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<document id="34F054BCECEB4DD29F373626F69CFA5A" ID-DOI="10.5281/zenodo.5390317" ID-ISSN="1638-9387" ID-Zenodo-Dep="5390317" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630312953056" checkinUser="marcus" docAuthor="Rahayu, Dwi Listyo &amp; Ng, Peter K. L." docDate="2004" docId="03BA8787FFA9FF99FE89FB1CFD282988" docLanguage="en" docName="Zoosystema.2004.26.1.87-94.pdf" docOrigin="Zoosystema 26 (1)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Amarinus pristes Rahayu &amp; Ng 2004, n. sp." docType="treatment" docVersion="4" lastPageNumber="93" masterDocId="FF83FFFFFFADFF9FFFAFFFB5FFA12F0D" masterDocTitle="The Hymenosomatidae (Crustacea, Decapoda, Brachyura) of Timika (Irian Jaya, Indonesia)" masterLastPageNumber="94" masterPageNumber="87" pageNumber="91" updateTime="1699222838032" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="0D12A02068C4FC6AA675D201DED10773" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="16991797AC9AD703FBB65939A56353F4">
<mods:title id="14A1936BE93B02ECB9830E9727A803AC">The Hymenosomatidae (Crustacea, Decapoda, Brachyura) of Timika (Irian Jaya, Indonesia)</mods:title>
</mods:titleInfo>
<mods:name id="7AB6B947F4F7084BCBDDD4541ED76FD5" type="personal">
<mods:role id="1D89DCAE33E93EF67F7B7A25BC6A55C2">
<mods:roleTerm id="F5812A236ED4A94EC25758C72DF3C54F">Author</mods:roleTerm>
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<mods:namePart id="9460DEABFD35A325506B25DAAFDB532B">Rahayu, Dwi Listyo</mods:namePart>
<mods:affiliation id="1C7711A8B6ADC8EDC8B1B32399AAB567">Research Centre for Oceanography, Indonesian Institute of Sciences, Jalan Pasir Putih 1, Ancol Timur, Jakarta 11048 (Indonesia) dwilistyo @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="1D28A335D70901AA4FC1DC7FB9C79AE4" type="email">dwilistyo@yahoo.com</mods:nameIdentifier>
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<mods:role id="C454B390865B48A295DC7CDD00570178">
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<mods:namePart id="BCE0FA48FDDBCD9823F8F1DF25E0DA75">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="D96B0CE7A80B5F22F2D55CA4F4CD8C95">Department of Biological Sciences, National University of Singapore, Kent Ridge, Singapore 119260 (Republic of Singapore) peterng @ nus. edu. sg ..</mods:affiliation>
<mods:nameIdentifier id="18821794B2E4B929DF64C181A01EC03E" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<mods:title id="6911870C952EB4BB267F10BA0065E9AB">Zoosystema</mods:title>
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<mods:part id="E188450B7F77A55A00518786CCAE22C5">
<mods:date id="45B5D4AA0984D40AB0DC1C882C87083B">2004</mods:date>
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<mods:number id="EBC3CA213ACBC0D6DFAAE899BA3E7DD4">26</mods:number>
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<mods:identifier id="2062530E0F47B2BDB01FE96DB3CFF9AD" type="DOI">10.5281/zenodo.5390317</mods:identifier>
<mods:identifier id="6E858B851373BE997EE953AC233B0385" type="ISSN">1638-9387</mods:identifier>
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<treatment id="03BA8787FFA9FF99FE89FB1CFD282988" ID-DOI="http://doi.org/10.5281/zenodo.5467565" ID-Zenodo-Dep="5467565" LSID="urn:lsid:plazi:treatment:03BA8787FFA9FF99FE89FB1CFD282988" httpUri="http://treatment.plazi.org/id/03BA8787FFA9FF99FE89FB1CFD282988" lastPageId="6" lastPageNumber="93" pageId="4" pageNumber="91">
<subSubSection id="C309651AFFA9FF9BFE89FB1CFDBC2BC9" box="[294,541,1193,1220]" pageId="4" pageNumber="91" type="nomenclature">
<paragraph id="8BAC3691FFA9FF9BFE89FB1CFDBC2BC9" blockId="4.[294,541,1193,1253]" box="[294,541,1193,1220]" pageId="4" pageNumber="91">
<heading id="D0E481FDFFA9FF9BFE89FB1CFDBC2BC9" box="[294,541,1193,1220]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="91" reason="2">
<taxonomicName id="4C134D12FFA9FF9BFE89FB1CFE7A2BCE" ID-CoL="65YHV" authority="Rahayu &amp; Ng, 2004" authorityName="Rahayu &amp; Ng" authorityYear="2004" box="[294,475,1193,1219]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="pristes" status="sp. nov.">
