitation id="EF824B75FFE8973B72A7F981FE26F802" author="PACE R." box="[251,418,1545,1569]" pageId="15" pageNumber="570" pagination="521 - 543" refId="ref9666" refString="PACE R. 1984 c. - Aleocharinae del Madagascar raccolte da Yves Gomy (Coleoptera Staphylinidae) (XLVIII Contributo alla conoscenza delle Aleocharinae). Revue suisse de Zoologie 91: 521 - 543, 92 figs." type="journal article" year="1984">Pace, 1984c: 526</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB21095FFE8973B73EEF981FD8AF802" author="PACE R." box="[434,526,1545,1569]" page="46" pageId="15" pageNumber="570" year="1999">
<bibRefCitation id="EF824B75FFE8973B73EEF981FD8AF802" author="PACE R." box="[434,526,1545,1569]" pageId="15" pageNumber="570" refId="ref9743" refString="PACE R. 1999. - Insectes Coleopteres Staphylinidae Aleocharinae. Faune de Madagascar 89. Paris, 261 p." type="book" year="1999">1999: 46</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAC3684FFE8973B722DF9B5FEDBF84C" blockId="15.[112,647,1597,1647]" pageId="15" pageNumber="570">
MATÉRIEL EXAMINÉ. —
<emphasis id="B967EA96FFE8973B730CF9B5FE68F876" bold="true" box="[336,492,1597,1621]" pageId="15" pageNumber="570">
<collectingCountry id="F3047614FFE8973B730CF9B5FE47F876" box="[336,451,1597,1621]" name="Madagascar" pageId="15" pageNumber="570">Madagascar</collectingCountry>
Est.
</emphasis>
Ambodivoangy,
<date id="FFAD1044FFE8973B722CF9DFFF41F84C" box="[112,197,1623,1647]" pageId="15" pageNumber="570" value="1945-06">VI.1945</date>
, J. Vadon,
<specimenCount id="9D15FD0DFFE8973B736CF9DFFEDDF84D" box="[304,345,1623,1647]" pageId="15" pageNumber="570" type="male">1 ♂</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C309650FFFE8973B722DF903FC69FF30" pageId="15" pageNumber="570" type="description">
<paragraph id="8BAC3684FFE8973B722DF903FE39F880" blockId="15.[113,445,1675,1699]" box="[113,445,1675,1699]" pageId="15" pageNumber="570">
RÉPARTITION. —
<collectingCountry id="F3047614FFE8973B7342F903FE15F880" box="[286,401,1675,1699]" name="Madagascar" pageId="15" pageNumber="570">Madagascar</collectingCountry>
Est.
</paragraph>
<paragraph id="8BAC3684FFE8973B70A8FF5FFC69FF30" blockId="15.[756,1130,215,275]" pageId="15" pageNumber="570">
<taxonomicName id="4C134D07FFE8973B70A8FF5FFBADFED1" authority="Pace, 2008" authorityName="Pace" authorityYear="2008" box="[756,1065,215,243]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="570" phylum="Arthropoda" rank="species" species="malgascia" status="sp. nov." subGenus="Calpusa">
<emphasis id="B967EA96FFE8973B70A8FF5FFCC3FED1" bold="true" box="[756,839,215,242]" italics="true" pageId="15" pageNumber="570">Placusa</emphasis>
(
<emphasis id="B967EA96FFE8973B710BFF5FFC2BFED1" bold="true" box="[855,943,215,242]" italics="true" pageId="15" pageNumber="570">Calpusa</emphasis>
)
<emphasis id="B967EA96FFE8973B71E3FF5FFBADFED1" bold="true" box="[959,1065,215,242]" italics="true" pageId="15" pageNumber="570">malgascia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457EDFFE8973B7673FF50FBEEFED0" box="[1071,1130,216,243]" pageId="15" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13282A01FFE8973B7125FF70FC33FF30" box="[889,951,248,275]" captionStart="FIG" captionStartId="14.[113,124,1659,1676]" captionTargetBox="[124,1195,221,1586]" captionTargetId="figure-16@14.[98,1224,179,1634]" captionTargetPageId="14" captionText="FIG. 7. — Habitus (A, C, G), spermathèque (B, F, J) et édéage en vues latérale (D, H) et ventrale (E, I): A, B, Brachida hova n. sp.; C-F, Placusa (Calpusa) malgascia n. sp.; G-J, Placusa (Calpusa) madecassa n. sp. Échelles: A, C, G, 1 mm; B, D-F, H-J, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.5395881" httpUri="https://zenodo.org/record/5395881/files/figure.png" pageId="15" pageNumber="570">Fig. 7</figureCitation>
C-F)
</paragraph>
</subSubSection>
<subSubSection id="C309650FFFE8973B70F9FEA3FC32FCE7" pageId="15" pageNumber="570" type="materials_examined">
<paragraph id="8BAC3684FFE8973B70F9FEA3FC9BFF54" blockId="15.[674,1211,298,708]" pageId="15" pageNumber="570">
<materialsCitation id="3B7B3CD9FFE8973B70F9FEA3FC9BFF54" collectingDate="1972-04-27" collectionCode="CNRS" collectorName="J. - M. Betsch" country="Madagascar" elevation="2600" location="Boby" pageId="15" pageNumber="570" specimenCode="RCP 225" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A88826FFE8973B70F9FEA3FC8AFF62" box="[677,782,299,322]" pageId="15" pageNumber="570" type="holotype">HOLOTYPE</typeStatus>
. —
<emphasis id="B967EA96FFE8973B7160FEA3FB80FF61" bold="true" box="[828,1028,298,323]" pageId="15" pageNumber="570">
<collectingCountry id="F3047614FFE8973B7160FEA3FC36FF60" box="[828,946,299,323]" name="Madagascar" pageId="15" pageNumber="570">Madagascar</collectingCountry>
Centre.
