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<document id="D1740CC97DE8B24AF1485B9124A1B528" ID-CLB-Dataset="22295" ID-DOI="10.5852/ejt.2018.457" ID-GBIF-Dataset="38fa93d4-a7aa-45dc-8959-8b32070d950f" ID-Zenodo-Dep="3817911" ID-ZooBank="90509851-4A60-4A51-8D39-64457010B6EE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588968764291" checkinUser="valdenar" docAuthor="Gargiulo, Fabio de F., Brescovit, Antonio D. &amp; Lucas, Sylvia M." docDate="2018" docId="03BA8784FFD8FF8FFECE0F5DFB3A8F77" docLanguage="en" docName="ejt-457_gargiulo_brescovit_lucas.pdf.imf" docOrigin="European Journal of Taxonomy 457" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Umbyquyra acuminata Gargiulo &amp; Brescovit &amp; Lucas 2018, gen. et comb. nov." docType="treatment" docVersion="6" lastPageNumber="25" masterDocId="FF83FFFCFFCEFF97FFE70D2EFF838A58" masterDocTitle="Umbyquyra gen. nov., a new tarantula spider genus from the Neotropical region (Araneae, Mygalomorphae, Theraphosidae), with a description of eight new species" masterLastPageNumber="50" masterPageNumber="1" pageNumber="23" updateTime="1698839066950" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6A204A292E9E335641244DE6C1779A61">Umbyquyra gen. nov., a new tarantula spider genus from the Neotropical region (Araneae, Mygalomorphae, Theraphosidae), with a description of eight new species</mods:title>
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<mods:namePart id="6D5A5A1C6EB3287CE01772FD9C6AA523">Gargiulo, Fabio de F.</mods:namePart>
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<mods:affiliation id="6B97B8E221346C4B31DC93A346CA1D44">Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="23769C336B5989E472A142D9E6EE1A16">Brescovit, Antonio D.</mods:namePart>
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<mods:affiliation id="7855E0BD0CED04A527084E3260C58B8C">Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, São Paulo, Brazil.</mods:affiliation>
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<mods:namePart id="001E16451EF2F23E0E39BB63BB15B905">Lucas, Sylvia M.</mods:namePart>
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<mods:affiliation id="5F3399986EBE21834F05AE721BE8E224">Laboratório Especial de Coleções Zoológicas, Instituto Butantan, Av. Vital Brasil, 1500, São Paulo, Brazil.</mods:affiliation>
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<heading id="D0E481FEFFD8FF81FECE0F5DFA8888D6" box="[297,1291,627,654]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="23" reason="2">
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<emphasis id="B967EA80FFD8FF81FECE0F5DFDBC88D5" bold="true" box="[297,575,627,654]" italics="true" pageId="22" pageNumber="23">Umbyquyra acuminata</emphasis>
(Schmidt &amp; Tesmoingt in 
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)
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<subSubSection id="C3096519FFD8FF81FD410FB8FC0F88E8" box="[678,908,662,689]" pageId="22" pageNumber="23" type="description">
<paragraph id="8BAC3692FFD8FF81FD410FB8FC0F88E8" blockId="22.[297,1291,627,689]" box="[678,908,662,689]" pageId="22" pageNumber="23">
<figureCitation id="13282A17FFD8FF81FD410FB8FD7C88E9" box="[678,767,662,689]" captionStart="Fig" captionStartId="23.[189,232,1883,1909]" captionTargetBox="[189,1398,431,1840]" captionTargetId="figure@23.[189,1399,431,1840]" captionTargetPageId="23" captionText="Fig. 15. Umbyquyra acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov. A–C. ♂, Santa Cruz de la Sierra, Bolivia (ZMB 47188). A. Palp, prolateral. B. Retrolateral. C. Tibial apophysis, retrolateral. D–E. ♀, same locality (ZMB 47189). D. Spermathecae, dorsal. E. Carapace, lateral. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.3817941" httpUri="https://zenodo.org/record/3817941/files/figure.png" pageId="22" pageNumber="23">Figs 15</figureCitation>
, 
<figureCitation id="13282A17FFD8FF81FCEB0FB8FCAB88E8" box="[780,808,662,688]" captionStart="Fig" captionStartId="27.[189,232,1805,1831]" captionTargetId="figure@28.[219,1369,265,2019]" captionTargetPageId="28" captionText="Fig. 18 (opposite page). A–B. Umbyquyra acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov. Live specimen from Santa Cruz de la Sierra, Bolivia (photos by Cristian Grismado). C–F. Umbyquyra cuiaba gen. et sp. nov. C. Live specimen from Parque Nacional Chapada dos Guimarães, Mato Grosso, Brazil, dorsal. D. Defensive display. E. Inside burrow. F. Detail of burrow interior without web. G–H. Umbyquyra paranaiba gen. et sp. nov. G. Live specimen from São Paulo, São Paulo, Brazil, dorsal. H. Defensive display, frontal (C–H, photos by Rafael Indicatti)." figureDoi="http://doi.org/10.5281/zenodo.3817948" httpUri="https://zenodo.org/record/3817948/files/figure.png" pageId="22" pageNumber="23">18</figureCitation>
A–B, 30
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<treatmentCitationGroup id="AB0311BCFFD8FF81FF5A0FF6FB03894E" pageId="22" pageNumber="23">
<treatmentCitation id="0AB21083FFD8FF81FF5A0FF6FCE988AA" author="Schmidt G. &amp; Tesmoingt M." box="[189,874,728,754]" page="2" pageId="22" pageNumber="23" year="2000">
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<emphasis id="B967EA80FFD8FF81FF5A0FF6FE7488AA" box="[189,503,728,754]" italics="true" pageId="22" pageNumber="23">Acanthoscurria acuminata</emphasis>
<bibRefCitation id="EF824B63FFD8FF81FE190FF6FCE988AA" author="Schmidt G. &amp; Tesmoingt M." box="[510,874,728,754]" pageId="22" pageNumber="23" pagination="1 - 6" refId="ref24552" refString="Schmidt G. &amp; Tesmoingt M. 2000. Eine weitere Acanthoscurria- Art aus Brasilien und Bolivien (Araneae: Theraphosidae: Theraphosinae). Arachnologisches Magazin 8: 1 - 6." type="journal article" year="2000">Schmidt &amp; Tesmoingt, 2000: 2</bibRefCitation>
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, figs 1–2. (nom. nud., no 
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depository). 