<emphasis id="B967EA83FFA9FF9BFE89FB1CFE7A2BCE" bold="true" box="[294,475,1193,1219]" italics="true" pageId="4" pageNumber="91">Amarinus pristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F8FFA9FF9BFE4DFB1FFDBC2BC9" box="[482,541,1194,1220]" pageId="4" pageNumber="91" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C309651AFFA9FF9BFED6FB7EFE682BE8" box="[377,457,1227,1253]" pageId="4" pageNumber="91" type="description">
<paragraph id="8BAC3691FFA9FF9BFED6FB7EFE682BE8" blockId="4.[294,541,1193,1253]" box="[377,457,1227,1253]" pageId="4" pageNumber="91">
(
<figureCitation id="13282A14FFA9FF9BFE2EFB7EFE632BE8" box="[385,450,1227,1253]" captionStart="FIG" captionStartId="5.[151,162,1629,1646]" captionTargetBox="[221,1188,259,1558]" captionTargetId="figure-27@5.[181,1252,244,1614]" captionTargetPageId="5" captionText="FIG. 2. — Amarinus pristes n. sp.; A-F, holotype 4.2 × 4.3 mm (MZB); G, H, paratype 5.5 × 5.7 mm (ZRC 2002.596); A, G, carapace; B, left third maxilliped; C, right cheliped; D, left third pereiopod; E, abdomen; F, right G1; G, abdomen. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="4" pageNumber="91">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C309651AFFA9FF9BFF38FAA7FE312988" pageId="4" pageNumber="91" type="materials_examined">
<paragraph id="8BAC3691FFA9FF9BFF38FAA7FDA32A53" blockId="4.[151,685,1298,1669]" pageId="4" pageNumber="91">
<materialsCitation id="3B7B3CCCFFA9FF9BFF38FAA7FDA32A53" collectingDate="2001-09-21" collectionCode="MZB" collectorName="I. Ermayanti" country="Indonesia" location="Kamora" pageId="4" pageNumber="91" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A88833FFA9FF9BFF38FAA7FEA42A25" box="[151,261,1298,1321]" pageId="4" pageNumber="91" type="holotype">HOLOTYPE</typeStatus>
. —
<emphasis id="B967EA83FFA9FF9BFE98FAA6FE3F2A26" bold="true" box="[311,414,1299,1323]" pageId="4" pageNumber="91">
<collectingCountry id="F3047601FFA9FF9BFE98FAA6FE3F2A26" box="[311,414,1299,1323]" name="Indonesia" pageId="4" pageNumber="91">Indonesia</collectingCountry>
</emphasis>
.
<location id="8ECC604AFFA9FF9BFE02FAA7FDA02A24" LSID="urn:lsid:plazi:treatment:03BA8787FFA9FF99FE89FB1CFD282988:8ECC604AFFA9FF9BFE02FAA7FDA02A24" box="[429,513,1298,1321]" country="Indonesia" name="Kamora" pageId="4" pageNumber="91">Kamora</location>
, intertidal, mud substrate in mangrove area,
<date id="FFAD1051FFA9FF9BFE7CFA98FDF52A48" box="[467,596,1325,1349]" pageId="4" pageNumber="91" value="2001-09-21">
<collectingDate id="EFE9E9B9FFA9FF9BFE7CFA98FDF52A48" box="[467,596,1325,1349]" pageId="4" pageNumber="91" value="2001-09-21">21.IX.2001</collectingDate>
</date>
, coll.
<collectorName id="26E65347FFA9FF9BFD34FA99FEA12A53" pageId="4" pageNumber="91">I. Ermayanti</collectorName>
, 4.2 ×
<quantity id="4CEB9B74FFA9FF9BFEF0FAF3FE0C2A51" box="[351,429,1349,1374]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="91" unit="mm" value="4.3">4.3 mm</quantity>
(
<collectionCode id="ED02AE54FFA9FF9BFE14FAF3FE572A53" box="[443,502,1350,1374]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="91" type="Museum">MZB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAC3691FFA9FF9BFF38FADAFD302AAC" blockId="4.[151,685,1298,1669]" pageId="4" pageNumber="91">
<typeStatus id="54A88833FFA9FF9BFF38FADAFF5E2A88" box="[151,255,1391,1414]" pageId="4" pageNumber="91" type="paratype">PARATYPES</typeStatus>
. —
<emphasis id="B967EA83FFA9FF9BFE82FAC5FE2E2A85" bold="true" box="[301,399,1392,1416]" pageId="4" pageNumber="91">
<collectingCountry id="F3047601FFA9FF9BFE82FAC5FE2E2A85" box="[301,399,1392,1416]" name="Indonesia" pageId="4" pageNumber="91">Indonesia</collectingCountry>
</emphasis>
. Otakwa, estuary,
<date id="FFAD1051FFA9FF9BFDE7FAC5FD092A85" box="[584,680,1392,1416]" pageId="4" pageNumber="91" value="1999-02-05">5.II.1999</date>
, coll. A. Haris, 1 5.5 ×
<quantity id="4CEB9B74FFA9FF9BFE21FA3CFE7D2A92" box="[398,476,1416,1441]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="91" unit="mm" value="5.7">5.7 mm</quantity>
(
<collectionCode id="ED02AE54FFA9FF9BFE46FA3CFDBD2AAC" box="[489,540,1417,1441]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="91">ZRC</collectionCode>
2002.596).
