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<mods:title id="64D19BEB0430615AED4AB643AD32D539">The IndoWest Pacific Pilumnidae XVIII: On the taxonomy of Pilumnus scabriusculus Adams and White, 1849, and P. s l u i t e r i De Man, 1892, with a note on Cancer incanus Forskål, 1775 (Brachyura: Xanthoidea)</mods:title>
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<mods:namePart id="C62F9AA24AAEBD66F8E1C2A31126A320">Ng, Peter K. L.</mods:namePart>
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<heading id="4D67347D3B07B554A019F9C2FB97F95E" bold="true" box="[264,1105,1638,1664]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="24E45F033B07B554A019F9C2FB97F95E" bold="true" box="[264,1105,1638,1664]" pageId="4" pageNumber="5">
<taxonomicName id="D190F8923B07B554A019F9C2FD48F95E" ID-CoL="4HVX5" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[264,654,1638,1664]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="sluiteri">
<emphasis id="24E45F033B07B554A019F9C2FE1FF95E" bold="true" box="[264,473,1638,1664]" italics="true" pageId="4" pageNumber="5">Pilumnus sluiteri</emphasis>
<bibRefCitation id="7201FEE03B07B554A0F0F9C2FD48F95E" author="De" box="[481,654,1638,1664]" pageId="4" pageNumber="12" refString="De Man, J. G. (1892) Decapoden des Indischen Archipels. In: Weber, M. (Ed.) Zoologische Ergebnisse einer reise in Niederl: ndisch Ost-Indien, 2, 265 - 527, pls. 15 - 30." type="journal article" year="1892">De Man, 1892</bibRefCitation>
</taxonomicName>
(
<figureCitation id="8EAB9F943B07B554A38FF9C2FD36F95E" box="[670,752,1638,1664]" captionStart="FIGURE 1" captionStartId="2.[264,357,1825,1846]" captionTargetBox="[385,1201,405,1812]" captionTargetId="figure@2.[385,1203,405,1812]" captionTargetPageId="2" captionText="FIGURE 1. Overall dorsal view. A, Pilumnus scabriusculus Adams &amp; White, 1849, holotype male, 26.0 x 19.4 mm, NHM 1843.6; B, P. scabriusculus Adams &amp; White, 1849, male, 27.3 x 21.5 mm, NHM 1884.31; C, P. s l u i t e r i De Man, 1892, lectotype male, 27.2 x 20.5 mm, ZMA 242226." httpUri="https://zenodo.org/record/170700/files/figure.png" pageId="4" pageNumber="5">Figs. 1</figureCitation>
C, 2B, 3C, D, 4B, 5C, 6C, D)
</emphasis>
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<paragraph id="162F83113B07B554A019F915FBBDF916" blockId="4.[264,1325,1713,1896]" box="[264,1147,1713,1736]" pageId="4" pageNumber="5">
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<taxonomicName id="D190F8923B07B554A019F915FD62F916" ID-CoL="8YYB8" authority="De Man, 1888: 295" authorityName="De Man" authorityPageNumber="295" authorityYear="1888" box="[264,676,1713,1736]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="forskalii">
<emphasis id="24E45F033B07B554A019F915FE0AF916" box="[264,460,1713,1736]" italics="true" pageId="4" pageNumber="5">Pilumnus forskalii</emphasis>
<treatmentCitation id="9731A5003B07B554A0C2F915FD62F916" author="De" box="[467,676,1713,1736]" page="295" pageId="4" pageNumber="5" year="1888">
<bibRefCitation id="7201FEE03B07B554A0C2F915FDB6F916" author="De" box="[467,624,1713,1736]" pageId="4" pageNumber="12" refString="De Man, J. G. (1888) Bericht uber die von Herrn Dr. J. Brock im indischen Archipel gesammelten Decapoden und Stomatopoden. Archiv fur Naturgeschichte, Berlin, 53 (1), 215 - 600, pls. 7 - 22, 22 a [for publication date, see Clark et al. 1990]." type="journal article" year="1888">De Man, 1888</bibRefCitation>
: 295
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, pl. 12 fig. 1 (not H.
<bibRefCitation id="7201FEE03B07B554A295F915FBA9F916" author="Milne" box="[900,1135,1713,1736]" pageId="4" pageNumber="13" refString="Milne Edwards, H. (1834) Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, I, xxxv + 468 pp., tabs. 3. Librairie de Roret, Paris." type="book" year="1834">Milne Edwards, 1834</bibRefCitation>
).
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<emphasis id="24E45F033B07B554A019F975FE79F936" box="[264,447,1745,1768]" italics="true" pageId="4" pageNumber="5">Pilumnus sluiteri</emphasis>
<treatmentCitation id="9731A5003B07B554A0D7F975FD51F936" author="De" box="[454,663,1745,1768]" page="283" pageId="4" pageNumber="5" year="1892">
<bibRefCitation id="7201FEE03B07B554A0D7F975FDA5F936" author="De" box="[454,611,1745,1768]" pageId="4" pageNumber="12" refString="De Man, J. G. (1892) Decapoden des Indischen Archipels. In: Weber, M. (Ed.) Zoologische Ergebnisse einer reise in Niederl: ndisch Ost-Indien, 2, 265 - 527, pls. 15 - 30." type="journal article" year="1892">De Man, 1892</bibRefCitation>
: 283
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, pl. 15 fig. 2; 1929: 1;
<treatmentCitation id="9731A5003B07B554A29AF975FB95F936" author="Alcock" box="[907,1107,1745,1768]" page="195" pageId="4" pageNumber="5" year="1898">
<bibRefCitation id="7201FEE03B07B554A29AF975FBD8F936" author="Alcock" box="[907,1054,1745,1768]" pageId="4" pageNumber="12" refString="Alcock, A. (1898) Materials for a Carcinological Fauna of India. No. 3. The Brachyura Cyclometopa. Part I. The Family Xanthidae. Journal of the Asiatic Society of Bengal, Calcutta, 67 (2), No. 1, 67 - 233." type="journal article" year="1898">Alcock, 1898</bibRefCitation>
: 195
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.
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</paragraph>
<paragraph id="162F83113B07B554A019F955FB76F8B6" blockId="4.[264,1325,1713,1896]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="3680A43F3B07B554A019F955FB76F8B6" pageId="4" pageNumber="5">
<taxonomicName id="D190F8923B07B554A019F955FD39F8D6" ID-CoL="77JND" authority="Lanchester, 1901: 541" authorityName="Lanchester" authorityPageNumber="541" authorityYear="1901" box="[264,767,1777,1800]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="scabriusculus">
<emphasis id="24E45F033B07B554A019F955FDC1F8D6" box="[264,519,1777,1800]" italics="true" pageId="4" pageNumber="5">Pilumnus scabriusculus</emphasis>
<treatmentCitation id="9731A5003B07B554A31FF955FD39F8D6" author="Lanchester" box="[526,767,1777,1800]" page="541" pageId="4" pageNumber="5" year="1901">
<bibRefCitation id="7201FEE03B07B554A31FF955FD0FF8D6" author="Lanchester" box="[526,713,1777,1800]" pageId="4" pageNumber="12" refString="Lanchester, W. F. (1901) The Crustacea of the &quot; Skeat &quot; Expedition to the Malay Peninsula, together with a note on the genus Actaeopsis. Part 1, Brachyura, Stomatopoda and Macrura. Proceedings of the Zoological Society of London, 1901, 534 - 574, pls. 33, 34." type="proceedings paper" year="1901">Lanchester, 1901</bibRefCitation>
: 541
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</taxonomicName>
;
<treatmentCitation id="9731A5003B07B554A21AF955FC77F8D6" author="Balss" box="[779,945,1777,1800]" page="24" pageId="4" pageNumber="5" year="1933">
<bibRefCitation id="7201FEE03B07B554A21AF955FC4DF8D6" author="Balss" box="[779,907,1777,1800]" pageId="4" pageNumber="12" refString="Balss, H. (1933) Beitrage zur Kenntnis der Gattungen Pilumnus (Crustacea Dekapoda) und verwandter Gattungen. Capita Zoologica, 4 (3), 1 - 47, pls. 1 - 7." type="journal article" year="1933">Balss, 1933</bibRefCitation>
: 24
</treatmentCitation>
(part); 1938: 56;
<treatmentCitation id="9731A5003B07B554A57EF955FAE1F8D6" author="Sakai" box="[1135,1319,1777,1800]" page="533" pageId="4" pageNumber="5" year="1939">
<bibRefCitation id="7201FEE03B07B554A57EF955FB37F8D6" author="Sakai" box="[1135,1265,1777,1800]" pageId="4" pageNumber="13" refString="Sakai, T. (1939) Brachygnatha, Brachyrhyncha. Studies on the Crabs of Japan 4, 365 - 741, 1 - 129, pls. 42 - 111, tab. 1." type="journal article" year="1939">Sakai, 1939</bibRefCitation>
: 533
</treatmentCitation>
, pl. 100 fig. 5; 1976: 486, text figure 259;
<treatmentCitation id="9731A5003B07B554A21AF8B5FB85F8F6" author="Takeda" box="[779,1091,1809,1832]" page="24" pageId="4" pageNumber="5" year="1968">
<bibRefCitation id="7201FEE03B07B554A21AF8B5FBDFF8F6" author="Takeda" box="[779,1049,1809,1832]" lastPageId="4" lastPageNumber="14" pageId="4" pageNumber="13" refString="Takeda, M. &amp; Miyake, S. (1968) Pilumnid crabs of the family Xanthidae from the West Pacific. I. Twenty-three species of the genus Pilumnus, with description of four new species. Occasional Papers of the Zoological Laboratory, Faculty of Agriculture, Kyushu University, 1 (1), 1 - 60, pls. 1 - 4." type="journal article" year="1968">Takeda &amp; Miyake, 1968</bibRefCitation>
: 24
</treatmentCitation>
, Fig. 10c, d;
<treatmentCitation id="9731A5003B07B554A5C8F8B5FE62F896" author="Takeda" page="187" pageId="4" pageNumber="5" year="1982">
<bibRefCitation id="7201FEE03B07B554A5C8F8B5FEB7F896" author="Takeda" pageId="4" pageNumber="13" refString="Takeda, M. (1982) Keys to the Japanese and foreign crustaceans fully illustrated in colors. Hokuryukan, Tokyo. vi + 285 pp., pp. 1 - 58 (keys)." type="book chapter" year="1982">Takeda, 1982</bibRefCitation>
: 187
</treatmentCitation>
(with figure);
<treatmentCitation id="9731A5003B07B554A352F895FCD4F896" author="Miyake" box="[579,786,1841,1864]" page="137" pageId="4" pageNumber="5" year="1983">
<bibRefCitation id="7201FEE03B07B554A352F895FD1AF896" author="Miyake" box="[579,732,1841,1864]" pageId="4" pageNumber="13" refString="Miyake, S. (1983) Japanese crustacean decapods and stomatopods in color. Vol. II. Brachyura (Crabs). Osaka: Hoikusha Publishing Co., vii + 277 pp." type="book" year="1983">Miyake, 1983</bibRefCitation>
: 137
</treatmentCitation>
, pl. 46 fig. 2;
<treatmentCitation id="9731A5003B07B554A2BAF895FB22F896" author="Nagai" box="[939,1252,1841,1864]" page="189" pageId="4" pageNumber="5" year="1988">
<bibRefCitation id="7201FEE03B07B554A2BAF895FB68F896" author="Nagai" box="[939,1198,1841,1864]" pageId="4" pageNumber="13" refString="Nagai, S. &amp; Nomura, K. (1988) Crustacea (Brachyura) [The guide book of marine organisms of Okinawa], 7, 1 - 250. Shinsei Tosho, Okinawa. [In Japanese]" type="journal article" year="1988">Nagai &amp; Nomura, 1988</bibRefCitation>
: 189
</treatmentCitation>
;
<treatmentCitation id="9731A5003B07B554A5E1F895FE1CF8B6" author="Naiyanetr" page="86" pageId="4" pageNumber="5" year="1998">
<bibRefCitation id="7201FEE03B07B554A5E1F895FE74F8B6" author="Naiyanetr" pageId="4" pageNumber="13" refString="Naiyanetr, P. (1998) Checklist of crustacean fauna in Thailand (Decapoda and Stomatopoda). OEPP Biodiversity Series, vol. 5. Office of Environment Policy and Planning, 161 pp." type="book" year="1998">Naiyanetr, 1998</bibRefCitation>
: 86
</treatmentCitation>
; Ng et al., 2001: 30;
<treatmentCitation id="9731A5003B07B554A3ABF8F5FC65F8B6" author="Minemizu" box="[698,931,1873,1896]" page="281" pageId="4" pageNumber="5" year="2000">
<bibRefCitation id="7201FEE03B07B554A3ABF8F5FCA8F8B6" author="Minemizu" box="[698,878,1873,1896]" pageId="4" pageNumber="13" refString="Minemizu, R. (2000) Marine decapod and stomatopod crustaceans mainly from Japan. Bun-Itsu Sougou Shuppan Ltd., Tokyo, 344 pp. [In Japanese]" type="book" year="2000">Minemizu, 2000</bibRefCitation>
: 281
</treatmentCitation>
; Ng &amp; Davie, 2002: 377.
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</paragraph>
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<paragraph id="162F83113B06B555A019FEB8FD0EFE70" blockId="5.[264,1325,284,430]" pageId="5" pageNumber="6">
<emphasis id="24E45F033B06B555A019FEB8FE2FFEEB" box="[264,489,284,309]" italics="true" pageId="5" pageNumber="6">Material examined</emphasis>
.
<typeStatus id="C92B3DB33B06B555A0EDFEB8FD80FEE8" box="[508,582,284,310]" pageId="5" pageNumber="6">Types</typeStatus>
.
<collectingCountry id="6E87C3813B06B555A345FEB8FD31FEE8" box="[596,759,284,310]" name="Indonesia" pageId="5" pageNumber="6">INDONESIA</collectingCountry>
,
<typeStatus id="C92B3DB33B06B555A217FEB8FCB7FEE8" box="[774,881,284,310]" pageId="5" pageNumber="6" type="lectotype">lectotype</typeStatus>
male (here designated), 27.2 x
<quantity id="D1682EF43B06B555A5E8FEB8FEFFFE80" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="5" pageNumber="6" unit="mm" value="20.5">20.5 mm</quantity>
,
<collectionCode id="70811BD43B06B555A057FEE0FE41FE80" box="[326,391,324,350]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="5" pageNumber="6">ZMA</collectionCode>
242226, Enkhuizen Island, near
<collectingCountry id="6E87C3813B06B555A202FEE0FCAAFE80" box="[787,876,324,350]" name="Netherlands" pageId="5" pageNumber="6">Batavia</collectingCountry>
(= Jakarta), Java,
<collectingCountry id="6E87C3813B06B555A55AFEE0FB06FE80" box="[1099,1216,324,350]" name="Indonesia" pageId="5" pageNumber="6">Indonesia</collectingCountry>
, coll. M. Weber, 18881889.
<typeStatus id="C92B3DB33B06B555A0F6FEC8FD40FE58" box="[487,646,364,390]" pageId="5" pageNumber="6" type="paralectotype">Paralectotype</typeStatus>
female, 26.1 x
<quantity id="D1682EF43B06B555A250FEC8FC68FE58" box="[833,942,364,390]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="5" pageNumber="6" unit="mm" value="19.8">19.8 mm</quantity>
,
<collectionCode id="70811BD43B06B555A2ABFEC8FC3DFE58" box="[954,1019,364,390]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="5" pageNumber="6">ZMA</collectionCode>
242209, off Bay of
<collectingCountry id="6E87C3813B06B555A5FCFEC8FEEAFE70" name="Netherlands" pageId="5" pageNumber="6">Batavia</collectingCountry>
(= Jakarta Bay), Java, coll. Sluiter.
</paragraph>
<caption id="42EFD3993B06B555A019F88AFBABF8BF" httpUri="https://zenodo.org/record/170703/files/figure.png" pageId="5" pageNumber="6" targetBox="[264,1323,491,1825]" targetPageId="5">
<paragraph id="162F83113B06B555A019F88AFBABF8BF" blockId="5.[264,1323,1838,1889]" pageId="5" pageNumber="6">
<emphasis id="24E45F033B06B555A019F88AFEB8F89D" bold="true" box="[264,382,1838,1859]" pageId="5" pageNumber="6">FIGURE 4.</emphasis>
Ventral view. A,
<taxonomicName id="D190F8923B06B555A337F88BFC2BF89A" authority="Adams &amp; White, 1849" authorityName="Adams &amp; White" authorityYear="1849" box="[550,1005,1839,1860]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="scabriusculus">
<emphasis id="24E45F033B06B555A337F88BFCCDF89A" box="[550,779,1839,1860]" italics="true" pageId="5" pageNumber="6">Pilumnus scabriusculus</emphasis>
Adams &amp; White, 1849
</taxonomicName>
, holotype male, 26.0 x 19.4 mm, NHM 1843.6; B,
<taxonomicName id="D190F8923B06B555A0A0F8E8FD58F8BF" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[433,670,1868,1889]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sluiteri">
<emphasis id="24E45F033B06B555A0A0F8E8FDCAF8BF" box="[433,524,1868,1889]" italics="true" pageId="5" pageNumber="6">P. sluiteri</emphasis>
De Man, 1892
</taxonomicName>
, lectotype male, 27.2 x 20.5 mm, ZMA 242226.
</paragraph>
</caption>
<caption id="42EFD3993B05B556A019F8B5FBFDF8BC" httpUri="https://zenodo.org/record/170704/files/figure.png" pageId="6" pageNumber="7" targetBox="[396,1192,290,1782]" targetPageId="6">
<paragraph id="162F83113B05B556A019F8B5FBFDF8BC" blockId="6.[264,1323,1809,1890]" pageId="6" pageNumber="7">
<emphasis id="24E45F033B05B556A019F8B5FEBBF8F8" bold="true" box="[264,381,1809,1830]" pageId="6" pageNumber="7">FIGURE 5.</emphasis>
Outer view of third maxilliped. A,
<taxonomicName id="D190F8923B05B556A3DEF8B6FB55F8F8" authority="Adams &amp; White, 1849" authorityName="Adams &amp; White" authorityYear="1849" box="[719,1171,1809,1831]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="scabriusculus">
<emphasis id="24E45F033B05B556A3DEF8B6FC75F8F9" box="[719,947,1810,1831]" italics="true" pageId="6" pageNumber="7">Pilumnus scabriusculus</emphasis>
Adams &amp; White, 1849
</taxonomicName>
, holotype male, 26.0 x 19.4 mm, NHM 1843.6; B,
<taxonomicName id="D190F8923B05B556A37AF894FC3EF89A" authority="Adams &amp; White, 1849" authorityName="Adams &amp; White" authorityYear="1849" box="[619,1016,1839,1861]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="scabriusculus">
<emphasis id="24E45F033B05B556A37AF894FCCFF89B" box="[619,777,1840,1861]" italics="true" pageId="6" pageNumber="7">P. scabriusculus</emphasis>
Adams &amp; White, 1849
</taxonomicName>
, male, 27.3 x 21.5 mm, NHM 1884.31; C,
<taxonomicName id="D190F8923B05B556A06EF8E9FDAAF8BC" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[383,620,1869,1890]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sluiteri">
<emphasis id="24E45F033B05B556A06EF8E9FE1CF8BC" box="[383,474,1869,1890]" italics="true" pageId="6" pageNumber="7">P. sluiteri</emphasis>
De Man, 1892
</taxonomicName>
, lectotype male, 27.2 x 20.5 mm, ZMA 242226.
</paragraph>
</caption>
<paragraph id="162F83113B04B557A029FEB8FAE4FC00" blockId="7.[264,1325,284,1870]" pageId="7" pageNumber="8">
Nontypes.
<collectingCountry id="6E87C3813B04B557A0D2FEB8FDA1FEE8" box="[451,615,284,310]" name="Indonesia" pageId="7" pageNumber="8">INDONESIA</collectingCountry>
:
<specimenCount id="009648983B04B557A365FEB8FD15FEE8" box="[628,723,284,310]" pageId="7" pageNumber="8" type="male">2 males</specimenCount>
, 32.2 x
<quantity id="D1682EF43B04B557A227FEB8FC65FEE8" box="[822,931,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="7" pageNumber="8" unit="mm" value="24.3">24.3 mm</quantity>
, 26.3 x
<quantity id="D1682EF43B04B557A517FEB8FBB2FEE8" box="[1030,1140,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="7" pageNumber="8" unit="mm" value="19.4">19.4 mm</quantity>
,
<collectionCode id="70811BD43B04B557A590FEB8FB04FEE8" box="[1153,1218,284,310]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="7" pageNumber="8">ZMA</collectionCode>
242221,
<collectingCountry id="6E87C3813B04B557A019FEE0FE8CFE80" box="[264,330,324,350]" name="Iceland" pageId="7" pageNumber="8">Pulau</collectingCountry>
Berhala, coll. van der Meer, 1928.
<collectingCountry id="6E87C3813B04B557A216FEE0FC62FE80" box="[775,932,324,350]" name="Malaysia" pageId="7" pageNumber="8">MALAYSIA</collectingCountry>
:
<specimenCount id="009648983B04B557A2A2FEE0FBCFFE80" box="[947,1033,324,350]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
, 37.5 x
<quantity id="D1682EF43B04B557A563FEE0FB27FE80" box="[1138,1249,324,350]" metricMagnitude="-2" metricUnit="m" metricValue="2.78" pageId="7" pageNumber="8" unit="mm" value="27.8">27.8 mm</quantity>
,
<collectionCode id="70811BD43B04B557A5E3FEE0FAEDFE80" box="[1266,1323,324,350]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1987.35, Kuala Menggatal, Kota Kinabalu, Sabah, coll. N. Lee,
<date id="622EA5D13B04B557A2EBFEC8FB4BFE58" box="[1018,1165,364,390]" pageId="7" pageNumber="8" value="1986-11-05">5 Nov. 1986</date>
;
<specimenCount id="009648983B04B557A588FEC8FB2DFE58" box="[1177,1259,364,390]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
, 35.8 x
<quantity id="D1682EF43B04B557A00FFE30FE4DFE70" box="[286,395,404,430]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="7" pageNumber="8" unit="mm" value="26.5">26.5 mm</quantity>
,
<collectionCode id="70811BD43B04B557A086FE30FE16FE70" box="[407,464,404,430]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1987.854, Kuala Menggatal, Kota Kinabalu, Sabah, coll. N. Lee,
<date id="622EA5D13B04B557A5CCFE30FE80FE08" pageId="7" pageNumber="8" value="1986-11-05">5 Nov. 1986</date>
;
<specimenCount id="009648983B04B557A045FE18FE61FE08" box="[340,423,444,470]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
, 18.6 x
<quantity id="D1682EF43B04B557A319FE18FDB2FE08" box="[520,628,444,470]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="7" pageNumber="8" unit="mm" value="13.4">13.4 mm</quantity>
,
<collectionCode id="70811BD43B04B557A393FE18FD7DFE08" box="[642,699,444,470]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1987.855, Kuala Menggatal, Kota Kinabalu, Sabah, coll. N. Lee,
<date id="622EA5D13B04B557A0B6FE40FDFAFE20" box="[423,572,484,510]" pageId="7" pageNumber="8" value="1986-12-05">5 Dec. 1986</date>
;
<specimenCount id="009648983B04B557A35BFE40FD58FE20" box="[586,670,484,510]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
, 32.8 x
<quantity id="D1682EF43B04B557A211FE40FCA8FE20" box="[768,878,484,510]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="7" pageNumber="8" unit="mm" value="23.5">23.5 mm</quantity>
,
<collectionCode id="70811BD43B04B557A26AFE40FC72FE20" box="[891,948,484,510]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1987.1193,
<collectingCountry id="6E87C3813B04B557A55BFE40FB4BFE20" box="[1098,1165,484,510]" name="Iceland" pageId="7" pageNumber="8">Pulau</collectingCountry>
Tiga, Sabah, coll. N. Lee,
<date id="622EA5D13B04B557A0B6FDA8FD8FFDF8" box="[423,585,524,550]" pageId="7" pageNumber="8" value="1987-04-22">22 Apr. 1987</date>
.
<collectingCountry id="6E87C3813B04B557A347FDA8FD39FDF8" box="[598,767,524,550]" name="Singapore" pageId="7" pageNumber="8">SINGAPORE</collectingCountry>
:
<specimenCount id="009648983B04B557A21DFDA8FCB1FDF8" box="[780,887,524,550]" pageId="7" pageNumber="8" type="female">1 female</specimenCount>
, 26.9 x
<quantity id="D1682EF43B04B557A2CBFDA8FB81FDF8" box="[986,1095,524,550]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="7" pageNumber="8" unit="mm" value="20.3">20.3 mm</quantity>
,
<collectionCode id="70811BD43B04B557A544FDA8FB48FDF8" box="[1109,1166,524,550]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1965.9.9.49,
<collectingCountry id="6E87C3813B04B557A019FD90FE8CFD90" box="[264,330,564,590]" name="Iceland" pageId="7" pageNumber="8">Pulau</collectingCountry>
Pawai, southern
<collectingCountry id="6E87C3813B04B557A309FD90FD54FD90" box="[536,658,564,590]" name="Singapore" pageId="7" pageNumber="8">Singapore</collectingCountry>
, coll. M.W.F. Tweedie,
<date id="622EA5D13B04B557A2BCFD90FBEDFD90" box="[941,1067,564,590]" pageId="7" pageNumber="8" value="1933-11">Nov. 1933</date>
;
<specimenCount id="009648983B04B557A529FD90FB50FD90" box="[1080,1174,564,590]" pageId="7" pageNumber="8" type="male">2 males</specimenCount>
, 45.0 x
<quantity id="D1682EF43B04B557A5E6FD90FEFFFDA8" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" pageId="7" pageNumber="8" unit="mm" value="34.3">34.3 mm</quantity>
, 21.4 x
<quantity id="D1682EF43B04B557A084FDF8FDC6FDA8" box="[405,512,604,630]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="7" pageNumber="8" unit="mm" value="16.3">16.3 mm</quantity>
,
<collectionCode id="70811BD43B04B557A31DFDF8FD83FDA8" box="[524,581,604,630]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1965.9.9.4647, Sultan Shoal, southern
<collectingCountry id="6E87C3813B04B557A502FDF8FB4BFDA8" box="[1043,1165,604,630]" name="Singapore" pageId="7" pageNumber="8">Singapore</collectingCountry>
, coll. M.W.F. Tweedie,
<date id="622EA5D13B04B557A06CFD20FE3AFD40" box="[381,508,644,670]" pageId="7" pageNumber="8" value="1933-12">Dec. 1933</date>
; 1 ovig. female, 26.3 x
<quantity id="D1682EF43B04B557A235FD20FC55FD40" box="[804,915,644,670]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" pageId="7" pageNumber="8" unit="mm" value="18.9">18.9 mm</quantity>
,
<collectionCode id="70811BD43B04B557A2B3FD20FC1DFD40" box="[930,987,644,670]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1970.1.5.12, Raffles Lighthouse, southern
<collectingCountry id="6E87C3813B04B557A0DBFD08FD82FD18" box="[458,580,684,710]" name="Singapore" pageId="7" pageNumber="8">Singapore</collectingCountry>
, coll. A. Monteiro,
<date id="622EA5D13B04B557A23EFD08FC59FD18" box="[815,927,684,710]" pageId="7" pageNumber="8" value="1937-07">Jul. 1937</date>
;
<specimenCount id="009648983B04B557A2BCFD08FBD1FD18" box="[941,1047,684,710]" pageId="7" pageNumber="8" type="female">1 female</specimenCount>
, 29.6 x
<quantity id="D1682EF43B04B557A566FD08FB22FD18" box="[1143,1252,684,710]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="7" pageNumber="8" unit="mm" value="23.9">23.9 mm</quantity>
,
<collectionCode id="70811BD43B04B557A5E0FD08FAECFD18" box="[1265,1322,684,710]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1965.9.9.48, Horsburg Lighthouse, South
<collectingCountry id="6E87C3813B04B557A3E9FD70FCF8FD30" box="[760,830,724,750]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
Sea, coll. A. Monteiro,
<date id="622EA5D13B04B557A54AFD70FB12FD30" box="[1115,1236,724,750]" pageId="7" pageNumber="8" value="1938-04">Apr. 1938</date>
. PHIL IPPINES:
<specimenCount id="009648983B04B557A095FD58FE1EFCC8" box="[388,472,764,790]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
, 21.4 x
<quantity id="D1682EF43B04B557A32BFD58FD61FCC8" box="[570,679,764,790]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="7" pageNumber="8" unit="mm" value="15.8">15.8 mm</quantity>
,
<collectionCode id="70811BD43B04B557A3A5FD58FD36FCC8" box="[692,752,764,790]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
, Alona Beach, Panglao, Bohol, coll. P.K.L. Ng,
<date id="622EA5D13B04B557A019FC80FEBDFCE0" box="[264,379,804,830]" pageId="7" pageNumber="8" value="2003-07">Jul. 2003</date>
; 1 exovig. female, 26.8 x
<quantity id="D1682EF43B04B557A3DEFC80FCF9FCE0" box="[719,831,804,830]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="7" pageNumber="8" unit="mm" value="18.8">18.8 mm</quantity>
,
<collectionCode id="70811BD43B04B557A25EFC80FC4EFCE0" box="[847,904,804,830]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
2003.281, Alona Beach, Panglao, Bohol, coll. P.K.L. Ng &amp; P.F. Clark,
<date id="622EA5D13B04B557A3D2FCE8FCF2FCB8" box="[707,820,844,870]" pageId="7" pageNumber="8" value="2003-07">Jul. 2003</date>
.
<collectingCountry id="6E87C3813B04B557A250FCE8FC1BFCB8" box="[833,989,844,870]" name="Thailand" pageId="7" pageNumber="8">THAILAND</collectingCountry>
:
<specimenCount id="009648983B04B557A2FBFCE8FBF8FCB8" box="[1002,1086,844,870]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
, 26.5 x
<quantity id="D1682EF43B04B557A5B1FCE8FAC8FCB8" box="[1184,1294,844,870]" metricMagnitude="-2" metricUnit="m" metricValue="1.98" pageId="7" pageNumber="8" unit="mm" value="19.8">19.8 mm</quantity>
, 1 ovig. female, 31.6 x
<quantity id="D1682EF43B04B557A317FCD0FDB2FC50" box="[518,628,884,910]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="7" pageNumber="8" unit="mm" value="23.7">23.7 mm</quantity>
,
<collectionCode id="70811BD43B04B557A392FCD0FD7AFC50" box="[643,700,884,910]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
1999.519, southern Phuket, western
<collectingCountry id="6E87C3813B04B557A56FFCD0FB2FFC50" box="[1150,1257,884,910]" name="Thailand" pageId="7" pageNumber="8">Thailand</collectingCountry>
, coll. P.K.L. Ng,
<date id="622EA5D13B04B557A08BFC38FDDBFC68" box="[410,541,924,950]" pageId="7" pageNumber="8" value="1998-12">Dec. 1998</date>
;
<specimenCount id="009648983B04B557A321FC38FD4FFC68" box="[560,649,924,950]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
, 29.5 x
<quantity id="D1682EF43B04B557A3EBFC38FCAAFC68" box="[762,876,924,950]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="7" pageNumber="8" unit="mm" value="22.1">22.1 mm</quantity>
,
<specimenCount id="009648983B04B557A26EFC38FC29FC68" box="[895,1007,924,950]" pageId="7" pageNumber="8" type="female">1 female</specimenCount>
, 28.8 x
<quantity id="D1682EF43B04B557A571FC38FB15FC68" box="[1120,1235,924,950]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="7" pageNumber="8" unit="mm" value="21.2">21.2 mm</quantity>
, NHM 2001.194041, Cape Sai Yok, Phuket, western
<collectingCountry id="6E87C3813B04B557A230FC60FC4AFC00" box="[801,908,964,990]" name="Thailand" pageId="7" pageNumber="8">Thailand</collectingCountry>
, coll. P. Clark et al., 36
<date id="622EA5D13B04B557A5B2FC60FAD8FC00" box="[1187,1310,964,990]" pageId="7" pageNumber="8" value="2000-05">May 2000</date>
.
</paragraph>
</subSubSection>
<subSubSection id="5E8AD09A3B04B558A029FC48FC88FC68" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="162F83113B04B557A029FC48FB6AF908" blockId="7.[264,1325,284,1870]" pageId="7" pageNumber="8">
<emphasis id="24E45F033B04B557A029FC48FE04FBDB" box="[312,450,1004,1029]" italics="true" pageId="7" pageNumber="8">Description</emphasis>
. Carapace broader than long; dorsal surface convex transversely and longitudinally, more so anteriorly; regions discernible; surface covered with small rounded granules, relatively denser anteriorly; bases of most granules surrounded with very short setae, most granules on anterior half of carapace with a long, simple seta each, those on posterior surface mostly without the long seta; setae do not obscure surface, setae on margins and below teeth longer, denser, obscuring surface. Frontal margin with 2 truncate lobes separated by deep cleft, margin with granular; lateral lobes triangular, distinct, separated from frontal lobes by deep Vshaped cleft. Subfrontal surface with uneven transverse row of small granules. Supraorbital margin with low tooth on inner margin, margin granulated, with 2 shallow clefts on outer half. Infraorbital margin granulated, inner part with triangular tooth. External orbital angle low, broadly triangular, with sharp basal granule; anterolateral margin with 3 triangular teeth, all spineor tubercletipped, third tooth smallest. Posterolateral margin gently convex to almost straight, sharply converging towards gently convex posterior carapace margin. Suborbital, subhepatic and pterygostomial regions with low granules. Posterior margin of epistome with median part acutely triangular, lateral parts prominently concave. Endostomial ridges strong, complete. Third maxillipeds relatively longer; ischium subrectangular with shallow submedian sulcus; merus almost square with dorsal margin gently concave, submedian part concave; exopod relatively stout, distal edge not reaching dorsal edge of merus, with long flagellum.
</paragraph>
<paragraph id="162F83113B04B558A029F940FC67FE20" blockId="7.[264,1325,284,1870]" lastBlockId="8.[264,1323,284,950]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">Chelipeds unequal; outer surfaces of chelae with numerous conical granules of varying sizes, those medially relatively lower, in larger specimens, median granules not discernible; with short setae between distinct granules that do not obscure surface; inner surface if chelae smooth. Fingers shorter than palm, cutting edges with distinct teeth. Inner distal angle of carpus with 1 low tubercle, inner ventral angle with a low tubercle, with low granule between them; outer surface covered with low rounded granules amongst short setae. Merus short, with distal angle of dorsal margin spiniform, subdistal part of dorsal margin expanded into projection with spiniform tip; remainder of dorsal margin granulated. Dorsal margin of basisischium with low granules.</paragraph>
<paragraph id="162F83113B0BB558A029FDA8FB41FD40" blockId="8.[264,1323,284,950]" pageId="8" pageNumber="9">Ambulatory legs relatively short, margins densely lined with setae; second leg longest. Carpus with distinct dorsal sulcus, dorsal margin entire. Merus of last leg short, stout, without subdistal dorsal angle; meri of first to third legs relatively more elongate, with distinct subdistal dorsal angle that is rounded, not spiniform; margins unarmed.</paragraph>
<paragraph id="162F83113B0BB558A029FD08FC88FC68" blockId="8.[264,1323,284,950]" pageId="8" pageNumber="9">Thoracic sternum with scattered setae, surface uneven but not prominently granulated. Sternites 1 and 2 separated by granulated suture; sternites 2 and 3 separated by distinct suture; sternites 3 and 4 effectively fused, suture not discernible but sternites separated by deep furrow. Male abdomen narrowly triangular longitudinally with all segments freely mobile; telson triangular with gently concave lateral margins; segments 3–6 progressively less trapezoidal, lateral margins slightly sinuous to almost straight. G1 prominently sinuous, distal part gently curved; tip relatively sharp.</paragraph>
</subSubSection>
</treatment>
</document>