<treatmentCitation id="0AB21083FFD8FF81FF5A0FD5FC72894D" author="Schmidt G." box="[189,1009,763,790]" page="4" pageId="22" pageNumber="23" year="2005">
<taxonomicName id="4C134D11FFD8FF81FF5A0FD5FC72894D" ID-CoL="92R4K" authority="Schmidt &amp; Tesmoingt in Schmidt, 2005: 4" authorityName="Schmidt &amp; Tesmoingt in Schmidt" authorityPageNumber="4" authorityYear="2005" box="[189,1009,763,790]" class="Arachnida" family="Theraphosidae" genus="Acanthoscurria" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="acuminata">
<emphasis id="B967EA80FFD8FF81FF5A0FD5FE74894D" box="[189,503,763,789]" italics="true" pageId="22" pageNumber="23">Acanthoscurria acuminata</emphasis>
Schmidt &amp; Tesmoingt in 
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, figs 1–2, 7.
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<treatmentCitationGroup id="AB0311BCFFD8FF81FF5A0E12FBF4890F" box="[189,1143,828,855]" pageId="22" pageNumber="23">
<emphasis id="B967EA80FFD8FF81FF5A0E12FD8E890E" box="[189,525,828,854]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C134D11FFD8FF81FF5A0E12FE74890E" ID-CoL="92R4K" authorityName="Schmidt &amp; Tesmoingt in Schmidt" authorityYear="2005" box="[189,503,828,854]" class="Arachnida" family="Theraphosidae" genus="Acanthoscurria" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="acuminata">Acanthoscurria acuminata</taxonomicName>
–
</emphasis>
<treatmentCitation id="0AB21083FFD8FF81FDF30E12FCC6890F" box="[532,837,828,855]" pageId="22" pageNumber="23">Schmidt 2000: 1, figs 1–2</treatmentCitation>
; 2003: 141, figs 235–238.
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</paragraph>
</subSubSection>
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<paragraph id="8BAC3692FFD8FF81FF5A0E50FEB589C0" blockId="22.[189,310,894,920]" box="[189,310,894,920]" pageId="22" pageNumber="23">
<heading id="D0E481FEFFD8FF81FF5A0E50FEB589C0" bold="true" box="[189,310,894,920]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B967EA80FFD8FF81FF5A0E50FEB589C0" bold="true" box="[189,310,894,920]" pageId="22" pageNumber="23">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3692FFD8FF81FF5A0E82FCF38E0C" blockId="22.[189,1398,940,1108]" pageId="22" pageNumber="23">
Males and females of 
<taxonomicName id="4C134D11FFD8FF81FE500E83FDD6899E" authorityName="Gargiulo &amp; Brescovit &amp; Lucas" authorityYear="2018" baseAuthorityName="Schmidt &amp; Tesmoingt in Schmidt" baseAuthorityYear="2005" box="[439,597,941,966]" class="Arachnida" family="Theraphosidae" genus="Acanthoscurria" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="acuminata" status="gen. et comb. nov.">
<emphasis id="B967EA80FFD8FF81FE500E83FDD6899E" box="[439,597,941,966]" italics="true" pageId="22" pageNumber="23">U. acuminata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457FBFFD8FF81FDBD0E83FCA8899F" box="[602,811,940,967]" pageId="22" pageNumber="23" rank="species">gen. et comb. nov.</taxonomicNameLabel>
differ from those of the other species by a procurve fovea overlapped by a large projection of the cephalic region (
<figureCitation id="13282A17FFD8FF81FC7D0EFEFB8189B2" box="[922,1026,975,1002]" captionStart="Fig" captionStartId="23.[189,232,1883,1909]" captionTargetBox="[189,1398,431,1840]" captionTargetId="figure@23.[189,1399,431,1840]" captionTargetPageId="23" captionText="Fig. 15. Umbyquyra acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov. A–C. ♂, Santa Cruz de la Sierra, Bolivia (ZMB 47188). A. Palp, prolateral. B. Retrolateral. C. Tibial apophysis, retrolateral. D–E. ♀, same locality (ZMB 47189). D. Spermathecae, dorsal. E. Carapace, lateral. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.3817941" httpUri="https://zenodo.org/record/3817941/files/figure.png" pageId="22" pageNumber="23">Fig. 15E</figureCitation>
) and by a retrolateral branch of the tibial apophysis with three strong spines (
<figureCitation id="13282A17FFD8FF81FD3C0EDDFCC48E56" box="[731,839,1011,1038]" captionStart="Fig" captionStartId="23.[189,232,1883,1909]" captionTargetBox="[189,1398,431,1840]" captionTargetId="figure@23.[189,1399,431,1840]" captionTargetPageId="23" captionText="Fig. 15. Umbyquyra acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov. A–C. ♂, Santa Cruz de la Sierra, Bolivia (ZMB 47188). A. Palp, prolateral. B. Retrolateral. C. Tibial apophysis, retrolateral. D–E. ♀, same locality (ZMB 47189). D. Spermathecae, dorsal. E. Carapace, lateral. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.3817941" httpUri="https://zenodo.org/record/3817941/files/figure.png" pageId="22" pageNumber="23">Fig. 15C</figureCitation>
). Female spermathecae have a cordiform basal area resembling those of 
<taxonomicName id="4C134D11FFD8FF81FE0F0939FDF38E68" authorityName="Gargiulo &amp; Brescovit &amp; Lucas" authorityYear="2018" baseAuthorityName="Rudloff" baseAuthorityYear="1996" box="[488,624,1046,1072]" class="Arachnida" family="Theraphosidae" genus="Umbyquyra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="schmidti" status="gen. et comb. nov.">
<emphasis id="B967EA80FFD8FF81FE0F0939FDF38E68" box="[488,624,1046,1072]" italics="true" pageId="22" pageNumber="23">U. schmidti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457FBFFD8FF81FD9E0939FCD58E69" box="[633,854,1046,1073]" pageId="22" pageNumber="23" rank="species">gen. et comb. nov.</taxonomicNameLabel>
, but differs from this species by the RS being more apart when compared to the other species (
<figureCitation id="13282A17FFD8FF81FD110917FCE08E0C" box="[758,867,1081,1108]" captionStart="Fig" captionStartId="23.[189,232,1883,1909]" captionTargetBox="[189,1398,431,1840]" captionTargetId="figure@23.[189,1399,431,1840]" captionTargetPageId="23" captionText="Fig. 15. Umbyquyra acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov. A–C. ♂, Santa Cruz de la Sierra, Bolivia (ZMB 47188). A. Palp, prolateral. B. Retrolateral. C. Tibial apophysis, retrolateral. D–E. ♀, same locality (ZMB 47189). D. Spermathecae, dorsal. E. Carapace, lateral. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.3817941" httpUri="https://zenodo.org/record/3817941/files/figure.png" pageId="22" pageNumber="23">Fig. 15D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3096519FFD8FF81FF5A0954FB7A8FCA" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BAC3692FFD8FF81FF5A0954FE298ECC" blockId="22.[189,426,1146,1172]" box="[189,426,1146,1172]" pageId="22" pageNumber="23">
<heading id="D0E481FEFFD8FF81FF5A0954FE298ECC" bold="true" box="[189,426,1146,1172]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B967EA80FFD8FF81FF5A0954FE298ECC" bold="true" box="[189,426,1146,1172]" pageId="22" pageNumber="23">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B7B3CCFFFD8FF81FF0D0987FE048F52" ID-GBIF-Occurrence="2611493316" collectionCode="SMF" collectorName="Verdez" country="Bolivia" latitude="-16.460335" location="Province Sara" longLatPrecision="1" longitude="-63.181225" pageId="22" pageNumber="23" specimenCode="SMF 3979" specimenCount="1" specimenCount-female="1" stateProvince="Santa Cruz" typeStatus="holotype">
<paragraph id="8BAC3692FFD8FF81FF0D0987FEDF8E9B" blockId="22.[189,1398,1193,1290]" box="[234,348,1193,1219]" pageId="22" pageNumber="23">
<emphasis id="B967EA80FFD8FF81FF0D0987FEDF8E9B" bold="true" box="[234,348,1193,1219]" pageId="22" pageNumber="23">
<typeStatus id="54A88830FFD8FF81FF0D0987FEDF8E9B" box="[234,348,1193,1219]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAC3692FFD8FF81FF5A09E3FE048F52" blockId="22.[189,1398,1193,1290]" pageId="22" pageNumber="23">
<collectingCountry id="F3047602FFD8FF81FF5A09E3FEB98EBF" box="[189,314,1229,1255]" name="Bolivia" pageId="22" pageNumber="23">BOLIVIA</collectingCountry>
: 
<specimenCount id="9D15FD1BFFD8FF81FEA009E3FEF68EBF" box="[327,373,1229,1255]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
, 
<location id="8ECC6049FFD8FF81FE6109E2FDAC8EBF" LSID="urn:lsid:plazi:treatment:03BA8784FFD8FF8FFECE0F5DFB3A8F77:8ECC6049FFD8FF81FE6109E2FDAC8EBF" box="[390,559,1228,1255]" country="Bolivia" latitude="-16.460335" longLatPrecision="1" longitude="-63.181225" name="Province Sara" pageId="22" pageNumber="23" stateProvince="Santa Cruz">Province Sara</location>
, 
<collectingRegion id="49D7F870FFD8FF81FDDB09E3FCD58EBE" box="[572,854,1228,1255]" country="Bolivia" name="Santa Cruz" pageId="22" pageNumber="23">Santa Cruz de la Sierra</collectingRegion>
, 
<geoCoordinate id="EE275055FFD8FF81FC8309E5FB8F8EBF" box="[868,1036,1227,1255]" degrees="16" direction="south" minutes="27" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="37.2" value="-16.460335">16°27′37.2″ S</geoCoordinate>
, 
<geoCoordinate id="EE275055FFD8FF81FBFC09E5FB4F8EBF" box="[1051,1228,1227,1255]" degrees="63" direction="west" minutes="10" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="52.4" value="-63.181225">63°10′52.4″ W</geoCoordinate>
, 1995, 
<collectorName id="26E65344FFD8FF81FAC209E2FAF58EBE" box="[1317,1398,1228,1254]" pageId="22" pageNumber="23">Verdez</collectorName>
leg. (
<specimenCode id="DBB59EE9FFD8FF81FF1009DEFEF88F52" box="[247,379,1264,1290]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="22" pageNumber="23">SMF 3979</specimenCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="3B7B3CCFFFD8FF81FF0D081FFB7A8FCA" ID-GBIF-Occurrence="2611493305" pageId="22" pageNumber="23">
<paragraph id="8BAC3692FFD8FF81FF0D081FFE628F13" blockId="22.[189,1398,1329,1426]" box="[234,481,1329,1355]" pageId="22" pageNumber="23">
<emphasis id="B967EA80FFD8FF81FF0D081FFE628F13" bold="true" box="[234,481,1329,1355]" pageId="22" pageNumber="23">Additional material</emphasis>
</paragraph>
<paragraph id="8BAC3692FFD8FF81FF5A087AFB7A8FCA" blockId="22.[189,1398,1329,1426]" pageId="22" pageNumber="23">
<collectingCountry id="F3047602FFD8FF81FF5A087AFEBA8F36" box="[189,313,1364,1390]" name="Bolivia" pageId="22" pageNumber="23">BOLIVIA</collectingCountry>
, 1 ♂, Province Sara, 
<collectingRegion id="49D7F870FFD8FF81FDDF087AFCCC8F36" box="[568,847,1364,1390]" country="Bolivia" name="Santa Cruz" pageId="22" pageNumber="23">Santa Cruz de la Sierra</collectingRegion>
, 
<geoCoordinate id="EE275055FFD8FF81FCBB087CFB808F36" box="[860,1027,1362,1390]" degrees="16" direction="south" minutes="58" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="16.4" value="-16.971222">16°58′16.4″ S</geoCoordinate>
, 
<geoCoordinate id="EE275055FFD8FF81FBF6087CFB418F37" box="[1041,1218,1362,1391]" degrees="63" direction="west" minutes="34" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="22.5" value="-63.572914">63°34′22.5″ W</geoCoordinate>
, 
<date id="FFAD1052FFD8FF81FB28087AFAF18F37" box="[1231,1394,1364,1391]" pageId="22" pageNumber="23" value="2005-12-29">29 Dec. 2005</date>
, Steinbach leg. (
<collectionCode id="ED02AE57FFD8FF81FE950856FE308FCA" box="[370,435,1400,1426]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="22" pageNumber="23" type="Museum">ZMB</collectionCode>
47188); 
<specimenCount id="9D15FD1BFFD8FF81FDF90856FDE28FCA" box="[542,609,1400,1426]" pageId="22" pageNumber="23" type="female">2 ♀♀</specimenCount>
, same data as for preceding (
<collectionCode id="ED02AE57FFD8FF81FC510856FC748FCA" box="[950,1015,1400,1426]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="22" pageNumber="23" type="Museum">ZMB</collectionCode>
47189, 
<collectionCode id="ED02AE57FFD8FF81FBB00856FB148FCA" box="[1111,1175,1400,1426]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="22" pageNumber="23" type="Museum">ZMB</collectionCode>
47190).
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C3096519FFD8FF8FFF5A0897FACC894B" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="description">
<paragraph id="8BAC3692FFD8FF81FF5A0897FECD8F8B" blockId="22.[189,334,1465,1491]" box="[189,334,1465,1491]" pageId="22" pageNumber="23">
<heading id="D0E481FEFFD8FF81FF5A0897FECD8F8B" bold="true" box="[189,334,1465,1491]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B967EA80FFD8FF81FF5A0897FECD8F8B" bold="true" box="[189,334,1465,1491]" pageId="22" pageNumber="23">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3692FFD8FF81FF0D08C6FE568C5A" blockId="22.[189,1399,1512,1996]" box="[234,469,1512,1538]" pageId="22" pageNumber="23">
<emphasis id="B967EA80FFD8FF81FF0D08C6FEA98C5A" bold="true" box="[234,298,1512,1538]" pageId="22" pageNumber="23">Male</emphasis>
(ZMB 47188)
</paragraph>
<paragraph id="8BAC3692FFD8FF80FF5A0B25FE708B34" blockId="22.[189,1399,1512,1996]" lastBlockId="23.[189,1399,267,364]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
In life: dark brown with light brown joints and golden hairs completely covering dorsal side of body (
<figureCitation id="13282A17FFD8FF81FF210B00FE9E8C11" box="[198,285,1582,1609]" captionStart="Fig" captionStartId="25.[189,232,1879,1905]" captionTargetBox="[191,1399,446,1840]" captionTargetId="figure@25.[191,1401,446,1840]" captionTargetPageId="25" captionText="Fig. 16. Umbyquyra cuiaba gen. et sp. nov. A–C. Holotype, ♂, Cuiabá, Mato Grosso, Brasil (IBSP 167423). A. Palp prolateral. B. Retrolateral. C. Tibial apophysis, retrolateral. D–F. Paratype, ♀, Canarana, Mato Grosso, Brazil (IBSP 108933). D. Spermathecae, dorsal. E. Eye tubercle, dorsal. F. Abdomen, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3817944" httpUri="https://zenodo.org/record/3817944/files/figure.png" pageId="22" pageNumber="23">Fig. 16</figureCitation>
). In ethanol: same color as 
<taxonomicName id="4C134D11FFD8FF81FDBA0B01FD798C10" authority="Gargiulo &amp; Brescovit &amp; Lucas, 2018" authorityName="Gargiulo &amp; Brescovit &amp; Lucas" authorityYear="2018" box="[605,762,1582,1608]" class="Arachnida" family="Theraphosidae" genus="Umbyquyra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="paranaiba" status="gen. et sp. nov.">
<emphasis id="B967EA80FFD8FF81FDBA0B01FD798C10" box="[605,762,1582,1608]" italics="true" pageId="22" pageNumber="23">U. paranaiba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457FBFFD8FF81FCE50B00FC328C10" box="[770,945,1582,1608]" pageId="22" pageNumber="23" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Total length 37.8. Carapace 17.4 long, 15 wide. Clypeus 0.3. Eye tubercle 2.2 long, 1.7 wide. AME 0.4 long, 0.4 wide; PME 0.28 long, 0.23 wide; ALE 0.4 long, 0.3 wide; PLE 0.4 long, 0.25 wide. Basal segment of chelicerae with 12 teeth. Stridulatory bristles with around 15 bristles on palp trochanter and 45 on leg I (
<figureCitation id="13282A17FFD8FF81FBB50BB9FB278CEA" box="[1106,1188,1687,1714]" captionStart="Fig" captionStartId="26.[189,232,1879,1905]" captionTargetBox="[189,1398,265,1834]" captionTargetId="figure@26.[189,1398,265,1834]" captionTargetPageId="26" captionText="Fig. 17. Umbyquyra cuiaba gen. et sp. nov., holotype, ♂, Cuiabá, Mato Grosso, Brazil (IBSP 167423). A. Palp, prolateral. B. Anterolateral. C. Retrolateral. D. Tibial apophysis, retrolateral. E. Stridulatory organ, palp trochanter. F. Trochanter I. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.3817946" httpUri="https://zenodo.org/record/3817946/files/figure.png" pageId="22" pageNumber="23">Fig. 17</figureCitation>
E–F). Palp: femur 8.3, patella 3.7, tibia 8.4, tarsus 3.7, total 24.1. Leg I femur 13.4, patella 4.5, tibia 12.5, metatarsus 12.7, tarsus 7.8, total 51. II 13.3, 4.6, 11.2, 6, 7.2, 42.3. III 11.6, 3.1, 9.5, 11.5, 6.8, 42.5. IV 15.8, 5.5, 12.8, 16.8, 7.5, 58.4. Spination: palp: tibia v0-1p-0ap, r0-1-2-1-2-1-1-1ap. Leg I femur d0-0-1app, patella v0-1-1-1-0ap, tibia d0-1p-0-0ap, v2-1r-1-0-1-1-1-1r-1-2apr, p0-0-1-0ap, r0-0-0-1-0ap, metatarsus v0- 1r-0-0ap. II femur, 0-0-0-1app, tibia v3-1r-1-1-1-1r-1-0-1apr-1app-1ap, p0-1-0-1-0-1ap, metatarsus v0-1p-1-0-0-0-1app-1ap-1apr, p0-0-0ap. III femur d0-0-1apr, tibia v2-1-0-1-2-1p-1r-0-1apr-1app, p0-1- 1-1-1-1ap, r0-1-0-1-0-0ap, metatarsus v0-0-1p-1r-0-1p-1r-0-2apr-1ap-1app, p2-1-1-1-1-0-0ap, r0-0-1- 0-0ap. IV femur d0-0-1apr, tibia v2-1-1p-1-1-1-2-1r-1apr-1app, r0-0-1-0ap, metatarsus d0-0-1r-1-1p-0- 0ap, r0-0-1-0-0ap, v1p-1r-1r-1-1p-1r-1-1-1p-1-1-1apr-1app. Tibial apophysis of leg I: prolateral branch inconspicuous with distal and sub distal spines. (
<figureCitation id="13282A17FFD9FF80FD110C00FCE28B11" box="[758,865,302,329]" captionStart="Fig" captionStartId="23.[189,232,1883,1909]" captionTargetBox="[189,1398,431,1840]" captionTargetId="figure@23.[189,1399,431,1840]" captionTargetPageId="23" captionText="Fig. 15. Umbyquyra acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov. A–C. ♂, Santa Cruz de la Sierra, Bolivia (ZMB 47188). A. Palp, prolateral. B. Retrolateral. C. Tibial apophysis, retrolateral. D–E. ♀, same locality (ZMB 47189). D. Spermathecae, dorsal. E. Carapace, lateral. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.3817941" httpUri="https://zenodo.org/record/3817941/files/figure.png" pageId="23" pageNumber="24">Fig. 15C</figureCitation>
). PLS basal, median and apical segments 1.8, 1.1, 2.8 long, respectively.
</paragraph>
<caption id="DF6C661AFFD9FF80FF5A0A75FD248D86" ID-DOI="http://doi.org/10.5281/zenodo.3817941" ID-Zenodo-Dep="3817941" httpUri="https://zenodo.org/record/3817941/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,232,1883,1909]" targetBox="[189,1398,431,1840]" targetPageId="23">
<paragraph id="8BAC3692FFD9FF80FF5A0A75FD248D86" blockId="23.[189,1399,1883,2015]" pageId="23" pageNumber="24">
<emphasis id="B967EA80FFD9FF80FF5A0A75FE998D2D" bold="true" box="[189,282,1883,1909]" pageId="23" pageNumber="24">Fig. 15.</emphasis>
<taxonomicName id="4C134D11FFD9FF80FEC40A75FBA68D2E" authority="(Schmidt &amp; Tesmoingt in Schmidt, 2005)" authorityName="Gargiulo &amp; Brescovit &amp; Lucas" authorityYear="2018" baseAuthorityName="Schmidt &amp; Tesmoingt in Schmidt" baseAuthorityYear="2005" box="[291,1061,1883,1910]" class="Arachnida" family="Theraphosidae" genus="Umbyquyra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="acuminata" status="gen. et comb. nov.">
<emphasis id="B967EA80FFD9FF80FEC40A75FDB28D2D" box="[291,561,1883,1909]" italics="true" pageId="23" pageNumber="24">Umbyquyra acuminata</emphasis>
(Schmidt &amp; Tesmoingt in 
<bibRefCitation id="EF824B63FFD9FF80FC880A75FB9E8D2E" author="Schmidt G." box="[879,1053,1883,1910]" pageId="23" pageNumber="24" pagination="3 - 49" refId="ref24512" refString="Schmidt G. 2005. Neupublikation einiger von mir in &quot; Arachnol. Mag. &quot; und &quot; Tarantulas of the World &quot; zwischen 2000 und 2005 veroffentlichten Arbeiten. Tarantulas of the World 110: 3 - 49." type="journal article" year="2005">Schmidt, 2005</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25457FBFFD9FF80FBC90A75FA8B8D2D" box="[1070,1288,1883,1909]" pageId="23" pageNumber="24" rank="species">gen. et comb. nov.</taxonomicNameLabel>
<emphasis id="B967EA80FFD9FF80FAF70A75FACA8D2D" bold="true" box="[1296,1353,1883,1909]" pageId="23" pageNumber="24">A–C</emphasis>
. ♂, Santa Cruz de la Sierra, Bolivia (ZMB 47188). 
<emphasis id="B967EA80FFD9FF80FD0D0A50FD7C8DC0" bold="true" box="[746,767,1918,1944]" pageId="23" pageNumber="24">A</emphasis>
. Palp, prolateral. 
<emphasis id="B967EA80FFD9FF80FC280A50FC608DC0" bold="true" box="[975,995,1918,1944]" pageId="23" pageNumber="24">B</emphasis>
. Retrolateral. 
<emphasis id="B967EA80FFD9FF80FB6D0A50FB1C8DC0" bold="true" box="[1162,1183,1918,1944]" pageId="23" pageNumber="24">C</emphasis>
. Tibial apophysis, retrolateral. 
<emphasis id="B967EA80FFD9FF80FEB40A8FFE098DE3" bold="true" box="[339,394,1953,1979]" pageId="23" pageNumber="24">D–E</emphasis>
. ♀, same locality (ZMB 47189). 
<emphasis id="B967EA80FFD9FF80FCDD0A8FFCCC8DE3" bold="true" box="[826,847,1953,1979]" pageId="23" pageNumber="24">D</emphasis>
. Spermathecae, dorsal. 
<emphasis id="B967EA80FFD9FF80FB9C0A8FFB0C8DE3" bold="true" box="[1147,1167,1953,1979]" pageId="23" pageNumber="24">E</emphasis>
. Carapace, lateral. Abbreviations: see Material and methods.
</paragraph>
</caption>
<paragraph id="8BAC3692FFD6FF8FFF0D0C22FE738B7D" blockId="24.[188,1399,267,787]" box="[234,496,267,294]" pageId="24" pageNumber="25">
<emphasis id="B967EA80FFD6FF8FFF0D0C22FEC68B7E" bold="true" box="[234,325,268,294]" pageId="24" pageNumber="25">Female</emphasis>
(ZMB 47189)
</paragraph>
<paragraph id="8BAC3692FFD6FF8FFF5B0C01FACC894B" blockId="24.[188,1399,267,787]" pageId="24" pageNumber="25">
In ethanol: same color as 
<taxonomicName id="4C134D11FFD6FF8FFE0D0C01FD0B8B11" authority="Gargiulo &amp; Brescovit &amp; Lucas, 2018" authorityName="Gargiulo &amp; Brescovit &amp; Lucas" authorityYear="2018" box="[490,648,303,329]" class="Arachnida" family="Theraphosidae" genus="Umbyquyra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="paranaiba" status="gen. et sp. nov.">
<emphasis id="B967EA80FFD6FF8FFE0D0C01FD0B8B11" box="[490,648,303,329]" italics="true" pageId="24" pageNumber="25">U. paranaiba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457FBFFD6FF8FFD770C01FCC28B11" box="[656,833,303,329]" pageId="24" pageNumber="25" rank="species">gen. et sp. nov.</taxonomicNameLabel>
Total length 32.2. Carapace 16.6 long, 13 wide. Clypeus 0.3. Eyes: anterior row slightly procurved and posterior recurved. AME 0.4 long, 0.4 wide; PME 0.27 long, 0.27 wide; ALE 0.48 long, 0.33 wide; PLE 0.35 long, 0.22 wide. Basal segment of chelicerae with 12 teeth. 7.0 long, 7.0 wide. Eye tubercle 1.7 long, 2.4 wide. Stridulatory bristles as in male. Palp: femur 7.6, patella 2.7, tibia 6.0, tarsus 5.5, total 21.8. Legs I femur 9.7, patella 3.5, tibia 5.3, metatarsus 4.6, tarsus 4.6, total 27.7. II femur 9.0, patella 3.0, tibia 7.0, metatarsus 5.5, tarsus 3.7, total 28.2. III femur 8.3, patella 3.0, tibia 5.5, metatarsus 6.5, tarsus 4.1, total 27.4. IV femur 8.6, patella 4.0, tibia 4.6, metatarsus 6.4, tarsus 4.2, total 27.8; Spination: palp: tibia v0-0-1p-1apr-2app, r0-0-1-0-1ap. Leg I femur d0-0-1app, tibia v0-1-0-1-1-1-1-0-1ap-1app, metatarsus v0-1-1-0-1app-1ap-1apr. II femur d0-0-1app, tibia v0-1-0-1-1-1-0-1app-1apr, metatarsus v0-1-1-0-1app-1ap-1apr. III femur d0-0-1apr, tibia v0-2-0-1-2app-1apr, p0-1-1-0-2ap, metatarsus d0-1p-1r-0-1app-2apr, p0-1-1-1-0-2ap. IV v0-1- 1-0-1app-2apr, p0-1-0-1ap, r1-1-1-0ap, metatarsus d0-1p-1r-0-1app-2apr, v1-1p-1-1p-1r-0-1apr-1ap- 1app, p1-1-1-0-1ap, r0-0-0-1ap. PLS basal, median and apical segments 2.5, 1.4, 2.0 long, respectively. Spermathecae with oval RS and long ducts, apart by approximately twice their diameter (
<figureCitation id="13282A17FFD6FF8FFB320FD6FAC2894B" box="[1237,1345,760,787]" captionStart="Fig" captionStartId="23.[189,232,1883,1909]" captionTargetBox="[189,1398,431,1840]" captionTargetId="figure@23.[189,1399,431,1840]" captionTargetPageId="23" captionText="Fig. 15. Umbyquyra acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov. A–C. ♂, Santa Cruz de la Sierra, Bolivia (ZMB 47188). A. Palp, prolateral. B. Retrolateral. C. Tibial apophysis, retrolateral. D–E. ♀, same locality (ZMB 47189). D. Spermathecae, dorsal. E. Carapace, lateral. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.3817941" httpUri="https://zenodo.org/record/3817941/files/figure.png" pageId="24" pageNumber="25">Fig. 15D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3096519FFD6FF8FFF5B0E14FEFE8E98" pageId="24" pageNumber="25" type="discussion">
<paragraph id="8BAC3692FFD6FF8FFF5B0E14FEA0890C" blockId="24.[188,291,826,852]" box="[188,291,826,852]" pageId="24" pageNumber="25">
<heading id="D0E481FEFFD6FF8FFF5B0E14FEA0890C" bold="true" box="[188,291,826,852]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B967EA80FFD6FF8FFF5B0E14FEA0890C" bold="true" box="[188,291,826,852]" pageId="24" pageNumber="25">Remark</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3692FFD6FF8FFF5B0E46FEFE8E98" blockId="24.[188,1399,872,1216]" pageId="24" pageNumber="25">
<bibRefCitation id="EF824B63FFD6FF8FFF5B0E46FD9E89DA" author="Schmidt G. &amp; Tesmoingt M." box="[188,541,872,898]" pageId="24" pageNumber="25" pagination="1 - 6" refId="ref24552" refString="Schmidt G. &amp; Tesmoingt M. 2000. Eine weitere Acanthoscurria- Art aus Brasilien und Bolivien (Araneae: Theraphosidae: Theraphosinae). Arachnologisches Magazin 8: 1 - 6." type="journal article" year="2000">Schmidt &amp; Tesmoingt (2000)</bibRefCitation>
described 
<taxonomicName id="4C134D11FFD6FF8FFD460E46FC6389DA" authorityName="Gargiulo &amp; Brescovit &amp; Lucas" authorityYear="2018" baseAuthorityName="Schmidt &amp; Tesmoingt in Schmidt" baseAuthorityYear="2005" box="[673,992,872,898]" class="Arachnida" family="Theraphosidae" genus="Acanthoscurria" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="acuminata">
<emphasis id="B967EA80FFD6FF8FFD460E46FC6389DA" box="[673,992,872,898]" italics="true" pageId="24" pageNumber="25">Acanthoscurria acuminata</emphasis>
</taxonomicName>
based on a female and a juvenile male and exuvia, respectively from 
<collectingRegion id="49D7F870FFD6FF8FFD8E0EA5FC0389FD" box="[617,896,907,933]" country="Bolivia" name="Santa Cruz" pageId="24" pageNumber="25">Santa Cruz de la Sierra</collectingRegion>
, Province Sara, 
<collectingCountry id="F3047602FFD6FF8FFBA50EA5FB1A89FD" box="[1090,1177,907,933]" name="Bolivia" pageId="24" pageNumber="25">Bolívia</collectingCountry>
and Cuiabá, 
<collectingRegion id="49D7F870FFD6FF8FFADF0EA2FE908991" country="Brazil" name="Mato Grosso" pageId="24" pageNumber="25">Mato Grosso</collectingRegion>
, 
<collectingCountry id="F3047602FFD6FF8FFEC60E80FEEE8990" box="[289,365,942,968]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
. They presented figures of the carapace with a large projection over the fovea and the aspect of the SR. Later, Schmidt &amp; Tesmoingt (2005) described another male of this species, collected in 
<collectingRegion id="49D7F870FFD6FF8FFF380EDBFD858E57" box="[223,518,1013,1039]" country="Bolivia" name="Santa Cruz" pageId="24" pageNumber="25">Santa Cruz de la Sierra</collectingRegion>
, 
<collectingCountry id="F3047602FFD6FF8FFDF10EDBFDF38E57" box="[534,624,1013,1039]" name="Bolivia" pageId="24" pageNumber="25">Bolivia</collectingCountry>
, and deposited it in the regional museum of 
<collectingRegion id="49D7F870FFD6FF8FFB400EDBFAB18E57" box="[1191,1330,1013,1039]" country="Bolivia" name="Santa Cruz" pageId="24" pageNumber="25">Santa Cruz</collectingRegion>
. This specimen could not be located. The female 
<typeStatus id="54A88830FFD6FF8FFD280936FCB48E6A" box="[719,823,1048,1074]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
is deposited in the SMF and does not have the spermathecae anymore. Moreover, the juvenile male and exuvia could not be located. The species here is transferred to 
<taxonomicName id="4C134D11FFD6FF8FFE9B0971FD8B8E21" authority="Gargiulo &amp; Brescovit &amp; Lucas, 2018" authorityName="Gargiulo &amp; Brescovit &amp; Lucas" authorityYear="2018" box="[380,520,1119,1145]" class="Arachnida" family="Theraphosidae" genus="Umbyquyra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B967EA80FFD6FF8FFE9B0971FD8B8E21" box="[380,520,1119,1145]" italics="true" pageId="24" pageNumber="25">Umbyquyra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457FBFFD6FF8FFDE80971FDF98E21" box="[527,634,1119,1145]" pageId="24" pageNumber="25" rank="genus">gen. nov.</taxonomicNameLabel>
due to the presence of stridulatory bristles on the palp trochanter and leg I, male palpal bulb with short embolus, four keels and spermathecae emerging from a common base (
<figureCitation id="13282A17FFD6FF8FFEE6098BFEF38E98" box="[257,368,1189,1216]" captionStart="Fig" captionStartId="23.[189,232,1883,1909]" captionTargetBox="[189,1398,431,1840]" captionTargetId="figure@23.[189,1399,431,1840]" captionTargetPageId="23" captionText="Fig. 15. Umbyquyra acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov. A–C. ♂, Santa Cruz de la Sierra, Bolivia (ZMB 47188). A. Palp, prolateral. B. Retrolateral. C. Tibial apophysis, retrolateral. D–E. ♀, same locality (ZMB 47189). D. Spermathecae, dorsal. E. Carapace, lateral. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.3817941" httpUri="https://zenodo.org/record/3817941/files/figure.png" pageId="24" pageNumber="25">Fig. 15D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3096519FFD6FF8FFF5B09C8FB3A8F77" box="[188,1209,1254,1327]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BAC3692FFD6FF8FFF5B09C8FED68F58" blockId="24.[188,341,1254,1280]" box="[188,341,1254,1280]" pageId="24" pageNumber="25">
<heading id="D0E481FEFFD6FF8FFF5B09C8FED68F58" bold="true" box="[188,341,1254,1280]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B967EA80FFD6FF8FFF5B09C8FED68F58" bold="true" box="[188,341,1254,1280]" pageId="24" pageNumber="25">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC3692FFD6FF8FFF5B083BFB3A8F77" blockId="24.[188,1209,1301,1327]" box="[188,1209,1301,1327]" pageId="24" pageNumber="25">
<collectingCountry id="F3047602FFD6FF8FFF5B083BFE8B8F77" box="[188,264,1301,1327]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
: Cuiabá, 
<collectingRegion id="49D7F870FFD6FF8FFE92083BFD918F77" box="[373,530,1301,1327]" country="Brazil" name="Mato Grosso" pageId="24" pageNumber="25">Mato Grosso</collectingRegion>
; 
<collectingCountry id="F3047602FFD6FF8FFDFA083BFDFB8F77" box="[541,632,1301,1327]" name="Bolivia" pageId="24" pageNumber="25">Bolivia</collectingCountry>
: 
<collectingRegion id="49D7F870FFD6FF8FFD64083BFC178F77" box="[643,916,1301,1327]" country="Bolivia" name="Santa Cruz" pageId="24" pageNumber="25">Santa Cruz de la Sierra</collectingRegion>
, Province Sara (
<figureCitation id="13282A17FFD6FF8FFBB3083BFB2F8F77" box="[1108,1196,1301,1327]" captionStart="Fig" captionStartId="45.[189,232,1919,1945]" captionTargetBox="[201,1377,1173,1856]" captionTargetId="figure@45.[189,1399,1153,1879]" captionTargetPageId="45" captionText="Fig. 30. Geographic distribution records of species of Umbyquyra gen. nov.: U. acuminata (Schmidt &amp; Tesmoingt in Schmidt, 2005) gen. et comb. nov., U. palmarum (Schiapelli &amp; Gerschman, 1945) gen. et comb. nov., U. araguaia gen. et sp. nov., U. paranaiba gen. et sp. nov. and U. sapezal gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3817972" httpUri="https://zenodo.org/record/3817972/files/figure.png" pageId="24" pageNumber="25">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>