</paragraph>
<paragraph id="8BAC3691FFA9FF9BFF38FA17FE312988" blockId="4.[151,685,1298,1669]" pageId="4" pageNumber="91">
Kamora, intertidal, mud substrate in mangrove area,
<date id="FFAD1051FFA9FF9BFF38FA09FEAB2AD9" box="[151,266,1468,1492]" pageId="4" pageNumber="91" value="2001-09-21">21.IX.2001</date>
, coll. I. Ermayanti, 2 3.0 ×
<quantity id="4CEB9B74FFA9FF9BFDF9FA09FD092ADF" box="[598,680,1467,1492]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="91" unit="mm" value="2.9">2.9 mm</quantity>
, 4 ×
<quantity id="4CEB9B74FFA9FF9BFF6EFA60FEB02AE6" box="[193,273,1492,1517]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="4" pageNumber="91" unit="mm" value="4.1">4.1 mm</quantity>
(
<collectionCode id="ED02AE54FFA9FF9BFE8DFA60FEFD2AE0" box="[290,348,1493,1517]" pageId="4" pageNumber="91">RCO</collectionCode>
);
<date id="FFAD1051FFA9FF9BFEDFFA60FE7A2AE0" box="[368,475,1493,1517]" pageId="4" pageNumber="91" value="2001-10-17">17.X.2001</date>
, 1 4.4 ×
<quantity id="4CEB9B74FFA9FF9BFDF4FA60FD0D2AE6" box="[603,684,1492,1517]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="4" pageNumber="91" unit="mm" value="4.4">4.4 mm</quantity>
(
<collectionCode id="ED02AE54FFA9FF9BFF31FA5AFF71290A" box="[158,208,1519,1543]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="91">ZRC</collectionCode>
2002.598);
<date id="FFAD1051FFA9FF9BFE90FA5AFE3A290A" box="[319,411,1519,1543]" pageId="4" pageNumber="91" value="2001-10-05">5.X.2001</date>
,1 4.1 ×4.0mm (
<collectionCode id="ED02AE54FFA9FF9BFDE3FA5AFD03290A" box="[588,674,1519,1543]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="91">MNHN</collectionCode>
). Ajkwa, intertidal, mud substrate in mangrove area,
<date id="FFAD1051FFA9FF9BFF38F994FEA32934" box="[151,258,1569,1593]" pageId="4" pageNumber="91" value="2001-10-16">16.X.2001</date>
, coll. I. Ermayanti, 1 5.1 × 5.0 mm, 1 3.7 ×
<quantity id="4CEB9B74FFA9FF9BFF7DF98EFE80295C" box="[210,289,1594,1619]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="91" unit="mm" value="3.5">3.5 mm</quantity>
(
<collectionCode id="ED02AE54FFA9FF9BFE80F98EFEC3295E" box="[303,354,1595,1619]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="91">ZRC</collectionCode>
2002.597);
<date id="FFAD1051FFA9FF9BFE71F98EFD9C295E" box="[478,573,1595,1619]" pageId="4" pageNumber="91" value="2001-10-03">3.X.2001</date>
, 1 3.4 ×
<quantity id="4CEB9B74FFA9FF9BFF38F9E1FF472967" box="[151,230,1619,1644]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="91" unit="mm" value="3.5">3.5 mm</quantity>
(
<collectionCode id="ED02AE54FFA9FF9BFF5AF9E1FE912961" box="[245,304,1620,1644]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="91" type="Museum">MZB</collectionCode>
);
<date id="FFAD1051FFA9FF9BFEEDF9E1FE0F2961" box="[322,430,1620,1644]" pageId="4" pageNumber="91" value="2001-10-22">22.X.2001</date>
, 1 4.5 ×
<quantity id="4CEB9B74FFA9FF9BFD89F9E1FDD62967" box="[550,631,1619,1644]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="91" unit="mm" value="4.3">4.3 mm</quantity>
, 1 4.5 ×
<quantity id="4CEB9B74FFA9FF9BFF7EF9D8FEBE298E" box="[209,287,1644,1669]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="4" pageNumber="91" unit="mm" value="4.3">4.3 mm</quantity>
(
<collectionCode id="ED02AE54FFA9FF9BFE82F9D8FE242988" box="[301,389,1645,1669]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="91">MNHN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C309651AFFA9FF9BFD65FF1CFB982FF8" pageId="4" pageNumber="91" type="etymology">
<paragraph id="8BAC3691FFA9FF9BFD65FF1CFB982FF8" blockId="4.[714,1247,169,245]" pageId="4" pageNumber="91">
ETYMOLOGY. — The name refers to the lateral margin of the carapace (from the Greek
<emphasis id="B967EA83FFA9FF9BFBB5FF76FBEC2FD7" box="[1050,1101,195,218]" italics="true" pageId="4" pageNumber="91">prion</emphasis>
for saw). The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C309651AFFA9FF99FD65FE9FFEC02C35" lastPageId="6" lastPageNumber="93" pageId="4" pageNumber="91" type="description">
<paragraph id="8BAC3691FFA9FF9BFD65FE9FFCFA2E4F" blockId="4.[714,1247,298,1669]" box="[714,859,298,324]" pageId="4" pageNumber="91">DESCRIPTION</paragraph>
<paragraph id="8BAC3691FFA9FF9BFD65FEFEFCFB2E68" blockId="4.[714,1247,298,1669]" box="[714,858,331,357]" pageId="4" pageNumber="91">
<emphasis id="B967EA83FFA9FF9BFD65FEFEFCFB2E68" box="[714,858,331,357]" italics="true" pageId="4" pageNumber="91">
Male
<typeStatus id="54A88833FFA9FF9BFCA8FEFEFCFB2E68" box="[775,858,331,357]" pageId="4" pageNumber="91" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAC3691FFA9FF9BFD65FEDEFC862C28" blockId="4.[714,1247,298,1669]" pageId="4" pageNumber="91">
Carapace (
<figureCitation id="13282A14FFA9FF9BFC92FEDEFC302E88" box="[829,913,363,389]" captionStart="FIG" captionStartId="5.[151,162,1629,1646]" captionTargetBox="[221,1188,259,1558]" captionTargetId="figure-27@5.[181,1252,244,1614]" captionTargetPageId="5" captionText="FIG. 2. — Amarinus pristes n. sp.; A-F, holotype 4.2 × 4.3 mm (MZB); G, H, paratype 5.5 × 5.7 mm (ZRC 2002.596); A, G, carapace; B, left third maxilliped; C, right cheliped; D, left third pereiopod; E, abdomen; F, right G1; G, abdomen. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="4" pageNumber="91">Fig. 2A</figureCitation>
) flat, circular, as long as broad or very slightly longer or shorter than broad, surrounded by complete and distinct rim, not interrupted at base of rostrum. Dorsal surface smooth, cervical, thoracic and gastrocardiac grooves distinct; cervical and thoracic grooves approaching but not reaching antero- and posterolateral margins respectively. Lateral margins crenulated, with strong, distinct teeth on anterior margin and between coxae of third and fourth pereiopods. Rostrum unilobed, broad, surface concave, not continuous with dorsal surface of carapace. Eyestalk prominent, clearly visible dorsally.
</paragraph>
<paragraph id="8BAC3691FFA9FF9BFD65FC9EFC362B29" blockId="4.[714,1247,298,1669]" pageId="4" pageNumber="91">
Third maxillipeds (
<figureCitation id="13282A14FFA9FF9BFC05FC9EFBA22C48" box="[938,1027,811,837]" captionStart="FIG" captionStartId="5.[151,162,1629,1646]" captionTargetBox="[221,1188,259,1558]" captionTargetId="figure-27@5.[181,1252,244,1614]" captionTargetPageId="5" captionText="FIG. 2. — Amarinus pristes n. sp.; A-F, holotype 4.2 × 4.3 mm (MZB); G, H, paratype 5.5 × 5.7 mm (ZRC 2002.596); A, G, carapace; B, left third maxilliped; C, right cheliped; D, left third pereiopod; E, abdomen; F, right G1; G, abdomen. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="4" pageNumber="91">Fig. 2B</figureCitation>
) almost fill mouth field when closed. Ischium of third maxilliped shorter than merus along outer lateral edge; short setae on inner lateral margin of ischium; longer setae on inner lateral margin of merus; inner lateral margins of ischium and merus meeting when closed. Palp not longer than merus, exopod much longer than merus.
</paragraph>
<paragraph id="8BAC3691FFA9FF9BFD65FB9EFCB12A29" blockId="4.[714,1247,298,1669]" pageId="4" pageNumber="91">
Chelipeds (
<figureCitation id="13282A14FFA9FF9BFCE6FB9EFC002B48" box="[841,929,1067,1093]" captionStart="FIG" captionStartId="5.[151,162,1629,1646]" captionTargetBox="[221,1188,259,1558]" captionTargetId="figure-27@5.[181,1252,244,1614]" captionTargetPageId="5" captionText="FIG. 2. — Amarinus pristes n. sp.; A-F, holotype 4.2 × 4.3 mm (MZB); G, H, paratype 5.5 × 5.7 mm (ZRC 2002.596); A, G, carapace; B, left third maxilliped; C, right cheliped; D, left third pereiopod; E, abdomen; F, right G1; G, abdomen. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="4" pageNumber="91">Fig. 2C</figureCitation>
) equal, slightly shorter than ambulatory legs; cutting edges of fingers serrated with denticles, tip of fingers sharp, gaping proximally when closed, strong median tooth on dactylus. Dactylus slightly shorter than palm, pollex gently curved inwards. Upper and lower margin of dactylus, propodus, carpus and merus scarcely setose.
