226 lines
21 KiB
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226 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne & Haring, Elisabeth" docDate="2021" docId="E3DDE51075F55ADAA0EA09297D66A35E" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Siciliaria tiberii subsp. armettensis De Mattia & Reier & Haring 2021, ssp. nov." docType="treatment" docUuid="BF64153C-F65F-4EA2-8794-269213F5A345" docUuidSource="ZooBank" docVersion="4" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>De Mattia, Willy</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reier, Susanne</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Haring, Elisabeth</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1077</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>175</mods:end>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
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<mods:identifier type="ZooBank">C28AD65A76F242CFBED7DFB3702CABCE</mods:identifier>
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<mods:identifier type="Pensoft-UUID">734088641608531C8E2CC69397B000ED</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="191645968" LSID="urn:lsid:zoobank.org:act:BF64153C-F65F-4EA2-8794-269213F5A345" httpUri="http://treatment.plazi.org/id/E3DDE51075F55ADAA0EA09297D66A35E" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/BF64153C-F65F-4EA2-8794-269213F5A345" authority="De Mattia & Reier & Haring, 2021" authorityName="De Mattia & Reier & Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. armettensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" status="ssp. nov." subSpecies="armettensis">Siciliaria tiberii armettensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">ssp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wagner's (1913, 1925) and Brandt's (1961) Siciliaria genital drawings A Siciliaria septemplicata, Wagner, 1913: plate 572, fig. 14 B Siciliaria grohmanniana grohmanniana, Wagner, 1925: plate 1, fig. 8 C Siciliaria calcarae calcarae, 1925: plate 3, fig. 25 D Siciliaria ferrox, Brandt, 1961: fig. 1 E overall Siciliaria / Sicania and Sicilian Mauritanica distribution F Siciliaria / Sicania and Sicilian Mauritanica distribution of taxa." figureDoi="10.3897/zookeys.1077.67081.figure1" httpUri="https://binary.pensoft.net/fig/622793" pageId="0" pageNumber="1">Figs 1.F, 27.1-27.3, 28.1-28.3, 29.6-29.8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Italy, Sicily, Carini, Grotta dei Puntali o
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<normalizedToken originalValue="dell’Armetta">dell'Armetta</normalizedToken>
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, 90 m asl,
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<geoCoordinate degrees="38" direction="north" minutes="8" orientation="latitude" precision="1" seconds="58.56" value="38.1496">38°8'58.56"N</geoCoordinate>
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,
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<geoCoordinate degrees="13" direction="east" minutes="9" orientation="longitude" precision="1" seconds="25.83" value="13.157175">13°9'25.83"E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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1
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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(NHMW 113621) [Lab ID 38_1, COI: MW758880, ITS2: MW757098, MW757099, MW757100] and 4
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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(NHMW 113622): Italy, Sicily, Carini, Grotta dei Puntali o
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<normalizedToken originalValue="dell’Armetta">dell'Armetta</normalizedToken>
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, 90 m asl,
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<geoCoordinate degrees="38" direction="north" minutes="08" orientation="latitude" precision="1" seconds="58.56" value="38.1496">38°08'58.56"N</geoCoordinate>
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,
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<geoCoordinate degrees="13" direction="east" minutes="09" orientation="longitude" precision="1" seconds="25.83" value="13.157175">13°09'25.83"E</geoCoordinate>
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, [Lab ID 38_2, COI: MW758881], W. De Mattia and J. Macor leg., 16.iv.2017. 3 dissected spm. 8
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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(CMWDM 18227): same locality.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="shell diagnosis">
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<paragraph pageId="0" pageNumber="1">Shell diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell decollate; whorls moderately ribbed; dorsal keel prominent; inferior lamella high; anterior upper palatal plica present, knob present at distal end of the upper palatal plica, detached from the lunella; lower anterior palatal plica (ALPP) present; palatal edge of clausilium plate distally not receding, palatal edge distally strongly bent upwards.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Shell description</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Figs
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Siciliaria tiberii armettensis ssp. nov., Grotta dei Puntali, Carini 28.1 shell 28.2 shell clausiliar plate double side 28.3 detail of the aperture 28.4 clausiliar plate double side. Siciliaria tiberii scalettensis Beckmann, 2004 Portella Scaletta, Villagrazia 28.5 shell 28.6 detail of the aperture 28.7 shell 28.8 detail of the aperture 28.9 clausiliar plate double side." figureDoi="10.3897/zookeys.1077.67081.figure28" httpUri="https://binary.pensoft.net/fig/622820" pageId="0" pageNumber="1">28</figureCitation>
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.1-28.3, 29.6-29.8).
