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<document id="31A0AA73E54EDCCDD332EA7D7DAD09C7" ID-CLB-Dataset="32249" ID-DOI="10.11646/zootaxa.4313.1.1" ID-GBIF-Dataset="d8cc2cd8-8410-49aa-a75d-7a41d9f52b26" ID-ISSN="1175-5326" ID-Zenodo-Dep="883161" ID-ZooBank="A5Fdfba5-F992-44A8-84C2-1756C943C19B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504182989959" checkinUser="plazi" docAuthor="Bush, Sarah E." docDate="2017" docId="832187E9FF10FF5EFF746548FB89F98F" docLanguage="en" docName="zootaxa.4313.1.1.pdf" docOrigin="Zootaxa 4313 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Harpactrox loeiensis Bush 2017, new species" docType="treatment" docVersion="17" lastPageNumber="158" masterDocId="7F18FF91FF8AFFC3FFE36200FFC9FFA3" masterDocTitle="Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key" masterLastPageNumber="443" masterPageNumber="1" pageNumber="155" updateTime="1698467717597" updateUser="ExternalLinkService">
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<mods:titleInfo id="E3F71625B88DC7CEF569678A01BC439B">
<mods:title id="5321AE1B1E0ADB77711F4A0A85F7DF5F">Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key</mods:title>
</mods:titleInfo>
<mods:name id="D6769563D15431D04930163207A1F95A" type="personal">
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<mods:namePart id="391D6BB6477D7E71B90CA7D9A251E9C4">Bush, Sarah E.</mods:namePart>
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<mods:title id="2A3050AE71E0169A73E28F89DA165D51">Zootaxa</mods:title>
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<mods:part id="AADC3ED85E434ECC64E6F258DC68CC9D">
<mods:date id="294899816FB82DC1FF8EBFBF04F4F7BF">2017</mods:date>
<mods:detail id="249E42D58B94CEE9FA95C6DB1FF7E7A6" type="pubDate">
<mods:number id="BE030E036A050B8BD745F1323ACC6E47">2017-08-31</mods:number>
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<mods:number id="530E9B89C0F3FE47F16CF855B8A0ADE4">4313</mods:number>
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<mods:classification id="B89520B01F8F689A544ED330201C356C">journal article</mods:classification>
<mods:identifier id="724DD7ADEDB9B0ED9638015CEFB3E016" type="CLB-Dataset">32249</mods:identifier>
<mods:identifier id="299DFE4C70D6AA8980B4D4A9F36981CF" type="DOI">10.11646/zootaxa.4313.1.1</mods:identifier>
<mods:identifier id="9767563C6BB6FCCECCCE7FFC3CFE2F2D" type="GBIF-Dataset">d8cc2cd8-8410-49aa-a75d-7a41d9f52b26</mods:identifier>
<mods:identifier id="FEC34455484DB9E74EC13D56A61E1684" type="ISSN">1175-5326</mods:identifier>
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<treatment id="832187E9FF10FF5EFF746548FB89F98F" ID-DOI="http://doi.org/10.5281/zenodo.5296963" ID-GBIF-Taxon="133100190" ID-Zenodo-Dep="5296963" LSID="urn:lsid:plazi:treatment:832187E9FF10FF5EFF746548FB89F98F" httpUri="http://treatment.plazi.org/id/832187E9FF10FF5EFF746548FB89F98F" lastPageId="157" lastPageNumber="158" pageId="154" pageNumber="155">
<subSubSection id="43926574FF10FF59FF746548FE8AF820" pageId="154" pageNumber="155" type="nomenclature">
<paragraph id="0B3736FFFF10FF59FF746548FCE0F8C1" blockId="154.[151,809,1864,1923]" box="[151,809,1864,1890]" pageId="154" pageNumber="155">
<heading id="507F8193FF10FF59FF746548FCE0F8C1" bold="true" box="[151,809,1864,1890]" fontSize="11" level="1" pageId="154" pageNumber="155" reason="1">
<emphasis id="39FCEAEDFF10FF59FF746548FCE0F8C1" bold="true" box="[151,809,1864,1890]" pageId="154" pageNumber="155">
<taxonomicName id="CC884D7CFF10FF59FF746548FD45F8C1" authority="Gustafsson &amp; Bush" authorityName="Bush" authorityYear="2017" box="[151,652,1864,1890]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="loeiensis" status="sp. nov.">
<emphasis id="39FCEAEDFF10FF59FF746548FE5BF8C1" bold="true" box="[151,402,1864,1890]" italics="true" pageId="154" pageNumber="155">Harpactrox loeiensis</emphasis>
Gustafsson &amp; Bush
</taxonomicName>
,
<taxonomicNameLabel id="22CF5796FF10FF59FD7A6548FCE0F8C1" box="[665,809,1864,1890]" pageId="154" pageNumber="155" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0B3736FFFF10FF59FF74656AFE8AF820" blockId="154.[151,809,1864,1923]" box="[151,323,1898,1923]" pageId="154" pageNumber="155">
(
<figureCitation id="93B32A7AFF10FF59FF7C656AFEF2F820" box="[159,315,1898,1923]" captionStart-0="FIGURES 246 – 247" captionStart-1="FIGURES 248 – 252" captionStartId-0="155.[151,264,1922,1944]" captionStartId-1="156.[151,264,1915,1937]" captionTargetBox-0="[199,1341,238,1835]" captionTargetBox-1="[213,1366,212,1871]" captionTargetId-0="figure@155.[164,1388,195,1874]" captionTargetId-1="figure@156.[192,1395,193,1894]" captionTargetPageId-0="155" captionTargetPageId-1="156" captionText-0="FIGURES 246 – 247. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 246, male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively. 247, female habitus, dorsal and ventral views." captionText-1="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri-0="https://zenodo.org/record/883292/files/figure.png" httpUri-1="https://zenodo.org/record/883294/files/figure.png" pageId="154" pageNumber="155">Figs 246–252</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43926574FF10FF59FF7465B2FBF6F844" pageId="154" pageNumber="155" type="materials_examined">
<paragraph id="0B3736FFFF10FF59FF7465B2FBF6F844" blockId="154.[151,1160,1970,2023]" pageId="154" pageNumber="155">
<materialsCitation id="BBE03CA2FF10FF59FF7465B2FBF6F844" ID-GBIF-Occurrence="1587279404" country="Thailand" county="Wang Saphung District" location="Krading Mountain" municipality="Srithan" pageId="154" pageNumber="155" specimenCount="2" stateProvince="Loei Province" typeStatus="holotype">
<emphasis id="39FCEAEDFF10FF59FF7465B2FECCF86B" bold="true" box="[151,261,1970,1992]" pageId="154" pageNumber="155">
<typeStatus id="D433885DFF10FF59FF7465B2FF05F86B" box="[151,204,1970,1992]" pageId="154" pageNumber="155">Type</typeStatus>
host.
</emphasis>
<taxonomicName id="CC884D7CFF10FF59FEEF65B3FC62F86B" authority="(Robinson &amp; Kloss, 1919)" authorityName="Bush" authorityYear="2017" baseAuthorityName="Robinson &amp; Kloss" baseAuthorityYear="1919" box="[268,939,1970,1992]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="154" pageNumber="155" phylum="Chordata" rank="subSpecies" species="erythrocephalus" subSpecies="annamensis">
<emphasis id="39FCEAEDFF10FF59FEEF65B3FD54F86B" box="[268,669,1970,1992]" italics="true" pageId="154" pageNumber="155">Harpactes erythrocephalus annamensis</emphasis>
(Robinson &amp; Kloss, 1919)
</taxonomicName>
—red-headed trogon.
<emphasis id="39FCEAEDFF10FF59FF7465D1FEECF844" bold="true" box="[151,293,2001,2023]" pageId="154" pageNumber="155">
<typeStatus id="D433885DFF10FF59FF7465D1FF05F844" box="[151,204,2001,2023]" pageId="154" pageNumber="155">Type</typeStatus>
locality.
</emphasis>
<location id="0E576024FF10FF59FECF65D1FE23F844" LSID="urn:lsid:plazi:treatment:832187E9FF10FF5EFF746548FB89F98F:0E576024FF10FF59FECF65D1FE23F844" box="[300,490,2001,2023]" country="Thailand" county="Wang Saphung District" municipality="Srithan" name="Krading Mountain" pageId="154" pageNumber="155" stateProvince="Loei Province">Krading Mountain</location>
,
<collectingMunicipality id="EB53AC85FF10FF59FE1765D1FD89F844" box="[500,576,2001,2023]" pageId="154" pageNumber="155">Srithan</collectingMunicipality>
,
<collectingCounty id="E2564E73FF10FF59FDA965D2FCF0F844" box="[586,825,2001,2023]" pageId="154" pageNumber="155">Wang Saphung District</collectingCounty>
,
<collectingRegion id="C94CF81DFF10FF59FCA165D1FC1DF844" box="[834,980,2001,2023]" country="Thailand" name="Loei" pageId="154" pageNumber="155">Loei Province</collectingRegion>
,
<collectingCountry id="739F766FFF10FF59FC3D65D1FBF2F844" box="[990,1083,2001,2023]" name="Thailand" pageId="154" pageNumber="155">Thailand</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="5FF76677FF11FF58FF746582FDFFF850" httpUri="https://zenodo.org/record/883292/files/figure.png" pageId="155" pageNumber="156" targetBox="[199,1341,238,1835]" targetPageId="155">
<paragraph id="0B3736FFFF11FF58FF746582FDFFF850" blockId="155.[151,1436,1922,2035]" pageId="155" pageNumber="156">
<emphasis id="39FCEAEDFF11FF58FF746582FEA6F83B" bold="true" box="[151,367,1922,1944]" pageId="155" pageNumber="156">FIGURES 246–247.</emphasis>
<taxonomicName id="CC884D7CFF11FF58FE9A6583FD98F83B" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[377,593,1922,1944]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="loeiensis" status="gen. nov.">
<emphasis id="39FCEAEDFF11FF58FE9A6583FD98F83B" box="[377,593,1922,1944]" italics="true" pageId="155" pageNumber="156">Harpactrox loeiensis</emphasis>
</taxonomicName>
<emphasis id="39FCEAEDFF11FF58FDB96582FCC1F83B" bold="true" box="[602,776,1922,1944]" pageId="155" pageNumber="156">
<taxonomicNameLabel id="22CF5796FF11FF58FDB96582FD6CF83B" box="[602,677,1922,1944]" pageId="155" pageNumber="156" rank="genus">n. gen.</taxonomicNameLabel>
&amp; n. sp.
</emphasis>
ex
<taxonomicName id="CC884D7CFF11FF58FCD76583FB02F83B" authorityName="Bush" authorityYear="2017" baseAuthorityName="Robinson &amp; Kloss" baseAuthorityYear="1919" box="[820,1227,1922,1944]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="155" pageNumber="156" phylum="Chordata" rank="subSpecies" species="erythrocephalus" subSpecies="annamensis">
<emphasis id="39FCEAEDFF11FF58FCD76583FB02F83B" box="[820,1227,1922,1944]" italics="true" pageId="155" pageNumber="156">Harpactes erythrocephalus annamensis</emphasis>
</taxonomicName>
:
<emphasis id="39FCEAEDFF11FF58FB3F6582FAC1F83B" bold="true" box="[1244,1288,1922,1944]" pageId="155" pageNumber="156">246,</emphasis>
male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively.
<emphasis id="39FCEAEDFF11FF58FA9365BFFA55F876" bold="true" box="[1392,1436,1983,2005]" pageId="155" pageNumber="156">247,</emphasis>
female habitus, dorsal and ventral views.
</paragraph>
</caption>
<caption id="5FF76677FF16FF5FFF74657BFD19F86C" httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="156" pageNumber="157" targetBox="[213,1366,212,1871]" targetPageId="156">
<paragraph id="0B3736FFFF16FF5FFF74657BFD19F86C" blockId="156.[151,1436,1915,1999]" pageId="156" pageNumber="157">
<emphasis id="39FCEAEDFF16FF5FFF74657BFEA5F832" bold="true" box="[151,364,1915,1937]" pageId="156" pageNumber="157">FIGURES 248–252.</emphasis>
<taxonomicName id="CC884D7CFF16FF5FFE97657CFD81F832" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[372,584,1915,1937]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="156" pageNumber="157" phylum="Arthropoda" rank="species" species="loeiensis" status="gen. nov.">
<emphasis id="39FCEAEDFF16FF5FFE97657CFD81F832" box="[372,584,1915,1937]" italics="true" pageId="156" pageNumber="157">Harpactrox loeiensis</emphasis>
</taxonomicName>
<emphasis id="39FCEAEDFF16FF5FFDAD657BFD38F832" bold="true" box="[590,753,1915,1937]" pageId="156" pageNumber="157">
<taxonomicNameLabel id="22CF5796FF16FF5FFDAD657BFD5FF832" box="[590,662,1915,1937]" pageId="156" pageNumber="157" rank="genus">n. gen.</taxonomicNameLabel>
&amp; n. sp.
</emphasis>
ex
<taxonomicName id="CC884D7CFF16FF5FFCF4657CFB61F832" authorityName="Bush" authorityYear="2017" baseAuthorityName="Robinson &amp; Kloss" baseAuthorityYear="1919" box="[791,1192,1915,1937]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="156" pageNumber="157" phylum="Chordata" rank="subSpecies" species="erythrocephalus" subSpecies="annamensis">
<emphasis id="39FCEAEDFF16FF5FFCF4657CFB61F832" box="[791,1192,1915,1937]" italics="true" pageId="156" pageNumber="157">Harpactes erythrocephalus annamensis</emphasis>
</taxonomicName>
:
<emphasis id="39FCEAEDFF16FF5FFB55657BFB2AF832" bold="true" box="[1206,1251,1915,1937]" pageId="156" pageNumber="157">248,</emphasis>
male head, dorsal and ventral views.
<emphasis id="39FCEAEDFF16FF5FFEBB659AFE4DF813" bold="true" box="[344,388,1946,1968]" pageId="156" pageNumber="157">249,</emphasis>
male genitalia, dorsal view.
<emphasis id="39FCEAEDFF16FF5FFD4A659AFD1FF813" bold="true" box="[681,726,1946,1968]" pageId="156" pageNumber="157">250,</emphasis>
male mesosome, ventral view.
<emphasis id="39FCEAEDFF16FF5FFBF4659AFB8AF813" bold="true" box="[1047,1091,1946,1968]" pageId="156" pageNumber="157">251,</emphasis>
male paramere, dorsal view.
<emphasis id="39FCEAEDFF16FF5FFA93659AFA55F813" bold="true" box="[1392,1436,1946,1968]" pageId="156" pageNumber="157">252,</emphasis>
female subgenital plate and vulval margin, ventral view.
</paragraph>
</caption>
<subSubSection id="43926574FF17FF5EFF746297FE76FD9E" pageId="157" pageNumber="158" type="diagnosis">
<paragraph id="0B3736FFFF17FF5EFF746297FE76FD9E" blockId="157.[151,1437,151,1581]" pageId="157" pageNumber="158">
<emphasis id="39FCEAEDFF17FF5EFF746297FED8FF13" bold="true" box="[151,273,151,176]" pageId="157" pageNumber="158">Diagnosis.</emphasis>
<taxonomicName id="CC884D7CFF17FF5EFEF96299FDCEFF13" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[282,519,151,176]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis" status="sp. nov.">
<emphasis id="39FCEAEDFF17FF5EFEF96299FDCEFF13" box="[282,519,151,176]" italics="true" pageId="157" pageNumber="158">Harpactrox loeiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF17FF5EFDF26297FD9BFF13" box="[529,594,151,176]" pageId="157" pageNumber="158" rank="species">
<emphasis id="39FCEAEDFF17FF5EFDF26297FD9BFF13" bold="true" box="[529,594,151,176]" pageId="157" pageNumber="158">n. sp.</emphasis>
</taxonomicNameLabel>
most similar to
<taxonomicName id="CC884D7CFF17FF5EFCF16299FC7FFF13" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[786,950,152,176]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="pontifrons" status="sp. nov.">
<emphasis id="39FCEAEDFF17FF5EFCF16299FCFDFF13" box="[786,820,153,176]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFCA76298FC7FFF13" box="[836,950,152,176]" italics="true" pageId="157" pageNumber="158">pontifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF17FF5EFC236297FBC8FF13" box="[960,1025,151,176]" pageId="157" pageNumber="158" rank="species">
<emphasis id="39FCEAEDFF17FF5EFC236297FBC8FF13" bold="true" box="[960,1025,151,176]" pageId="157" pageNumber="158">n. sp.</emphasis>
</taxonomicNameLabel>
, with which it shares the following characters: head broader than long in both species (
<figureCitation id="93B32A7AFF17FF5EFCE562BCFCBBFF77" box="[774,882,188,213]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Figs 248</figureCitation>
,
<figureCitation id="93B32A7AFF17FF5EFC6062BCFC67FF76" box="[899,942,188,213]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="157" pageNumber="158">260</figureCitation>
) (approximately as broad as long in
<taxonomicName id="CC884D7CFF17FF5EFA9062BDFED9FF5B" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="geminodus" status="sp. nov.">
<emphasis id="39FCEAEDFF17FF5EFA9062BDFA5CFF77" box="[1395,1429,189,212]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFF7462DFFED9FF5B" box="[151,272,223,248]" italics="true" pageId="157" pageNumber="158">geminodus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FF17FF5EFEF562DFFE9DFF5B" box="[278,340,223,248]" pageId="157" pageNumber="158" rank="species">
<emphasis id="39FCEAEDFF17FF5EFEF562DFFE9DFF5B" bold="true" box="[278,340,223,248]" pageId="157" pageNumber="158">n. sp.</emphasis>
</taxonomicNameLabel>
, Fig. 255); dorsal preantennal suture does not reach
<emphasis id="39FCEAEDFF17FF5EFC7862DFFC1AFF5B" box="[923,979,223,248]" italics="true" pageId="157" pageNumber="158">dsms</emphasis>
(suture reaches
<emphasis id="39FCEAEDFF17FF5EFB6B62DFFB09FF5B" box="[1160,1216,223,248]" italics="true" pageId="157" pageNumber="158">dsms</emphasis>
in
<taxonomicName id="CC884D7CFF17FF5EFB0162E1FA42FF5B" authorityName="Bush" authorityYear="2017" box="[1250,1419,223,248]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="geminodus">
<emphasis id="39FCEAEDFF17FF5EFB0162E1FACDFF5B" box="[1250,1284,225,248]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFAF162DFFA42FF5B" box="[1298,1419,223,248]" italics="true" pageId="157" pageNumber="158">geminodus</emphasis>
</taxonomicName>
);
<emphasis id="39FCEAEDFF17FF5EFF746304FF05FEBF" box="[151,204,260,284]" italics="true" pageId="157" pageNumber="158">
<taxonomicName id="CC884D7CFF17FF5EFF746304FF77FEBF" box="[151,190,260,284]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
the same length as
<emphasis id="39FCEAEDFF17FF5EFE456304FE31FEBF" box="[422,504,260,284]" italics="true" pageId="157" pageNumber="158">
<taxonomicName id="CC884D7CFF17FF5EFE456304FE07FEBF" box="[422,462,260,284]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
1–2
</emphasis>
and
<emphasis id="39FCEAEDFF17FF5EFDCE6304FDB7FEBF" box="[557,638,260,284]" italics="true" pageId="157" pageNumber="158">
<taxonomicName id="CC884D7CFF17FF5EFDCE6304FD9AFEBF" box="[557,595,260,284]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
4–5
</emphasis>
(
<emphasis id="39FCEAEDFF17FF5EFD6D6304FD0AFEBF" box="[654,707,260,284]" italics="true" pageId="157" pageNumber="158">
<taxonomicName id="CC884D7CFF17FF5EFD6D6304FD7CFEBF" box="[654,693,260,284]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
slightly longer than other
<taxonomicName id="CC884D7CFF17FF5EFC096305FBD8FEBF" box="[1002,1041,261,284]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF17FF5EFC096305FBD8FEBF" box="[1002,1041,261,284]" italics="true" pageId="157" pageNumber="158">mts</emphasis>
</taxonomicName>
in
<taxonomicName id="CC884D7CFF17FF5EFBD76305FB14FEBE" authorityName="Bush" authorityYear="2017" box="[1076,1245,260,285]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="geminodus">
<emphasis id="39FCEAEDFF17FF5EFBD76305FB9FFEBF" box="[1076,1110,261,284]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFB876304FB14FEBE" box="[1124,1245,260,285]" italics="true" pageId="157" pageNumber="158">geminodus</emphasis>
</taxonomicName>
n. sp). However,
<taxonomicName id="CC884D7CFF17FF5EFF746329FEE4FEE3" authorityName="Bush" authorityYear="2017" box="[151,301,295,320]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFF746329FF70FEE3" box="[151,185,297,320]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFF286327FEE4FEE3" box="[203,301,295,320]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF17FF5EFEA26327FE6DFEE3" box="[321,420,295,320]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 248</figureCitation>
) lacks the dorsal ridge at the osculum found in
<taxonomicName id="CC884D7CFF17FF5EFC016329FB40FEE3" authorityName="Bush" authorityYear="2017" box="[994,1161,296,320]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="pontifrons">
<emphasis id="39FCEAEDFF17FF5EFC016329FBCDFEE3" box="[994,1028,297,320]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFBF46328FB40FEE3" box="[1047,1161,296,320]" italics="true" pageId="157" pageNumber="158">pontifrons</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF17FF5EFB7F6327FACAFEE3" box="[1180,1283,295,320]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="157" pageNumber="158">Fig. 260</figureCitation>
).
<taxonomicName id="CC884D7CFF17FF5EFAF96329FEC0FEC7" authorityName="Bush" authorityYear="2017" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="pontifrons">
<emphasis id="39FCEAEDFF17FF5EFAF96329FEC0FEC7" italics="true" pageId="157" pageNumber="158">Harpactrox pontifrons</emphasis>
</taxonomicName>
has 2
<emphasis id="39FCEAEDFF17FF5EFEB7634DFE49FEC7" box="[340,384,333,356]" italics="true" pageId="157" pageNumber="158">vms</emphasis>
, 3
<emphasis id="39FCEAEDFF17FF5EFE46634DFE0EFEC7" box="[421,455,333,356]" italics="true" pageId="157" pageNumber="158">vss</emphasis>
, and 7–8
<emphasis id="39FCEAEDFF17FF5EFDDA634DFD97FEC7" box="[569,606,333,356]" italics="true" pageId="157" pageNumber="158">vos</emphasis>
(
<figureCitation id="93B32A7AFF17FF5EFD8C634CFD1BFEC6" box="[623,722,332,357]" captionStart="FIGURES 260 – 262" captionStartId="161.[151,264,1970,1992]" captionTargetBox="[194,1356,212,1926]" captionTargetId="figure@161.[163,1404,193,1927]" captionTargetPageId="161" captionText="FIGURES 260 – 262. Harpactrox pontifrons n. gen. &amp; n. sp. ex Harpactes ardens ardens: 260, female head, dorsal and ventral views. 261, female habitus, dorsal and ventral views. 262, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883298/files/figure.png" pageId="157" pageNumber="158">Fig. 262</figureCitation>
), whereas
<taxonomicName id="CC884D7CFF17FF5EFCA8634DFC17FEC6" authorityName="Bush" authorityYear="2017" box="[843,990,332,357]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFCA8634DFCA4FEC7" box="[843,877,333,356]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFC9F634CFC17FEC6" box="[892,990,332,357]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF17FF5EFC0D634CFB86FEC6" box="[1006,1103,332,357]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 252</figureCitation>
) has 4
<emphasis id="39FCEAEDFF17FF5EFB40634DFB06FEC7" box="[1187,1231,333,356]" italics="true" pageId="157" pageNumber="158">vms</emphasis>
, 5–6
<emphasis id="39FCEAEDFF17FF5EFAF3634DFAFBFEC7" box="[1296,1330,333,356]" italics="true" pageId="157" pageNumber="158">vss</emphasis>
, and 5–8
<emphasis id="39FCEAEDFF17FF5EFF746371FF75FE2B" box="[151,188,369,392]" italics="true" pageId="157" pageNumber="158">vos</emphasis>
. The ventral anterior plate of
<taxonomicName id="CC884D7CFF17FF5EFDF66371FD63FE2B" authorityName="Bush" authorityYear="2017" box="[533,682,367,392]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFDF66371FDFEFE2B" box="[533,567,369,392]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFDAB636FFD63FE2B" box="[584,682,367,392]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF17FF5EFD58636FFCD4FE2B" box="[699,797,367,392]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 248</figureCitation>
) is smaller than that of the other two species (Figs 255, 260). Males of
<taxonomicName id="CC884D7CFF17FF5EFEA66395FE11FE0E" authorityName="Bush" authorityYear="2017" box="[325,472,404,429]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFEA66395FEAEFE0F" box="[325,359,405,428]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFE956394FE11FE0E" box="[374,472,404,429]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF17FF5EFE0B6394FD4FFE0E" box="[488,646,404,429]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Figs 249–251</figureCitation>
) are separated from those of
<taxonomicName id="CC884D7CFF17FF5EFC306395FBB4FE0E" authorityName="Bush" authorityYear="2017" box="[979,1149,404,429]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="geminodus">
<emphasis id="39FCEAEDFF17FF5EFC306395FC3CFE0F" box="[979,1013,405,428]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFBE76394FBB4FE0E" box="[1028,1149,404,429]" italics="true" pageId="157" pageNumber="158">geminodus</emphasis>
</taxonomicName>
by the shape of the male genitalia(Figs 256–258), but males of
<taxonomicName id="CC884D7CFF17FF5EFDB763B9FD30FE73" authorityName="Bush" authorityYear="2017" box="[596,761,440,464]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="pontifrons">
<emphasis id="39FCEAEDFF17FF5EFDB763B9FDBFFE73" box="[596,630,441,464]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFD6463B8FD30FE73" box="[647,761,440,464]" italics="true" pageId="157" pageNumber="158">pontifrons</emphasis>
</taxonomicName>
are unknown. In
<taxonomicName id="CC884D7CFF17FF5EFC2F63B9FBABFE73" authorityName="Bush" authorityYear="2017" box="[972,1122,439,464]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFC2F63B9FC27FE73" box="[972,1006,441,464]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFBE363B7FBABFE73" box="[1024,1122,439,464]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
</taxonomicName>
the proximal mesosome is slender (
<figureCitation id="93B32A7AFF17FF5EFF1463DCFE93FE56" box="[247,346,476,501]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 250</figureCitation>
), with a narrowly rounded anterior margin, whereas in
<taxonomicName id="CC884D7CFF17FF5EFC2763DDFBA4FE56" authorityName="Bush" authorityYear="2017" box="[964,1133,476,501]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="geminodus">
<emphasis id="39FCEAEDFF17FF5EFC2763DDFC2FFE57" box="[964,998,477,500]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFC1763DCFBA4FE56" box="[1012,1133,476,501]" italics="true" pageId="157" pageNumber="158">geminodus</emphasis>
</taxonomicName>
it is broad (Fig. 257), with a largely flat anterior margin. In addition, the gonopore is extended distally in
<taxonomicName id="CC884D7CFF17FF5EFBF06001FB6FFDBB" authorityName="Bush" authorityYear="2017" box="[1043,1190,511,536]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFBF06001FBFCFDBB" box="[1043,1077,513,536]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFBA763FFFB6FFDBB" box="[1092,1190,511,536]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFF17FF5EFB5563FFFAD3FDBB" box="[1206,1306,511,536]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 250</figureCitation>
), but not in
<taxonomicName id="CC884D7CFF17FF5EFF746025FE89FD9E" authorityName="Bush" authorityYear="2017" box="[151,320,548,573]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="geminodus">
<emphasis id="39FCEAEDFF17FF5EFF746025FF70FD9F" box="[151,185,549,572]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFF246024FE89FD9E" box="[199,320,548,573]" italics="true" pageId="157" pageNumber="158">geminodus</emphasis>
</taxonomicName>
(Fig. 257).
</paragraph>
</subSubSection>
<subSubSection id="43926574FF17FF5EFF246047FED3FB4B" pageId="157" pageNumber="158" type="description">
<paragraph id="0B3736FFFF17FF5EFF246047FB78FD53" blockId="157.[151,1437,151,1581]" pageId="157" pageNumber="158">
<emphasis id="39FCEAEDFF17FF5EFF246047FE17FDFC" bold="true" box="[199,478,583,608]" pageId="157" pageNumber="158">
Description.
<emphasis id="39FCEAEDFF17FF5EFE826047FE17FDFC" bold="true" box="[353,478,583,608]" italics="true" pageId="157" pageNumber="158">Both sexes</emphasis>
</emphasis>
. Head shape, structure, and chaetotaxy as in
<figureCitation id="93B32A7AFF17FF5EFC016047FB8DFDC3" box="[994,1092,583,608]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 248</figureCitation>
. Ventral anterior plate slightly longer than marginal carina. Ventral carinae diffuse anterior to pulvinus. Dorsal preantennal suture encompasses
<emphasis id="39FCEAEDFF17FF5EFF74608FFF77FD0B" box="[151,190,655,680]" italics="true" pageId="157" pageNumber="158">ads</emphasis>
, and bends anteriorly at
<emphasis id="39FCEAEDFF17FF5EFE36608FFE32FD0B" box="[469,507,655,680]" italics="true" pageId="157" pageNumber="158">ads</emphasis>
, but does not reach
<emphasis id="39FCEAEDFF17FF5EFD02608FFCD1FD0B" box="[737,792,655,680]" italics="true" pageId="157" pageNumber="158">dsms</emphasis>
. Preantennal nodi large. All
<taxonomicName id="CC884D7CFF17FF5EFBBD6091FB4CFD0B" box="[1118,1157,657,680]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFF17FF5EFBBD6091FB4CFD0B" box="[1118,1157,657,680]" italics="true" pageId="157" pageNumber="158">mts</emphasis>
</taxonomicName>
of roughly equal length. Gular plate indistinct, but anterior end pointed. Thoracic and abdominal segments as in genus description and
<figureCitation id="93B32A7AFF17FF5EFA8F60B4FF35FD53" captionStart="FIGURES 246 – 247" captionStartId="155.[151,264,1922,1944]" captionTargetBox="[199,1341,238,1835]" captionTargetId="figure@155.[164,1388,195,1874]" captionTargetPageId="155" captionText="FIGURES 246 – 247. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 246, male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively. 247, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883292/files/figure.png" pageId="157" pageNumber="158">Figs 246–247</figureCitation>
. Pleurites broad anteriorly and irregular in male, more slender and regular in female.
</paragraph>
<paragraph id="0B3736FFFF17FF5EFF2460FCFED3FBB3" blockId="157.[151,1437,151,1581]" pageId="157" pageNumber="158">
<collectingRegion id="C94CF81DFF17FF5EFF2460FCFECBFCB6" box="[199,258,764,789]" country="Maldives" name="Maale" pageId="157" pageNumber="158">
<emphasis id="39FCEAEDFF17FF5EFF2460FCFECBFCB6" bold="true" box="[199,258,764,789]" italics="true" pageId="157" pageNumber="158">Male</emphasis>
</collectingRegion>
. Subgenital plate trapezoidal (
<figureCitation id="93B32A7AFF17FF5EFD9360FCFD10FCB6" box="[624,729,764,789]" captionStart="FIGURES 246 – 247" captionStartId="155.[151,264,1922,1944]" captionTargetBox="[199,1341,238,1835]" captionTargetId="figure@155.[164,1388,195,1874]" captionTargetPageId="155" captionText="FIGURES 246 – 247. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 246, male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively. 247, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883292/files/figure.png" pageId="157" pageNumber="158">Fig. 246</figureCitation>
). Abdominal chaetotaxy as in
<tableCitation id="460A0344FF17FF5EFBA860FCFB6DFCB7" box="[1099,1188,764,789]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="157" pageNumber="158">Table 2</tableCitation>
and
<figureCitation id="93B32A7AFF17FF5EFB0560FCFA84FCB6" box="[1254,1357,764,789]" captionStart="FIGURES 246 – 247" captionStartId="155.[151,264,1922,1944]" captionTargetBox="[199,1341,238,1835]" captionTargetId="figure@155.[164,1388,195,1874]" captionTargetPageId="155" captionText="FIGURES 246 – 247. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 246, male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively. 247, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883292/files/figure.png" pageId="157" pageNumber="158">Fig. 246</figureCitation>
. Basal apodeme (
<figureCitation id="93B32A7AFF17FF5EFEEC611FFEBBFC9B" box="[271,370,799,824]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 249</figureCitation>
) roughly as wide distally as proximally. Proximal mesosome (
<figureCitation id="93B32A7AFF17FF5EFBB3611FFB7DFC9B" box="[1104,1204,799,824]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 250</figureCitation>
) slender, elongated, anterior margin rounded, narrow. Gonopore narrowly open distally, elongated. Mesosomal lobes slight, rounded; 3
<emphasis id="39FCEAEDFF17FF5EFF746169FF18FC23" box="[151,209,873,896]" italics="true" pageId="157" pageNumber="158">ames</emphasis>
microsetae submedianly on each side anterior to gonopore; 2
<emphasis id="39FCEAEDFF17FF5EFC206169FC34FC23" box="[963,1021,873,896]" italics="true" pageId="157" pageNumber="158">pmes</emphasis>
microsetae on each side lateral to gonopore. Parameral heads (
<figureCitation id="93B32A7AFF17FF5EFE3D618CFD89FC06" box="[478,576,908,933]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 251</figureCitation>
) rounded. Parameral blades blunt, slender;
<emphasis id="39FCEAEDFF17FF5EFBDB618CFB4AFC07" box="[1080,1155,908,932]" italics="true" pageId="157" pageNumber="158">pst1–2</emphasis>
as in genus description. Measurements ex
<taxonomicName id="CC884D7CFF17FF5EFE8561B1FCECFC6B" authorityName="Bush" authorityYear="2017" baseAuthorityName="Robinson &amp; Kloss" baseAuthorityYear="1919" box="[358,805,943,968]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="157" pageNumber="158" phylum="Chordata" rank="subSpecies" species="erythrocephalus" subSpecies="annamensis">
<emphasis id="39FCEAEDFF17FF5EFE8561B1FCECFC6B" box="[358,805,943,968]" italics="true" pageId="157" pageNumber="158">Harpactes erythrocephalus annamensis</emphasis>
</taxonomicName>
(n = 12 except n = 10 for TL): TL = 1.77–2.06 (1.85); HL = 0.41–0.45 (0.43); HW = 0.52–0.55 (0.53); PRW = 0.28–0.32 (0.30); PTW = 0.45–0.48 (0.46); AW = 0.51– 0.72 (0.59).
</paragraph>
<paragraph id="0B3736FFFF17FF5EFF24661CFED3FB4B" blockId="157.[151,1437,151,1581]" pageId="157" pageNumber="158">
<emphasis id="39FCEAEDFF17FF5EFF24661CFED7FB96" bold="true" box="[199,286,1052,1077]" italics="true" pageId="157" pageNumber="158">Female</emphasis>
. Abdominal chaetotaxy as in
<tableCitation id="460A0344FF17FF5EFD99661CFD06FB97" box="[634,719,1052,1077]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="157" pageNumber="158">Table 2</tableCitation>
and
<figureCitation id="93B32A7AFF17FF5EFCE8661CFCA6FB97" box="[779,879,1052,1077]" captionStart="FIGURES 246 – 247" captionStartId="155.[151,264,1922,1944]" captionTargetBox="[199,1341,238,1835]" captionTargetId="figure@155.[164,1388,195,1874]" captionTargetPageId="155" captionText="FIGURES 246 – 247. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 246, male habitus, dorsal and ventral views; only examined male with distorted abdomen, and shape of abdomen here reproduced as closely as possible; anterior sternites and some ventral setae missing in only available male specimen, and here included tentatively. 247, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883292/files/figure.png" pageId="157" pageNumber="158">Fig. 247</figureCitation>
. Subgenital plate broadly trapezoidal (
<figureCitation id="93B32A7AFF17FF5EFAC8661CFA59FB96" box="[1323,1424,1052,1077]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 252</figureCitation>
), reaching vulval margin. Vulval margin (
<figureCitation id="93B32A7AFF17FF5EFDBF663FFD75FBFB" box="[604,700,1087,1112]" captionStart="FIGURES 248 – 252" captionStartId="156.[151,264,1915,1937]" captionTargetBox="[213,1366,212,1871]" captionTargetId="figure@156.[192,1395,193,1894]" captionTargetPageId="156" captionText="FIGURES 248 – 252. Harpactrox loeiensis n. gen. &amp; n. sp. ex Harpactes erythrocephalus annamensis: 248, male head, dorsal and ventral views. 249, male genitalia, dorsal view. 250, male mesosome, ventral view. 251, male paramere, dorsal view. 252, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883294/files/figure.png" pageId="157" pageNumber="158">Fig. 252</figureCitation>
) gently rounded, with 4 short, slender
<emphasis id="39FCEAEDFF17FF5EFB906641FB57FBFB" box="[1139,1182,1089,1112]" italics="true" pageId="157" pageNumber="158">vms</emphasis>
on each side, and 5–6 short, thorn-like
<emphasis id="39FCEAEDFF17FF5EFEB56665FEB1FBDF" box="[342,376,1125,1148]" italics="true" pageId="157" pageNumber="158">vss</emphasis>
on each side; 5–8
<emphasis id="39FCEAEDFF17FF5EFDB06665FDB1FBDF" box="[595,632,1125,1148]" italics="true" pageId="157" pageNumber="158">vos</emphasis>
on each side; distal 3
<emphasis id="39FCEAEDFF17FF5EFC9E6665FC6BFBDF" box="[893,930,1125,1148]" italics="true" pageId="157" pageNumber="158">vos</emphasis>
median to
<emphasis id="39FCEAEDFF17FF5EFBC66665FB8EFBDF" box="[1061,1095,1125,1148]" italics="true" pageId="157" pageNumber="158">vss</emphasis>
. Measurements ex
<taxonomicName id="CC884D7CFF17FF5EFAC46665FE11FB03" authorityName="Bush" authorityYear="2017" baseAuthorityName="Robinson &amp; Kloss" baseAuthorityYear="1919" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="157" pageNumber="158" phylum="Chordata" rank="subSpecies" species="erythrocephalus" subSpecies="annamensis">
<emphasis id="39FCEAEDFF17FF5EFAC46665FE11FB03" italics="true" pageId="157" pageNumber="158">Harpactes erythrocephalus annamensis</emphasis>
</taxonomicName>
(n = 15 except n = 14 for TL and PTW and n = 13 for AW): TL = 1.91–2.37 (2.09); HL = 0.46–0.51 (0.48); HW = 0.59–0.64 (0.60); PRW = 0.32–0.37 (0.34); PTW = 0.51–0.55 (0.52); AW = 0.60– 0.76 (0.68).
</paragraph>
</subSubSection>
<subSubSection id="43926574FF17FF5EFF2466F4FE40FA3E" pageId="157" pageNumber="158" type="discussion">
<paragraph id="0B3736FFFF17FF5EFF2466F4FE40FA3E" blockId="157.[151,1437,151,1581]" pageId="157" pageNumber="158">
<emphasis id="39FCEAEDFF17FF5EFF2466F4FEF2FAAE" bold="true" box="[199,315,1268,1293]" pageId="157" pageNumber="158">Remarks.</emphasis>
We have specimens examined from the host subspecies
<taxonomicName id="CC884D7CFF17FF5EFC0F66F5FA55FAAF" authority="(Gould, 1834)" baseAuthorityName="Gould" baseAuthorityYear="1834" box="[1004,1436,1268,1293]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="157" pageNumber="158" phylum="Chordata" rank="subSpecies" species="erythorcephalus" subSpecies="erythrocephalus">
<emphasis id="39FCEAEDFF17FF5EFC0F66F5FBC9FAAF" box="[1004,1024,1269,1292]" italics="true" pageId="157" pageNumber="158">H</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFBF766F5FBE9FAAF" box="[1044,1056,1269,1292]" italics="true" pageId="157" pageNumber="158">e</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFBD666F4FB23FAAE" box="[1077,1258,1268,1293]" italics="true" pageId="157" pageNumber="158">erythrocephalus</emphasis>
(Gould, 1834)
</taxonomicName>
collected in
<collectingCountry id="739F766FFF17FF5EFEC06719FE59FA92" box="[291,400,1305,1329]" name="Myanmar" pageId="157" pageNumber="158">Myanmar</collectingCountry>
which do not belong to
<taxonomicName id="CC884D7CFF17FF5EFD486719FC67FA93" authorityName="Bush" authorityYear="2017" box="[683,942,1303,1328]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="geminodus">
<emphasis id="39FCEAEDFF17FF5EFD486719FC67FA93" box="[683,942,1303,1328]" italics="true" pageId="157" pageNumber="158">Harpactrox geminodus</emphasis>
</taxonomicName>
, but are more similar in head shape to
<taxonomicName id="CC884D7CFF17FF5EFA906719FEF3FAF6" authority="They" authorityName="They" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFA906719FA5CFA93" box="[1395,1429,1305,1328]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFF74673CFF30FAF6" box="[151,249,1340,1365]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
They
</taxonomicName>
differ from
<taxonomicName id="CC884D7CFF17FF5EFE26673DFD91FAF6" box="[453,600,1340,1365]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFE26673DFE2EFAF7" box="[453,487,1341,1364]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFE15673CFD91FAF6" box="[502,600,1340,1365]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
</taxonomicName>
in the shape of the male genitalia and the extent of the dorsal preantennal suture. We presently cannot assess whether this material is conspecific with
<taxonomicName id="CC884D7CFF17FF5EFBEA6761FB56FADB" box="[1033,1183,1375,1400]" class="Insecta" family="Philopteridae" genus="Harpactrox" kingdom="Animalia" order="Phthiraptera" pageId="157" pageNumber="158" phylum="Arthropoda" rank="species" species="loeiensis">
<emphasis id="39FCEAEDFF17FF5EFBEA6761FBE2FADB" box="[1033,1067,1377,1400]" italics="true" pageId="157" pageNumber="158">Ha</emphasis>
.
<emphasis id="39FCEAEDFF17FF5EFBDF675FFB56FADB" box="[1084,1183,1375,1400]" italics="true" pageId="157" pageNumber="158">loeiensis</emphasis>
</taxonomicName>
or not, and leave this material undescribed.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF17FF5EFF2467A7FB1BFA63" box="[199,1234,1447,1473]" pageId="157" pageNumber="158" type="etymology">
<paragraph id="0B3736FFFF17FF5EFF2467A7FB1BFA63" blockId="157.[151,1437,151,1581]" box="[199,1234,1447,1473]" pageId="157" pageNumber="158">
<emphasis id="39FCEAEDFF17FF5EFF2467A7FE87FA63" bold="true" box="[199,334,1447,1472]" pageId="157" pageNumber="158">Etymology.</emphasis>
The species epithet is derived from the
<typeStatus id="D433885DFF17FF5EFCEE67A9FCF4FA62" box="[781,829,1449,1473]" pageId="157" pageNumber="158">type</typeStatus>
locality,
<collectingRegion id="C94CF81DFF17FF5EFC4667A7FB8AFA63" box="[933,1091,1447,1472]" country="Thailand" name="Loei" pageId="157" pageNumber="158">Loei Province</collectingRegion>
in
<collectingCountry id="739F766FFF17FF5EFB8567A7FB04FA63" box="[1126,1229,1447,1472]" name="Thailand" pageId="157" pageNumber="158">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="43926574FF17FF5EFF2467CCFB89F98F" pageId="157" pageNumber="158" type="materials_examined">
<paragraph id="0B3736FFFF17FF5EFF2467CCFB89F98F" blockId="157.[151,1437,151,1581]" pageId="157" pageNumber="158">
<materialsCitation id="BBE03CA2FF17FF5EFF2467CCFA51F9AB" ID-GBIF-Occurrence="1587279383" collectingDate="1954-01" collectionCode="OSUS" collectorName="R. E. Elbel &amp; B. LeKagul" country="Thailand" county="Wang Saphung District" location="Krading Mountain" municipality="Srithan" pageId="157" pageNumber="158" specimenCount="1" specimenCount-male="1" stateProvince="Loei Province" typeStatus="holotype">
<emphasis id="39FCEAEDFF17FF5EFF2467CCFC39FA46" bold="true" box="[199,1008,1484,1509]" pageId="157" pageNumber="158">
<typeStatus id="D433885DFF17FF5EFF2467CCFECBFA46" box="[199,258,1484,1509]" pageId="157" pageNumber="158">Type</typeStatus>
material. Ex
<taxonomicName id="CC884D7CFF17FF5EFE4567CCFCBBFA46" authorityName="Bush" authorityYear="2017" baseAuthorityName="Robinson &amp; Kloss" baseAuthorityYear="1919" box="[422,882,1484,1509]" class="Aves" family="Trogonidae" genus="Harpactes" kingdom="Animalia" order="Trogoniformes" pageId="157" pageNumber="158" phylum="Chordata" rank="subSpecies" species="erythrocephalus" subSpecies="annamensis">
<emphasis id="39FCEAEDFF17FF5EFE4567CCFCBBFA46" bold="true" box="[422,882,1484,1509]" italics="true" pageId="157" pageNumber="158">Harpactes erythrocephalus annamensis</emphasis>
</taxonomicName>
:
<typeStatus id="D433885DFF17FF5EFC6067CCFC39FA46" box="[899,1008,1484,1509]" pageId="157" pageNumber="158" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="1D8EFD76FF17FF5EFC1B67CDFBC6FA47" box="[1016,1039,1485,1508]" pageId="157" pageNumber="158" type="male">♂</specimenCount>
,
<location id="0E576024FF17FF5EFBFF67CCFB26FA46" LSID="urn:lsid:plazi:treatment:832187E9FF10FF5EFF746548FB89F98F:0E576024FF17FF5EFBFF67CCFB26FA46" box="[1052,1263,1484,1509]" country="Thailand" county="Wang Saphung District" municipality="Srithan" name="Krading Mountain" pageId="157" pageNumber="158" stateProvince="Loei Province">Krading Mountain</location>
,
<collectingMunicipality id="EB53AC85FF17FF5EFB1F67CCFA99FA46" box="[1276,1360,1484,1509]" pageId="157" pageNumber="158">Srithan</collectingMunicipality>
,
<collectingCounty id="E2564E73FF17FF5EFAB867CDFE90F9AB" pageId="157" pageNumber="158">Wang Saphung District</collectingCounty>
,
<collectingRegion id="C94CF81DFF17FF5EFE8567EFFDC3F9AB" box="[358,522,1519,1544]" country="Thailand" name="Loei" pageId="157" pageNumber="158">Loei Province</collectingRegion>
,
<collectingCountry id="739F766FFF17FF5EFDF467EFFDB4F9AB" box="[535,637,1519,1544]" name="Thailand" pageId="157" pageNumber="158">Thailand</collectingCountry>
, 1,
<collectingDate id="6F72E9D7FF17FF5EFD4A67F1FCD0F9AB" box="[681,793,1519,1544]" pageId="157" pageNumber="158" value="1954-01">
<date id="7F36103FFF17FF5EFD4A67F1FCD0F9AB" box="[681,793,1519,1544]" pageId="157" pageNumber="158" value="1954-01">Jan. 1954</date>
</collectingDate>
,
<collectorName id="A67D5329FF17FF5EFCC467F1FC54F9AB" box="[807,925,1519,1544]" pageId="157" pageNumber="158">R.E. Elbel</collectorName>
&amp;
<collectorName id="A67D5329FF17FF5EFC2667F1FB84F9AB" box="[965,1101,1519,1544]" pageId="157" pageNumber="158">B. LeKagul</collectorName>
, RE-3252, B-22640 (
<collectionCode id="6D99AE3AFF17FF5EFAA067F0FA59F9AB" box="[1347,1424,1520,1544]" country="USA" httpUri="http://grbio.org/cool/x70a-ez93" name="Oklahoma State University" pageId="157" pageNumber="158">OSUS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="BBE03CA2FF17FF5EFF746414FD6AF98F" ID-GBIF-Occurrence="1587279377" box="[151,675,1556,1581]" collectingDate="1954-01" collectionCode="OSUS" collectorName="R. E. Elbel &amp; B. LeKagul" country="Thailand" county="Wang Saphung District" location="Krading Mountain" municipality="Srithan" pageId="157" pageNumber="158" specimenCount="1" specimenCount-female="1" stateProvince="Loei Province" typeStatus="paratype">
<emphasis id="39FCEAEDFF17FF5EFF746414FED1F98E" bold="true" box="[151,280,1556,1581]" pageId="157" pageNumber="158">
<typeStatus id="D433885DFF17FF5EFF746414FEDAF98E" box="[151,275,1556,1581]" pageId="157" pageNumber="158" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="1D8EFD76FF17FF5EFEFD6414FE88F98F" box="[286,321,1556,1580]" pageId="157" pageNumber="158" type="female">1♀</specimenCount>
same data as holotype (
<collectionCode id="6D99AE3AFF17FF5EFDAD6414FD52F98F" box="[590,667,1556,1580]" country="USA" httpUri="http://grbio.org/cool/x70a-ez93" name="Oklahoma State University" pageId="157" pageNumber="158">OSUS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="BBE03CA2FF17FF5EFD4C6414FB89F98F" ID-GBIF-Occurrence="1587279376" box="[687,1088,1556,1581]" collectingDate="1954-01" collectionCode="OSUS" collectorName="R. E. Elbel &amp; B. LeKagul" country="Thailand" county="Wang Saphung District" location="Krading Mountain" municipality="Srithan" pageId="157" pageNumber="158" specimenCount="2" specimenCount-female="2" stateProvince="Loei Province" typeStatus="paratype">
<specimenCount id="1D8EFD76FF17FF5EFD4C6414FD1AF98F" box="[687,723,1556,1580]" pageId="157" pageNumber="158" type="female">2♀</specimenCount>
, same data as holotype (PIPeR).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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