</paragraph>
<paragraph id="8BAC3691FFA9FF9BFD65FA9EFCDE2929" blockId="4.[714,1247,298,1669]" pageId="4" pageNumber="91">Ambulatory legs (pereiopods 2-5) stout, crosssection circular, dorsal and ventral edges sparsely setose. Dactylus slightly shorter than propodus, slightly curved with tip sharply hooked, recurved subterminal tooth, ventral edge more densely lined with long and short setae. Propodus slightly longer than carpus; merus about 1.75 times length of carpus.</paragraph>
<paragraph id="8BAC3691FFA9FF99FD65F99EFEEF2E69" blockId="4.[714,1247,298,1669]" lastBlockId="6.[151,684,170,357]" lastPageId="6" lastPageNumber="93" pageId="4" pageNumber="91">
Abdomen (
<figureCitation id="13282A14FFA9FF9BFCEBF99EFC372948" box="[836,918,1579,1605]" captionStart="FIG" captionStartId="5.[151,162,1629,1646]" captionTargetBox="[221,1188,259,1558]" captionTargetId="figure-27@5.[181,1252,244,1614]" captionTargetPageId="5" captionText="FIG. 2. — Amarinus pristes n. sp.; A-F, holotype 4.2 × 4.3 mm (MZB); G, H, paratype 5.5 × 5.7 mm (ZRC 2002.596); A, G, carapace; B, left third maxilliped; C, right cheliped; D, left third pereiopod; E, abdomen; F, right G1; G, abdomen. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="4" pageNumber="91">Fig. 2E</figureCitation>
) five-segmented, with pleotelson free, triangular, segment 1 shortest, widest; segment 3 longest; segments 4 and 5 similar length, tapering to pleotelson; pleotelson bluntly triangular, approximately same length as segment 3, with lateral intercalated plates basally; press-button system as described by
<bibRefCitation id="EF824B60FFABFF99FD96FEBEFEFD2E48" author="GUINOT D. &amp; BOUCHARD J. - M." pageId="6" pageNumber="93" pagination="613 - 694" refId="ref4111" refString="GUINOT D. &amp; BOUCHARD J. - M. 1998. - Evolution of the abdominal holding systems of brachyuran crabs (Crustacea, Decapoda, Brachyura). Zoosystema 20 (4): 613 - 694." type="journal article" year="1998">Guinot &amp; Bouchard (1998)</bibRefCitation>
. Male G1 (
<figureCitation id="13282A14FFABFF99FE70FE9EFD922E48" box="[479,563,299,325]" captionStart="FIG" captionStartId="5.[151,162,1629,1646]" captionTargetBox="[221,1188,259,1558]" captionTargetId="figure-27@5.[181,1252,244,1614]" captionTargetPageId="5" captionText="FIG. 2. — Amarinus pristes n. sp.; A-F, holotype 4.2 × 4.3 mm (MZB); G, H, paratype 5.5 × 5.7 mm (ZRC 2002.596); A, G, carapace; B, left third maxilliped; C, right cheliped; D, left third pereiopod; E, abdomen; F, right G1; G, abdomen. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="6" pageNumber="93">Fig. 2F</figureCitation>
) stout, tip curved, truncate.
</paragraph>
<caption id="DF6C6619FFA8FF9AFF38F9E8FB1C2989" ID-DOI="http://doi.org/10.5281/zenodo.5390323" ID-Zenodo-Dep="5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="5" pageNumber="92" startId="5.[151,162,1629,1646]" targetBox="[221,1188,259,1558]" targetPageId="5">
<paragraph id="8BAC3691FFA8FF9AFF38F9E8FB1C2989" blockId="5.[151,1249,1629,1668]" pageId="5" pageNumber="92">
FIG. 2. —
<taxonomicName id="4C134D12FFA8FF9AFF4BF9E8FECA2963" authority="Rahayu &amp; Ng, 2004" authorityName="Rahayu &amp; Ng" authorityYear="2004" box="[228,363,1629,1646]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="pristes" status="sp. nov.">
<emphasis id="B967EA83FFA8FF9AFF4BF9E8FECA2963" box="[228,363,1629,1646]" italics="true" pageId="5" pageNumber="92">Amarinus pristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F8FFA8FF9AFEDFF9E8FE012963" box="[368,416,1629,1646]" pageId="5" pageNumber="92" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="B967EA83FFA8FF9AFE09F9E8FE642963" bold="true" box="[422,453,1629,1646]" pageId="5" pageNumber="92">A -F</emphasis>
, holotype 4.2 × 4.3 mm (MZB);
<emphasis id="B967EA83FFA8FF9AFD77F9E8FD472963" bold="true" box="[728,742,1629,1646]" pageId="5" pageNumber="92">G</emphasis>
,
<emphasis id="B967EA83FFA8FF9AFD5FF9E8FD5F2963" bold="true" box="[752,766,1629,1646]" pageId="5" pageNumber="92">H</emphasis>
, paratype 5.5 × 5.7 mm (ZRC 2002.596);
<emphasis id="B967EA83FFA8FF9AFBCFF9E8FBCC2963" bold="true" box="[1120,1133,1629,1646]" pageId="5" pageNumber="92">A</emphasis>
,
<emphasis id="B967EA83FFA8FF9AFBD9F9E8FB252963" bold="true" box="[1142,1156,1629,1646]" pageId="5" pageNumber="92">G</emphasis>
, carapace;
<emphasis id="B967EA83FFA8FF9AFF38F9C6FF052989" bold="true" box="[151,164,1651,1668]" pageId="5" pageNumber="92">B</emphasis>
, left third maxilliped;
<emphasis id="B967EA83FFA8FF9AFEF8F9C6FEC42989" bold="true" box="[343,357,1651,1668]" pageId="5" pageNumber="92">C</emphasis>
, right cheliped;
<emphasis id="B967EA83FFA8FF9AFE45F9C6FE592989" bold="true" box="[490,504,1651,1668]" pageId="5" pageNumber="92">D</emphasis>
, left third pereiopod;
<emphasis id="B967EA83FFA8FF9AFD03F9C6FD192989" bold="true" box="[684,696,1651,1668]" pageId="5" pageNumber="92">E</emphasis>
, abdomen;
<emphasis id="B967EA83FFA8FF9AFC9FF9C6FC9A2989" bold="true" box="[816,827,1651,1668]" pageId="5" pageNumber="92">F</emphasis>
, right G1;
<emphasis id="B967EA83FFA8FF9AFC3CF9C6FC002989" bold="true" box="[915,929,1651,1668]" pageId="5" pageNumber="92">G</emphasis>
, abdomen. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BAC3691FFABFF99FF38FE3DFF402EAF" blockId="6.[151,684,392,824]" box="[151,225,392,418]" pageId="6" pageNumber="93">
<emphasis id="B967EA83FFABFF99FF38FE3DFF402EAF" box="[151,225,392,418]" italics="true" pageId="6" pageNumber="93">Female</emphasis>
</paragraph>
<paragraph id="8BAC3691FFABFF99FF38FE18FEC02C35" blockId="6.[151,684,392,824]" pageId="6" pageNumber="93">
In larger female specimens, crenulations on lateral carapace margin stronger (
<figureCitation id="13282A14FFABFF99FE74FE78FD902EEA" box="[475,561,461,487]" captionStart="FIG" captionStartId="5.[151,162,1629,1646]" captionTargetBox="[221,1188,259,1558]" captionTargetId="figure-27@5.[181,1252,244,1614]" captionTargetPageId="5" captionText="FIG. 2. — Amarinus pristes n. sp.; A-F, holotype 4.2 × 4.3 mm (MZB); G, H, paratype 5.5 × 5.7 mm (ZRC 2002.596); A, G, carapace; B, left third maxilliped; C, right cheliped; D, left third pereiopod; E, abdomen; F, right G1; G, abdomen. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="6" pageNumber="93">Fig. 2G</figureCitation>
); in smaller females, crenulations as in male. Chelipeds more slender than those of males, cutting edges of fingers not serrated, without tooth on dactylus. Abdomen (
<figureCitation id="13282A14FFABFF99FEB8FDF2FED22D6C" box="[279,371,583,609]" captionStart="FIG" captionStartId="5.[151,162,1629,1646]" captionTargetBox="[221,1188,259,1558]" captionTargetId="figure-27@5.[181,1252,244,1614]" captionTargetPageId="5" captionText="FIG. 2. — Amarinus pristes n. sp.; A-F, holotype 4.2 × 4.3 mm (MZB); G, H, paratype 5.5 × 5.7 mm (ZRC 2002.596); A, G, carapace; B, left third maxilliped; C, right cheliped; D, left third pereiopod; E, abdomen; F, right G1; G, abdomen. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5390323" httpUri="https://zenodo.org/record/5390323/files/figure.png" pageId="6" pageNumber="93">Fig. 2H</figureCitation>
) five-segmented (excluding pleotelson), submedial groove running almost entire length, defining convex central region; all intersegmental sutures distinct, covering entire sternum for larger specimens (e.g., 5.5 ×
<quantity id="4CEB9B74FFABFF99FDE0FD77FD092DD6" box="[591,680,705,732]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="6" pageNumber="93" unit="mm" value="5.7">5.7 mm</quantity>
, ZRC 2002.596), not covering entire sternum for smaller specimens, longer than broad, pleotelson broadly triangular.
</paragraph>
</subSubSection>
<subSubSection id="C309651AFFABFF99FF38FCDFFD282988" pageId="6" pageNumber="93" type="discussion">
<paragraph id="8BAC3691FFABFF99FF38FCDFFF5A2C8F" blockId="6.[151,684,874,1669]" box="[151,251,874,900]" pageId="6" pageNumber="93">REMARKS</paragraph>
<paragraph id="8BAC3691FFABFF99FF38FC3EFEAE2948" blockId="6.[151,684,874,1669]" pageId="6" pageNumber="93">
The genus
<taxonomicName id="4C134D12FFABFF99FEBEFC3FFED72CA9" authorityName="Lucas" authorityYear="1980" box="[273,374,906,932]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="B967EA83FFABFF99FEBEFC3FFED72CA9" box="[273,374,906,932]" italics="true" pageId="6" pageNumber="93">Amarinus</emphasis>
</taxonomicName>
(
<typeStatus id="54A88833FFABFF99FE29FC3EFE142CA8" box="[390,437,907,933]" pageId="6" pageNumber="93">type</typeStatus>
species
<taxonomicName id="4C134D12FFABFF99FDA1FC3EFEFE2CC8" authority="Chilton, 1882" authorityName="Chilton" authorityYear="1882" class="Malacostraca" family="Hymenosomatidae" genus="Elamena" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="lacustris">
<emphasis id="B967EA83FFABFF99FDA1FC3EFF192CCE" italics="true" pageId="6" pageNumber="93">Elamena lacustris</emphasis>
Chilton, 1882
</taxonomicName>
) is a small genus of wholly to mostly freshwater species with a Gondwanic distribution. Only one species has previously been reported from
<collectingCountry id="F3047601FFABFF99FE99FBBEFE002B28" box="[310,417,1035,1061]" name="Indonesia" pageId="6" pageNumber="93">Indonesia</collectingCountry>
, viz.
<taxonomicName id="4C134D12FFABFF99FE75FBBFFE9D2B48" authority="Ng &amp; Chuang, 1996" authorityName="Ng &amp; Chuang" authorityYear="1996" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B967EA83FFABFF99FE75FBBFFDC32B28" box="[474,610,1034,1061]" italics="true" pageId="6" pageNumber="93">A. crenulatus</emphasis>
<bibRefCitation id="EF824B60FFABFF99FDC5FBBFFE9D2B48" author="NG P. K. L. &amp; CHUANG C. T. N." pageId="6" pageNumber="93" pagination="1 - 82" refId="ref4370" refString="NG P. K. L. &amp; CHUANG C. T. N. 1996. - The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology Supplement 3: 1 - 82." type="journal article" year="1996">Ng &amp; Chuang, 1996</bibRefCitation>
</taxonomicName>
, from
<collectingRegion id="49D7F873FFABFF99FE25FB9EFE4C2B48" box="[394,493,1067,1093]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="93">Sulawesi</collectingRegion>
. With regards to the uneven to serrated lateral carapace margin,
<taxonomicName id="4C134D12FFABFF99FF38FBDFFF582B8E" authority="Rahayu &amp; Ng, 2004" authorityName="Rahayu &amp; Ng" authorityYear="2004" box="[151,249,1130,1156]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="pristes" status="sp. nov.">
<emphasis id="B967EA83FFABFF99FF38FBDFFF582B8E" box="[151,249,1130,1156]" italics="true" pageId="6" pageNumber="93">A. pristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F8FFABFF99FEACFBDFFEE32B89" box="[259,322,1130,1156]" pageId="6" pageNumber="93" rank="species">n. sp.</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C134D12FFABFF99FDB2FBDFFE602BA8" authority="(Balss, 1934) (Philippines)" baseAuthorityName="Balss" baseAuthorityYear="1934" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="wolterecki">
<emphasis id="B967EA83FFABFF99FDB2FBDFFD0D2B88" box="[541,684,1130,1157]" italics="true" pageId="6" pageNumber="93">A. wolterecki</emphasis>
(Balss, 1934) (
<collectingCountry id="F3047601FFABFF99FE95FB3EFE1B2BA8" box="[314,442,1163,1189]" name="Philippines" pageId="6" pageNumber="93">Philippines</collectingCountry>
)
</taxonomicName>
and
<taxonomicName id="4C134D12FFABFF99FE53FB3FFD282BA8" authorityName="Ng &amp; Chuang" authorityYear="1996" box="[508,649,1162,1189]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B967EA83FFABFF99FE53FB3FFD282BA8" box="[508,649,1162,1189]" italics="true" pageId="6" pageNumber="93">A. crenulatus</emphasis>
</taxonomicName>
. It can be distinguished from
<taxonomicName id="4C134D12FFABFF99FE63FB1FFDFB2BC8" baseAuthorityName="Balss" baseAuthorityYear="1934" box="[460,602,1194,1221]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="wolterecki">
<emphasis id="B967EA83FFABFF99FE63FB1FFDFB2BC8" box="[460,602,1194,1221]" italics="true" pageId="6" pageNumber="93">A. wolterecki</emphasis>
</taxonomicName>
by the form of male G1 that is curved and truncate at tip while it is straight and tapering at tip in
<taxonomicName id="4C134D12FFABFF99FDE0FB5FFF692A28" baseAuthorityName="Balss" baseAuthorityYear="1934" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="wolterecki">
<emphasis id="B967EA83FFABFF99FDE0FB5FFF692A28" italics="true" pageId="6" pageNumber="93">A. wolterecki</emphasis>
</taxonomicName>
. The male abdomen of
<taxonomicName id="4C134D12FFABFF99FE78FABFFDC12A28" baseAuthorityName="Balss" baseAuthorityYear="1934" box="[471,608,1290,1317]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="wolterecki">
<emphasis id="B967EA83FFABFF99FE78FABFFDC12A28" box="[471,608,1290,1317]" italics="true" pageId="6" pageNumber="93">A. wolterecki</emphasis>
</taxonomicName>
is also proportionally more slender than in
<taxonomicName id="4C134D12FFABFF99FDE8FA9FFD0D2A4E" authority="Rahayu &amp; Ng, 2004" authorityName="Rahayu &amp; Ng" authorityYear="2004" box="[583,684,1322,1348]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="pristes" status="sp. nov.">
<emphasis id="B967EA83FFABFF99FDE8FA9FFD0D2A4E" box="[583,684,1322,1348]" italics="true" pageId="6" pageNumber="93">A. pristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F8FFABFF99FF38FAFFFF702A69" box="[151,209,1354,1380]" pageId="6" pageNumber="93" rank="species">n. sp.</taxonomicNameLabel>