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</emphasis>
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The shell is elongated, fusiform, sinistral, decollate and robust. It is brown to dark reddish in colour. The shell surface has small ribs, more densely packed towards the decollate tip and gradually spreading towards the aperture. There are whitish papillae along the moderately deep sutures throughout the whole teleoconch. The spire is slowly and regularly growing, with 7
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<normalizedToken originalValue="½">1/2</normalizedToken>
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(after decollation) slightly convex whorls. The basal and cervical keels are well distinct. The umbilicus is closed. The aperture is roundish to subovoid and its height is ~ 1/4 to 1/5 of shell height. The PRI is short and it ends at the level of the L. It is not fused with the L and presents a slight thickening along its posterior end. The PRI is not visible from a frontal view of the aperture. The L is dorso-lateral to lateral. The PUPP is short or obsolete, knob-like and connected to the L. The AUPP is strong and long, detached from the L and clearly visible from the aperture. The distal end of the AUPP has a well-developed knob. The BAS starts directly from the L and it is long and strong, well visible from the aperture. The SCL is absent or resembling an irregular small thickening, connected with the lunella. The IL is very high to extremely high. The SUL is tooth-like, long and overlapping with the SPL. The SPL is long and it ends at 4/5 of the last whorl. The SCOL is not visible. The peristome is continuous, markedly thickened and reflected. It is not superiorly fused to the wall of the first whorl. The palatal edge of the clausilium is distally receding and bent upwards. The plate is cylindrical in shape. The palatal edge against the distal end is bent upwards and more or less blunt.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: decollate shell height 20.1, whorl width 4.8, aperture height 4.6, aperture width 3.1.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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(n = 35, decollate): shell height 19.7
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<normalizedToken originalValue="±">+/-</normalizedToken>
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1.6, whorl width 4.6
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.2, aperture height 4.2
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.3, aperture width 2.8
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<normalizedToken originalValue="±">+/-</normalizedToken>
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0.3.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs">
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<paragraph pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Fig.
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Siciliaria tiberii armettensis ssp. nov., Grotta dei Puntali, Carini 27.1 whole distal genital organs 27.2 internal distal part of genital organs 27.3 detail of the pseudopapilla. Siciliaria tiberii scalettensis Beckmann, 2004 Portella Scaletta, Villagrazia 27.4 whole distal genital organs 27.5 internal distal part of genital organs 27.6 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure27" httpUri="https://binary.pensoft.net/fig/622819" pageId="0" pageNumber="1">27</figureCitation>
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.1).
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</emphasis>
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The FO is longer than the V (FO/V range 1.5-1.8). The VD is thin along its whole course. The FDBC is slightly longer than the BC+SDBC (FDBC/BC+SDBC range 1.0-1.3). The BC+SDBC is irregularly spindle-like and longer than the V (BC+SDBC/V range 1.6-1.9), with no distinction between the SDBC and the BC. The apex is pointed. The D is longer than the V (D/V range 3.0-3.2) and longer that the BC+SDBC (D/BC+SDBC range 2.1-2.3), much thinner than the BC+SDBC and with a small but round apex. The V is short, cylindrical and large in diameter. The A is large. The PC is much longer than the V (P+E/V range 2.6-2.8). The PR is short and robust. The ET is not clearly visible. The E is longer than the P (E/P range 1.7-1.9), gradually shrinking and turning into the VD.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs">
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<paragraph pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(Figs
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Siciliaria tiberii armettensis ssp. nov., Grotta dei Puntali, Carini 27.1 whole distal genital organs 27.2 internal distal part of genital organs 27.3 detail of the pseudopapilla. Siciliaria tiberii scalettensis Beckmann, 2004 Portella Scaletta, Villagrazia 27.4 whole distal genital organs 27.5 internal distal part of genital organs 27.6 detail of the pseudopapilla." figureDoi="10.3897/zookeys.1077.67081.figure27" httpUri="https://binary.pensoft.net/fig/622819" pageId="0" pageNumber="1">27</figureCitation>
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.2, 27.3).