</emphasis>
[Andringitra], pic
<location id="8ECC605FFFE8973B70FAFECDFD58FF7E" LSID="urn:lsid:plazi:treatment:03BA8792FFE8972572ADFA42FF58FA28:8ECC605FFFE8973B70FAFECDFD58FF7E" box="[678,732,325,349]" country="Madagascar" name="Boby" pageId="15" pageNumber="570">Boby</location>
,
<quantity id="4CEB9B61FFE8973B70BBFECDFCBFFF7E" box="[743,827,325,349]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="570" unit="m" value="2600.0">
<elevation id="003ED1B7FFE8973B70BBFECDFCBFFF7E" box="[743,827,325,349]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="15" pageNumber="570" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<date id="FFAD1044FFE8973B711BFECDFC39FF7E" box="[839,957,325,349]" pageId="15" pageNumber="570" value="1972-04-27">
<collectingDate id="EFE9E9ACFFE8973B711BFECDFC39FF7E" box="[839,957,325,349]" pageId="15" pageNumber="570" value="1972-04-27">27.IV.1972</collectingDate>
</date>
,
<collectionCode id="ED02AE41FFE8973B7195FECDFB89FF7E" box="[969,1037,325,349]" country="France" httpUri="http://grbio.org/cool/peix-xwi9" name="Centre National de la Recherche Scientifique" pageId="15" pageNumber="570">CNRS</collectionCode>
<specimenCode id="DBB59EFFFFE8973B764AFECDFBFCFF7E" box="[1046,1144,325,349]" collectionCode="RCP" pageId="15" pageNumber="570">RCP 225</specimenCode>
,
<collectorName id="26E65352FFE8973B76D8FECDFD6DFF54" pageId="15" pageNumber="570">J.-M. Betsch</collectorName>
,
<specimenCount id="9D15FD0DFFE8973B70AEFED7FC9DFF54" box="[754,793,351,375]" pageId="15" pageNumber="570" type="male">1 ♂</specimenCount>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAC3684FFE8973B70F9FE01FC32FCE7" blockId="15.[674,1211,298,708]" pageId="15" pageNumber="570">
<typeStatus id="54A88826FFE8973B70F9FE01FC81FFBC" box="[677,773,393,416]" pageId="15" pageNumber="570" type="paratype">PARATYPES</typeStatus>
. —
<emphasis id="B967EA96FFE8973B7177FE01FC60FF82" bold="true" box="[811,996,393,417]" pageId="15" pageNumber="570">
<collectingCountry id="F3047614FFE8973B7177FE01FC1EFF82" box="[811,922,393,417]" name="Madagascar" pageId="15" pageNumber="570">Madagascar</collectingCountry>
<collectingRegion id="49D7F866FFE8973B71C2FE01FC60FF82" box="[926,996,393,417]" pageId="15" pageNumber="570">Centre.</collectingRegion>
</emphasis>
Andringitra, Marositry,
<quantity id="4CEB9B61FFE8973B70FEFE2BFD76FF98" box="[674,754,419,443]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="570" unit="m" value="2000.0">2000 m</quantity>
,
<collectionCode id="ED02AE41FFE8973B70A1FE2BFCC8FF98" box="[765,844,419,443]" pageId="15" pageNumber="570">FDHM</collectionCode>
,
<date id="FFAD1044FFE8973B710BFE2BFC43FF98" box="[855,967,419,443]" pageId="15" pageNumber="570" value="1970-12-03">3.XII.1970</date>
,
<collectionCode id="ED02AE41FFE8973B718DFE2BFB97FF98" box="[977,1043,419,443]" country="France" httpUri="http://grbio.org/cool/peix-xwi9" name="Centre National de la Recherche Scientifique" pageId="15" pageNumber="570">CNRS</collectionCode>
<collectionCode id="ED02AE41FFE8973B7646FE2BFBCDFF98" box="[1050,1097,419,443]" pageId="15" pageNumber="570">RCP</collectionCode>
225, J.-M. Betsch,
<specimenCount id="9D15FD0DFFE8973B70AEFE36FC9DFFF6" box="[754,793,446,470]" pageId="15" pageNumber="570" type="male">1 ♂</specimenCount>
. — Marositry, est riv. Antsifotra,
<quantity id="4CEB9B61FFE8973B763AFE36FB31FFF5" box="[1126,1205,446,470]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="570" unit="m" value="2000.0">2000 m</quantity>
,
<collectionCode id="ED02AE41FFE8973B70F9FE50FD69FFD3" box="[677,749,472,496]" pageId="15" pageNumber="570">FDSM</collectionCode>
,
<date id="FFAD1044FFE8973B70A5FE50FCFBFFD3" box="[761,895,472,496]" pageId="15" pageNumber="570" value="1970-12-02" valueMax="1970-12-05" valueMin="1970-12-02">2-5.XII.1970</date>
, J.-M. Betsch,
<specimenCount id="9D15FD0DFFE8973B7644FE50FBCDFFD3" box="[1048,1097,472,496]" pageId="15" pageNumber="570" type="generic">1 ex.</specimenCount>
— Plateau Andohariana,
<quantity id="4CEB9B61FFE8973B7171FE7BFC30FC28" box="[813,948,499,523]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="15" pageNumber="570" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000-2100 m</quantity>
,
<collectionCode id="ED02AE41FFE8973B71E0FE7BFB85FC28" box="[956,1025,499,523]" pageId="15" pageNumber="570">BFAM</collectionCode>
,
<date id="FFAD1044FFE8973B7655FE7BFB3CFC28" box="[1033,1208,499,523]" pageId="15" pageNumber="570" value="1970-11-09" valueMax="1970-12-10" valueMin="1970-11-09">9.XI-10.XII.1970</date>
,
<collectionCode id="ED02AE41FFE8973B70FFFD85FD56FC06" box="[675,722,525,549]" pageId="15" pageNumber="570">RCP</collectionCode>
225, J.-M. Betsch,
<specimenCount id="9D15FD0DFFE8973B71CEFD85FC45FC06" box="[914,961,525,549]" pageId="15" pageNumber="570" type="generic">1 ex.</specimenCount>
— Andrianony,
<quantity id="4CEB9B61FFE8973B7639FD85FB37FC06" box="[1125,1203,525,549]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="15" pageNumber="570" unit="m" value="1650.0">1650 m</quantity>
,
<date id="FFAD1044FFE8973B70F8FDA0FC8FFC63" box="[676,779,552,576]" pageId="15" pageNumber="570" value="1970-11-03">3.XI.1970</date>
, J.-M. Betsch,
<specimenCount id="9D15FD0DFFE8973B71CBFDA0FC40FC63" box="[919,964,552,576]" pageId="15" pageNumber="570" type="generic">1 ex.</specimenCount>
— Anjavidilava,
<quantity id="4CEB9B61FFE8973B7634FDA0FB31FC63" box="[1128,1205,552,576]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="570" unit="m" value="2000.0">2000 m</quantity>
,
<collectionCode id="ED02AE41FFE8973B70FFFDCAFD75FC79" box="[675,753,578,602]" pageId="15" pageNumber="570">FDSMP</collectionCode>
,
<date id="FFAD1044FFE8973B70A5FDCAFCF4FC79" box="[761,880,578,602]" pageId="15" pageNumber="570" value="1970-12-20">20.XII.1970</date>
, J.-M. Betsch,
<specimenCount id="9D15FD0DFFE8973B71AAFDCAFB9EFC7A" box="[1014,1050,578,602]" pageId="15" pageNumber="570" type="female">1 ♀</specimenCount>
. — Anjavidilava,
<quantity id="4CEB9B61FFE8973B70FFFDD5FCAEFC56" box="[675,810,605,629]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="1.95" metricValueMin="1.85" pageId="15" pageNumber="570" unit="m" value="1900.0" valueMax="1950.0" valueMin="1850.0">1850-1950 m</quantity>
,
<collectionCode id="ED02AE41FFE8973B716FFDD5FC06FC56" box="[819,898,605,629]" pageId="15" pageNumber="570">FDHM</collectionCode>
2,
<date id="FFAD1044FFE8973B71CBFDD5FB92FC56" box="[919,1046,605,629]" pageId="15" pageNumber="570" value="1970-12-18">18.XII.1970</date>
-
<date id="FFAD1044FFE8973B7641FDD5FBFEFC56" box="[1053,1146,605,629]" pageId="15" pageNumber="570" value="1971-01-15">15.I.1971</date>
, J.-M. Betsch,
<specimenCount id="9D15FD0DFFE8973B70B3FDFFFC99FCAC" box="[751,797,631,655]" pageId="15" pageNumber="570" type="generic">1 ex.</specimenCount>
— Cirque Boby, zone sommitale,
<collectionCode id="ED02AE41FFE8973B7635FDFFFB30FCAC" box="[1129,1204,631,655]" pageId="15" pageNumber="570">HFAM</collectionCode>
,
<quantity id="4CEB9B61FFE8973B70F8FD1AFD7DFC89" box="[676,761,658,682]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="570" unit="m" value="2500.0">2500 m</quantity>
,
<date id="FFAD1044FFE8973B715AFD1AFC20FC89" box="[774,932,658,682]" pageId="15" pageNumber="570" value="1970-11-23" valueMax="1970-11-29" valueMin="1970-11-23">23-29.XI.1970</date>
,
<specimenCount id="9D15FD0DFFE8973B71EDFD1AFC60FC89" box="[945,996,658,682]" pageId="15" pageNumber="570" type="generic">5 ex.</specimenCount>
— Pic Boby, FBM,
<quantity id="4CEB9B61FFE8973B70FBFD24FD72FCE7" box="[679,758,684,708]" metricMagnitude="3" metricUnit="m" metricValue="2.55" pageId="15" pageNumber="570" unit="m" value="2550.0">2550 m</quantity>
,
<date id="FFAD1044FFE8973B715CFD24FCF8FCE7" box="[768,892,684,708]" pageId="15" pageNumber="570" value="1970-12-16">16.XII.1970</date>
,
<specimenCount id="9D15FD0DFFE8973B71DBFD24FC32FCE7" box="[903,950,684,708]" pageId="15" pageNumber="570" type="generic">1 ex.</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C309650FFFE8973B70F9FD54FC6FFBAA" pageId="15" pageNumber="570" type="description">
<paragraph id="8BAC3684FFE8973B70F9FD54FBA2FDE4" blockId="15.[675,1210,732,967]" pageId="15" pageNumber="570">
DIAGNOSE. — Cette nouvelle espèce est la première connue du sous-genre
<emphasis id="B967EA96FFE8973B71DDFD7EFC48FD2D" box="[897,972,758,782]" italics="true" pageId="15" pageNumber="570">Calpusa</emphasis>
Mulsant &amp; Rey, 1871 à
<collectingCountry id="F3047614FFE8973B70F9FC99FC9EFD0A" box="[677,794,785,809]" name="Madagascar" pageId="15" pageNumber="570">Madagascar</collectingCountry>
. Lespèce géographiquement la plus voisine est
<taxonomicName id="4C134D07FFE8973B709FFCA4FC2FFD60" authority="Pace, 1994" authorityName="Pace" authorityYear="1994" box="[707,939,811,835]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="570" phylum="Arthropoda" rank="species" species="ruandensis">
<emphasis id="B967EA96FFE8973B709FFCA4FCBFFD60" box="[707,827,811,835]" italics="true" pageId="15" pageNumber="570">P. ruandensis</emphasis>
<bibRefCitation id="EF824B75FFE8973B711CFCA3FC2FFD60" author="PACE R." box="[832,939,811,835]" pageId="15" pageNumber="570" pagination="117 - 193" refId="ref9706" refString="PACE R. 1994. - Aleocharinae della Sottoregione Africana Orientale al Museo di Ginevra (Coleoptera, Staphylinidae). Parte I. Revue suisse de Zoologie 100 (4): 117 - 193." type="journal article" year="1994">Pace, 1994</bibRefCitation>
</taxonomicName>
, du
<collectingCountry id="F3047614FFE8973B718FFCA3FBA2FD60" box="[979,1062,811,835]" name="Rwanda" pageId="15" pageNumber="570">Rwanda</collectingCountry>
. Cette nouvelle espèce sen distingue par les yeux plus développés (yeux plus courts que les tempes chez
<taxonomicName id="4C134D07FFE8973B719DFCE9FBB7FD5B" authorityName="Pace" authorityYear="1994" box="[961,1075,864,888]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="570" phylum="Arthropoda" rank="species" species="ruandensis">
<emphasis id="B967EA96FFE8973B719DFCE9FBB7FD5B" box="[961,1075,864,888]" italics="true" pageId="15" pageNumber="570">P. ruandensis</emphasis>
</taxonomicName>
), par labsence dune fossette sur la tête (présente chez
<taxonomicName id="4C134D07FFE8973B767DFCF3FB1EFDB1" authorityName="Pace" authorityYear="1994" box="[1057,1178,890,914]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="570" phylum="Arthropoda" rank="species" species="ruandensis">
<emphasis id="B967EA96FFE8973B767DFCF3FB1EFDB1" box="[1057,1178,890,914]" italics="true" pageId="15" pageNumber="570">P. ruandensis</emphasis>
</taxonomicName>
) et par la partie apicale de lédéage beaucoup plus large, en vue ventrale (étroite chez
<taxonomicName id="4C134D07FFE8973B71C2FC38FB93FDE4" authorityName="Pace" authorityYear="1994" box="[926,1047,943,967]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="570" phylum="Arthropoda" rank="species" species="ruandensis">
<emphasis id="B967EA96FFE8973B71C2FC38FB93FDE4" box="[926,1047,943,967]" italics="true" pageId="15" pageNumber="570">P. ruandensis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BAC3684FFE8973B70F9FC6CFCB1FDDF" blockId="15.[674,1212,996,1308]" box="[677,821,996,1022]" pageId="15" pageNumber="570">DESCRIPTION</paragraph>
<paragraph id="8BAC3684FFE8973B70F9FB8BFB37FB3F" blockId="15.[674,1212,996,1308]" pageId="15" pageNumber="570">
Longueur
<quantity id="4CEB9B61FFE8973B7150FB8BFCDBFA3D" box="[780,863,1027,1054]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="570" unit="mm" value="1.3">1,3 mm</quantity>
.Corps brillant et noir-brun, élytres bruns; antennes noires avec les deux antennomères basaux bruns; pattes brunes avec la base et lextrémité des tibias ainsi que les tarses jaunes. Ponctuation de la tête et du pronotum indistincte. Granulation des élytres et de labdomen superficielle. Réticulation du corps superficielle, seulement évidente sur les tempes, celle de labdomen à mailles très transverses. Édéage:
<figureCitation id="13282A01FFE8973B715CFA89FC0AFB3F" box="[768,910,1281,1308]" captionStart="FIG" captionStartId="14.[113,124,1659,1676]" captionTargetBox="[124,1195,221,1586]" captionTargetId="figure-16@14.[98,1224,179,1634]" captionTargetPageId="14" captionText="FIG. 7. — Habitus (A, C, G), spermathèque (B, F, J) et édéage en vues latérale (D, H) et ventrale (E, I): A, B, Brachida hova n. sp.; C-F, Placusa (Calpusa) malgascia n. sp.; G-J, Placusa (Calpusa) madecassa n. sp. Échelles: A, C, G, 1 mm; B, D-F, H-J, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.5395881" httpUri="https://zenodo.org/record/5395881/files/figure.png" pageId="15" pageNumber="570">Figure 7D, E</figureCitation>
; spermathèque:
<figureCitation id="13282A01FFE8973B761BFA89FB2BFB3F" box="[1095,1199,1281,1308]" captionStart="FIG" captionStartId="14.[113,124,1659,1676]" captionTargetBox="[124,1195,221,1586]" captionTargetId="figure-16@14.[98,1224,179,1634]" captionTargetPageId="14" captionText="FIG. 7. — Habitus (A, C, G), spermathèque (B, F, J) et édéage en vues latérale (D, H) et ventrale (E, I): A, B, Brachida hova n. sp.; C-F, Placusa (Calpusa) malgascia n. sp.; G-J, Placusa (Calpusa) madecassa n. sp. Échelles: A, C, G, 1 mm; B, D-F, H-J, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.5395881" httpUri="https://zenodo.org/record/5395881/files/figure.png" pageId="15" pageNumber="570">
Figure
<specimenCount id="9D15FD0DFFE8973B76CFFA89FB2BFB3F" box="[1171,1199,1281,1308]" pageId="15" pageNumber="570" type="generic">7F</specimenCount>
</figureCitation>
.
</paragraph>
<paragraph id="8BAC3684FFE8973B70ADFAC5FC6FFBAA" blockId="15.[753,1133,1357,1417]" pageId="15" pageNumber="570">
<taxonomicName id="4C134D07FFE8973B70ADFAC5FBAFFB4B" authority="Pace, 2008" authorityName="Pace" authorityYear="2008" box="[753,1067,1357,1385]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="570" phylum="Arthropoda" rank="species" species="madecassa" status="sp. nov." subGenus="Calpusa">
<emphasis id="B967EA96FFE8973B70ADFAC5FCC0FB4B" bold="true" box="[753,836,1357,1384]" italics="true" pageId="15" pageNumber="570">Placusa</emphasis>
(
<emphasis id="B967EA96FFE8973B7108FAC5FC28FB4B" bold="true" box="[852,940,1357,1384]" italics="true" pageId="15" pageNumber="570">Calpusa</emphasis>
)
<emphasis id="B967EA96FFE8973B71E0FAC5FBAFFB4B" bold="true" box="[956,1067,1357,1384]" italics="true" pageId="15" pageNumber="570">madecassa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457EDFFE8973B766EFAC6FBE9FB4A" box="[1074,1133,1358,1385]" pageId="15" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13282A01FFE8973B7127FAE6FC3DFBAA" box="[891,953,1390,1417]" captionStart="FIG" captionStartId="14.[113,124,1659,1676]" captionTargetBox="[124,1195,221,1586]" captionTargetId="figure-16@14.[98,1224,179,1634]" captionTargetPageId="14" captionText="FIG. 7. — Habitus (A, C, G), spermathèque (B, F, J) et édéage en vues latérale (D, H) et ventrale (E, I): A, B, Brachida hova n. sp.; C-F, Placusa (Calpusa) malgascia n. sp.; G-J, Placusa (Calpusa) madecassa n. sp. Échelles: A, C, G, 1 mm; B, D-F, H-J, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.5395881" httpUri="https://zenodo.org/record/5395881/files/figure.png" pageId="15" pageNumber="570">Fig. 7</figureCitation>
G-J)
</paragraph>
</subSubSection>
<subSubSection id="C309650FFFE8973B70F9FA2FFCDBF87A" pageId="15" pageNumber="570" type="materials_examined">
<paragraph id="8BAC3684FFE8973B70F9FA2FFCA6FBFA" blockId="15.[675,1210,1447,1497]" pageId="15" pageNumber="570">
<materialsCitation id="3B7B3CD9FFE8973B70F9FA2FFCA6FBFA" collectorName="Dr Ch. Nodier" country="Madagascar" location="Madagascar" pageId="15" pageNumber="570" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A88826FFE8973B70F9FA2FFC8AFB9E" box="[677,782,1447,1470]" pageId="15" pageNumber="570" type="holotype">HOLOTYPE</typeStatus>
. —
<emphasis id="B967EA96FFE8973B716AFA2FFC7FFB9C" bold="true" box="[822,1019,1447,1471]" pageId="15" pageNumber="570">
<collectingCountry id="F3047614FFE8973B716AFA2FFC28FB9C" box="[822,940,1447,1471]" name="Madagascar" pageId="15" pageNumber="570">Madagascar</collectingCountry>
Centre.
</emphasis>
Tananarive,
<collectorName id="26E65352FFE8973B7628FA2FFD6DFBFA" pageId="15" pageNumber="570">Dr Ch. Nodier</collectorName>
,
<specimenCount id="9D15FD0DFFE8973B70AFFA49FC98FBFA" box="[755,796,1473,1497]" pageId="15" pageNumber="570" type="male">1 ♂</specimenCount>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAC3684FFE8973B70F9FA7AFCDBF87A" blockId="15.[675,1210,1521,1625]" pageId="15" pageNumber="570">
<typeStatus id="54A88826FFE8973B70F9FA7AFC82F82B" box="[677,774,1522,1545]" pageId="15" pageNumber="570" type="paratype">PARATYPES</typeStatus>
. — Mêmes données que pour lholotype,
<specimenCount id="9D15FD0DFFE8973B76DEFA79FB37F82B" box="[1154,1203,1521,1545]" pageId="15" pageNumber="570" type="female">6 ♀♀</specimenCount>
. Lun des
<typeStatus id="54A88826FFE8973B70AAF984FCD4F807" box="[758,848,1548,1572]" pageId="15" pageNumber="570" type="paratype">paratypes</typeStatus>
femelles est porteur de Laboulbéniales (champignons parasites), qui ont été montées dans du baume du
<collectingCountry id="F3047614FFE8973B7151F9C9FCDFF87A" box="[781,859,1601,1625]" name="Canada" pageId="15" pageNumber="570">Canada</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C309650FFFE8972570F9F9F9FF58FA28" lastPageId="17" lastPageNumber="572" pageId="15" pageNumber="570" type="description">
<paragraph id="8BAC3684FFE8972470F9F9F9FCEBF881" blockId="15.[677,1211,1649,1699]" lastBlockId="16.[675,1209,1595,1698]" lastPageId="16" lastPageNumber="571" pageId="15" pageNumber="570">
DIAGNOSE. — Lédéage de la nouvelle espèce présente une lame ventrale insérée près de la « crista apicalis ». Une lame semblable se retrouve sur lédéage de
<taxonomicName id="4C134D07FFF79724706EF9B4FF64F84E" authority="Pace, 1994" authorityName="Pace" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="571" phylum="Arthropoda" rank="species" species="somala">
<emphasis id="B967EA96FFF79724706EF9B4FD02F870" box="[562,646,1595,1619]" italics="true" pageId="16" pageNumber="571">P. somala</emphasis>
<bibRefCitation id="EF824B75FFF79724722CF9DDFF64F84E" author="PACE R." box="[112,224,1621,1645]" pageId="16" pageNumber="571" pagination="117 - 193" refId="ref9706" refString="PACE R. 1994. - Aleocharinae della Sottoregione Africana Orientale al Museo di Ginevra (Coleoptera, Staphylinidae). Parte I. Revue suisse de Zoologie 100 (4): 117 - 193." type="journal article" year="1994">Pace, 1994</bibRefCitation>
</taxonomicName>
, de
<collectingCountry id="F3047614FFF797247350F9DDFEDAF84E" box="[268,350,1621,1645]" name="Somalia" pageId="16" pageNumber="571">Somalie</collectingCountry>
. Laffinité taxonomique entre ces deux espèces, cependant, reste faible. En effet, la pubescence du corps de
<taxonomicName id="4C134D07FFF797247308F903FE23F881" authorityName="Pace" authorityYear="1994" box="[340,423,1674,1698]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="571" phylum="Arthropoda" rank="species" species="somala">
<emphasis id="B967EA96FFF797247308F903FE23F881" box="[340,423,1674,1698]" italics="true" pageId="16" pageNumber="571">P. somala</emphasis>
</taxonomicName>
est dense. Lédéage et la spermathèque de la nouvelle espèce sont de toute évidence de forme différente: le bulbe distal de la spermathèque de
<taxonomicName id="4C134D07FFF79724709FF9F8FC9EF8AB" authorityName="Pace" authorityYear="1994" box="[707,794,1648,1672]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="571" phylum="Arthropoda" rank="species" species="somala">
<emphasis id="B967EA96FFF79724709FF9F8FC9EF8AB" box="[707,794,1648,1672]" italics="true" pageId="16" pageNumber="571">P. somala</emphasis>
</taxonomicName>
est long et réniforme, celui de la nouvelle espèce est sphérique.
</paragraph>
<caption id="DF6C660CFFF79724722DFA43FF52F829" ID-DOI="http://doi.org/10.5281/zenodo.5395883" ID-Zenodo-Dep="5395883" httpUri="https://zenodo.org/record/5395883/files/figure.png" pageId="16" pageNumber="571" startId="16.[113,124,1483,1500]" targetBox="[124,1194,223,1432]" targetPageId="16">
<paragraph id="8BAC3684FFF79724722DFA43FF52F829" blockId="16.[113,1209,1483,1546]" pageId="16" pageNumber="571">
FIG. 8. — Spermathèque (
<emphasis id="B967EA96FFF797247314FA43FED1FBFF" bold="true" box="[328,341,1483,1500]" pageId="16" pageNumber="571">A</emphasis>
,
<emphasis id="B967EA96FFF79724733CFA43FEEAFBFF" bold="true" box="[352,366,1483,1500]" pageId="16" pageNumber="571">C</emphasis>
,
<emphasis id="B967EA96FFF797247325FA43FE03FBFF" bold="true" box="[377,391,1483,1500]" pageId="16" pageNumber="571">G</emphasis>
), habitus (
<emphasis id="B967EA96FFF797247382FA43FE6FFBFF" bold="true" box="[478,491,1483,1500]" pageId="16" pageNumber="571">B</emphasis>
,
<emphasis id="B967EA96FFF7972473AAFA43FD80FBFF" bold="true" box="[502,516,1483,1500]" pageId="16" pageNumber="571">D</emphasis>
), édéage en vues latérale (
<emphasis id="B967EA96FFF7972470B9FA43FD75FBFF" bold="true" box="[741,753,1483,1500]" pageId="16" pageNumber="571">E</emphasis>
) et ventrale (
<emphasis id="B967EA96FFF797247102FA43FCEDFBFF" bold="true" box="[862,873,1483,1500]" pageId="16" pageNumber="571">F</emphasis>
), et sixième urotergite libre du mâle (
<emphasis id="B967EA96FFF7972476C3FA43FB29FBFF" bold="true" box="[1183,1197,1483,1500]" pageId="16" pageNumber="571">H</emphasis>
):
<emphasis id="B967EA96FFF79724722DFA6AFFFAFBD0" bold="true" box="[113,126,1506,1523]" pageId="16" pageNumber="571">A</emphasis>
,
<taxonomicName id="4C134D07FFF7972472D5FA6AFE22FBD0" authority="Fauvel" authorityName="Fauvel" authorityYear="1904" box="[137,422,1506,1523]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="571" phylum="Arthropoda" rank="species" species="simulans" subGenus="Placusa">
<emphasis id="B967EA96FFF7972472D5FA6AFF4EFBD0" box="[137,202,1506,1523]" italics="true" pageId="16" pageNumber="571">Placusa</emphasis>
(
<emphasis id="B967EA96FFF797247289FA6AFE92FBD0" box="[213,278,1506,1523]" italics="true" pageId="16" pageNumber="571">Placusa</emphasis>
)
<emphasis id="B967EA96FFF79724737DFA6AFEEEFBD0" box="[289,362,1506,1523]" italics="true" pageId="16" pageNumber="571">simulans</emphasis>
Fauvel
</taxonomicName>
;
<emphasis id="B967EA96FFF7972473EFFA6AFE44FBD0" bold="true" box="[435,448,1506,1523]" pageId="16" pageNumber="571">B</emphasis>
,
<emphasis id="B967EA96FFF797247397FA6AFE5DFBD0" bold="true" box="[459,473,1506,1523]" pageId="16" pageNumber="571">C</emphasis>
,
<taxonomicName id="4C134D07FFF7972473B8FA6AFDDFFBD0" authority="Pace, 2008" authorityName="Pace" authorityYear="2008" box="[484,603,1506,1523]" class="Insecta" family="Staphylinidae" genus="Placusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="571" phylum="Arthropoda" rank="species" species="convexa" status="sp. nov." subGenus="Placusa">
<emphasis id="B967EA96FFF7972473B8FA6AFE70FBD0" box="[484,500,1506,1523]" italics="true" pageId="16" pageNumber="571">P.</emphasis>
(
<emphasis id="B967EA96FFF7972473A1FA6AFD89FBD0" box="[509,525,1506,1523]" italics="true" pageId="16" pageNumber="571">P.</emphasis>
)
<emphasis id="B967EA96FFF79724704AFA6AFDDFFBD0" box="[534,603,1506,1523]" italics="true" pageId="16" pageNumber="571">convexa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457EDFFF79724703CFA6AFD15FBD0" box="[608,657,1506,1523]" pageId="16" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B967EA96FFF7972470C1FA6AFD44FBD0" bold="true" box="[669,704,1506,1523]" pageId="16" pageNumber="571">D -H</emphasis>
,
<taxonomicName id="4C134D07FFF797247097FA6AFC0EFBD0" authority="Pace, 2008" authorityName="Pace" authorityYear="2008" box="[715,906,1506,1523]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="571" phylum="Arthropoda" rank="species" species="antongilicola" status="sp. nov.">
<emphasis id="B967EA96FFF797247097FA6AFC0EFBD0" box="[715,906,1506,1523]" italics="true" pageId="16" pageNumber="571">Homalota antongilicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457EDFFF7972471D3FA6AFC44FBD0" box="[911,960,1506,1523]" pageId="16" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
Échelles: A, C, E-H, 0,1 mm; B, D, 1 mm.
</paragraph>
</caption>
<paragraph id="8BAC3684FFF69725722DFF50FE86FED3" blockId="17.[111,648,216,1035]" box="[113,258,216,242]" pageId="17" pageNumber="572">DESCRIPTION</paragraph>
<paragraph id="8BAC3684FFF69725722DFF70FF58FA28" blockId="17.[111,648,216,1035]" pageId="17" pageNumber="572">
Longueur
<quantity id="4CEB9B61FFF6972572BAFF70FEC7FF30" box="[230,323,248,275]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="17" pageNumber="572" unit="mm" value="2.3">2,3 mm</quantity>
. Corps brillant et noir-brun, avec des élytres bruns; antennes brunes avec les trois antennomères basaux jaune-rougeâtre et les antennomères quatre et cinq rougeâtres; pattes brunes avec des tarses jaune-rougeâtre. De la base de lantenne gauche à celle de droite sétend une ligne qui sépare la partie antérieure brillante de la postérieure fortement réticulée, à mailles très transverses seulement sur le tiers postérieur. Réticulation du pronotum, des élytres et de labdomen évidente, vigoureuse sur le cinquième urotergite libre du mâle. Ponctuation de la tête ombiliquée et absente sur la bande médiane longitudinale, celle du pronotum plus évidente que celle de la tête. Granulation des élytres et de labdomen bien visible. Sutures des élytres du mâle saillantes. Les premier et deuxième urotergites libres du mâle présentent un bourrelet médian transverse postérieur. Cinquième urotergite libre du mâle concave, sixième prolongé en forme de pelle, à lextrémité arrondie et rebordée, encadrée de chaque côté dune dent aiguë. Quatrième, cinquième et sixième antennomères aussi longs que larges, septième faiblement transverse et les trois suivants transverses. Édéage:
<figureCitation id="13282A01FFF697257312FC58FE50FDC8" box="[334,468,976,1003]" captionStart="FIG" captionStartId="14.[113,124,1659,1676]" captionTargetBox="[124,1195,221,1586]" captionTargetId="figure-16@14.[98,1224,179,1634]" captionTargetPageId="14" captionText="FIG. 7. — Habitus (A, C, G), spermathèque (B, F, J) et édéage en vues latérale (D, H) et ventrale (E, I): A, B, Brachida hova n. sp.; C-F, Placusa (Calpusa) malgascia n. sp.; G-J, Placusa (Calpusa) madecassa n. sp. Échelles: A, C, G, 1 mm; B, D-F, H-J, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.5395881" httpUri="https://zenodo.org/record/5395881/files/figure.png" pageId="17" pageNumber="572">Figure 7H, I</figureCitation>
; spermathèque:
<figureCitation id="13282A01FFF69725722DFC78FF53FA28" box="[113,215,1008,1035]" captionStart="FIG" captionStartId="14.[113,124,1659,1676]" captionTargetBox="[124,1195,221,1586]" captionTargetId="figure-16@14.[98,1224,179,1634]" captionTargetPageId="14" captionText="FIG. 7. — Habitus (A, C, G), spermathèque (B, F, J) et édéage en vues latérale (D, H) et ventrale (E, I): A, B, Brachida hova n. sp.; C-F, Placusa (Calpusa) malgascia n. sp.; G-J, Placusa (Calpusa) madecassa n. sp. Échelles: A, C, G, 1 mm; B, D-F, H-J, 0,1 mm." figureDoi="http://doi.org/10.5281/zenodo.5395881" httpUri="https://zenodo.org/record/5395881/files/figure.png" pageId="17" pageNumber="572">Figure 7J</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>