(see
<bibRefCitation id="EF824B60FFABFF99FEA9FAFFFE442A68" author="NG P. K. L. &amp; CHUANG C. T. N." box="[262,485,1354,1381]" pageId="6" pageNumber="93" pagination="1 - 82" refId="ref4370" refString="NG P. K. L. &amp; CHUANG C. T. N. 1996. - The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology Supplement 3: 1 - 82." type="journal article" year="1996">Ng &amp; Chuang 1996</bibRefCitation>
). From
<taxonomicName id="4C134D12FFABFF99FD95FAFFFF152A8E" authorityName="Ng &amp; Chuang" authorityYear="1996" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B967EA83FFABFF99FD95FAFFFF152A8E" italics="true" pageId="6" pageNumber="93">A. crenulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C134D12FFABFF99FF6BFADFFE832A8E" authority="Rahayu &amp; Ng, 2004" authorityName="Rahayu &amp; Ng" authorityYear="2004" box="[196,290,1386,1412]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="pristes" status="sp. nov.">
<emphasis id="B967EA83FFABFF99FF6BFADFFE832A8E" box="[196,290,1386,1412]" italics="true" pageId="6" pageNumber="93">A. pristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F8FFABFF99FE84FADFFEC92A89" box="[299,360,1386,1412]" pageId="6" pageNumber="93" rank="species">n. sp.</taxonomicNameLabel>
can easily be separated by its relatively shorter ambulatory pereiopods, notably the merus and dactylus (cf.
<bibRefCitation id="EF824B60FFABFF99FE12FA1FFD3D2AC8" author="NG P. K. L. &amp; CHUANG C. T. N." box="[445,668,1450,1477]" pageId="6" pageNumber="93" pagination="1 - 82" refId="ref4370" refString="NG P. K. L. &amp; CHUANG C. T. N. 1996. - The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology Supplement 3: 1 - 82." type="journal article" year="1996">Ng &amp; Chuang 1996</bibRefCitation>
). The male abdomen of
<taxonomicName id="4C134D12FFABFF99FE22FA7FFE4B2AEE" authority="Rahayu &amp; Ng, 2004" authorityName="Rahayu &amp; Ng" authorityYear="2004" box="[397,490,1482,1508]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="pristes" status="sp. nov.">
<emphasis id="B967EA83FFABFF99FE22FA7FFE4B2AEE" box="[397,490,1482,1508]" italics="true" pageId="6" pageNumber="93">A. pristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F8FFABFF99FE5DFA7FFD8C2AE9" box="[498,557,1482,1508]" pageId="6" pageNumber="93" rank="species">n. sp.</taxonomicNameLabel>
agrees with what has been described in detail for the genus by
<bibRefCitation id="EF824B60FFABFF99FF11F9BEFD062928" author="GUINOT D. &amp; RICHER DE FORGES B." box="[190,679,1546,1573]" pageId="6" pageNumber="93" pagination="453 - 502" refId="ref4151" refString="GUINOT D. &amp; RICHER DE FORGES B. 1997. - Affinites entre les Hymenosomatidae MacLeay, 1838 et les Inachoididae Dana, 1851 (Crustacea, Decapoda, Brachyura). Zoosystema 19 (2 - 3): 453 - 502." type="journal article" year="1997">Guinot &amp; Richer de Forges (1997: 467</bibRefCitation>
, fig. 4B-D).
</paragraph>
<paragraph id="8BAC3691FFABFF99FF38F9FEFD282988" blockId="6.[151,684,874,1669]" pageId="6" pageNumber="93">
At present
<taxonomicName id="4C134D12FFABFF99FEA3F9FFFEC9296E" authority="Rahayu &amp; Ng, 2004" authorityName="Rahayu &amp; Ng" authorityYear="2004" box="[268,360,1610,1636]" class="Malacostraca" family="Hymenosomatidae" genus="Amarinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="pristes" status="sp. nov.">
<emphasis id="B967EA83FFABFF99FEA3F9FFFEC9296E" box="[268,360,1610,1636]" italics="true" pageId="6" pageNumber="93">A. pristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F8FFABFF99FEC0F9FFFE0B2969" box="[367,426,1610,1636]" pageId="6" pageNumber="93" rank="species">n. sp.</taxonomicNameLabel>
is only known from the
<typeStatus id="54A88833FFABFF99FF38F9DFFF642989" box="[151,197,1642,1668]" pageId="6" pageNumber="93">type</typeStatus>
locality in Kamora,
<collectingRegion id="49D7F873FFABFF99FE0DF9DEFDAC2989" box="[418,525,1642,1669]" country="Indonesia" name="Papua Barat" pageId="6" pageNumber="93">Irian Jaya</collectingRegion>
,
<collectingCountry id="F3047601FFABFF99FDB7F9DEFD242988" box="[536,645,1643,1669]" name="Indonesia" pageId="6" pageNumber="93">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>