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</emphasis>
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The A shows a set of irregular fleshy folds. The P presents a main longitudinal fleshy fold that goes from the PP as far as the A. This fold is heavily segmented in an oblique way. Additionally, the P also shows a set of small longitudinal and irregular small pleats that reach the A. The fine structure of the wall is smooth. The PP is small, elongated and smooth. The P-E transition presents a first distal and a second median ER while both the PP and ELP originates from a third proximal ER. The epiphallar formula is: 1ER+2ER+3ER(PP+ELP). The E shows five to seven smooth longitudinal pleats that proximally turn into a pattern of coarse small weak pleats that gradually fade away towards the VD. The V shows a set of heavy and robust transverse smooth pleats that proximally begin somehow irregularly. The wall is smooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="comparative analysis">
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<paragraph pageId="0" pageNumber="1">Comparative analysis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. armettensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="armettensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii armettensis</emphasis>
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</taxonomicName>
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ssp. nov., like the nominate subspecies, has a ribbed shell and it is clearly distinguishable from all the remaining subspecies that present a more or less striated but ribless shell surface. Nonetheless, the nominate subspecies shows a stronger ribbing, with elevated and robust ribs. The distal end of the AUPP always presents a knob that is well visible from the frontal view (Figs
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Siciliaria tiberii armettensis ssp. nov., Grotta dei Puntali, Carini 28.1 shell 28.2 shell clausiliar plate double side 28.3 detail of the aperture 28.4 clausiliar plate double side. Siciliaria tiberii scalettensis Beckmann, 2004 Portella Scaletta, Villagrazia 28.5 shell 28.6 detail of the aperture 28.7 shell 28.8 detail of the aperture 28.9 clausiliar plate double side." figureDoi="10.3897/zookeys.1077.67081.figure28" httpUri="https://binary.pensoft.net/fig/622820" pageId="0" pageNumber="1">28</figureCitation>
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.1-28.2). This feature is shared also by the nominate subspecies (Fig.
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<figureCitation captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Siciliaria tiberii tiberii (A. Schmidt, 1868), Capo Rama, Terrasini 26.1 shell 26.2 clausiliar plate double side 26.3 shell 26.4 detail of the aperture 26.5 clausiliar plate double side. Siciliaria tiberii alcamoensis Brandt, 1961 comb. nov., Piano Margi, Carini 26.6 shell 26.7 clausiliar plate double side 26.8 detail of the aperture 26.9 shell." figureDoi="10.3897/zookeys.1077.67081.figure26" httpUri="https://binary.pensoft.net/fig/622818" pageId="0" pageNumber="1">26</figureCitation>
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.1) but only by the ≈25% of the investigated specimens (n = 40).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="ecology">
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<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. armettensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="armettensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii armettensis</emphasis>
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</taxonomicName>
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ssp. nov. inhabits east-exposed limestone walls and cliffs, hiding among tuft of vegetation, rocks cracks and crevices.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. armettensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="armettensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii armettensis</emphasis>
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</taxonomicName>
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ssp. nov. is exclusively known from the type locality, the surroundings of the Grotta dei Puntali o
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<normalizedToken originalValue="dell’Armetta">dell'Armetta</normalizedToken>
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. Further field investigation along the Monte Pecoraro eastern slopes is needed in order to determine the actual distribution of the taxon. Grotta dei Puntali o
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<normalizedToken originalValue="dell’Armetta">dell'Armetta</normalizedToken>
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are included in the protected area Riserva Naturale Grotta dei Puntali.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The taxon is named after the Grotte
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<normalizedToken originalValue="dell’Armetta">dell'Armetta</normalizedToken>
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, a complex of natural caves where wall engravings and stone handcrafted tools, dated from the Paleolithic to the Bronze Age, were discovered (
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<bibRefCitation author="Mannino, G" journalOrPublisher="Sicilia Archeologica" pageId="0" pageNumber="1" pagination="73 - 83" refId="B44" refString="Mannino, G, 1978. Le grotte di Armetta (Carini). Sicilia Archeologica 11 (38): 73 - 83" title="Le grotte di Armetta (Carini)." volume="11" year="1978">Mannino 1978</bibRefCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1077.67081.figure27" httpUri="https://binary.pensoft.net/fig/622819" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
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<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. armettensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="armettensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii armettensis</emphasis>
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</taxonomicName>
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ssp. nov., Grotta dei Puntali, Carini
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<emphasis bold="true" pageId="0" pageNumber="1">27.1</emphasis>
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whole distal genital organs
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<emphasis bold="true" pageId="0" pageNumber="1">27.2</emphasis>
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internal distal part of genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">27.3</emphasis>
|
||
detail of the pseudopapilla.
|
||
<taxonomicName authorityName="Beckmann" authorityYear="2004" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria tiberii subsp. scalettensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="tiberii" subSpecies="scalettensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria tiberii scalettensis</emphasis>
|
||
</taxonomicName>
|
||
Beckmann, 2004 Portella Scaletta, Villagrazia
|
||
<emphasis bold="true" pageId="0" pageNumber="1">27.4</emphasis>
|
||
whole distal genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">27.5</emphasis>
|
||
internal distal part of genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">27.6</emphasis>
|
||
detail of the pseudopapilla.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |