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<mods:title id="50065858F3DB4658E50A1E5745E34F33">Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key</mods:title>
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<taxonomicName id="CC884D7CFE94FEDDFF746298FE09FF11" authority="Conci, 1941" authorityName="Conci" authorityYear="1941" box="[151,448,152,178]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFF746298FE09FF11" bold="true" box="[151,448,152,178]" pageId="286" pageNumber="287">
<emphasis id="39FCEAEDFE94FEDDFF746298FEEEFF11" bold="true" box="[151,295,152,178]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
Conci, 1941
</emphasis>
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</heading>
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<paragraph id="0B3736FFFE94FEDDFF7462E0FDEBFF57" blockId="286.[151,600,222,306]" box="[151,546,222,245]" pageId="286" pageNumber="287">
<treatmentCitationGroup id="2B9811D1FE94FEDDFF7462E0FDEBFF57" box="[151,546,222,245]" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFF7462E0FE61FF57" authority="Nitzsch, 1818: 291" authorityName="Nitzsch" authorityPageNumber="291" authorityYear="1818" box="[151,424,222,245]" class="Insecta" family="Philopteridae" genus="Nirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFF7462E0FF2BFF56" box="[151,226,224,245]" italics="true" pageId="286" pageNumber="287">Nirmus</emphasis>
<treatmentCitation id="8A2910EEFE94FEDDFF0B62DEFE61FF57" author="Nitzsch" box="[232,424,222,244]" page="291" pageId="286" pageNumber="287" year="1818">
<bibRefCitation id="6F194B0EFE94FEDDFF0B62DEFEB1FF57" author="Nitzsch" box="[232,376,222,244]" pageId="286" pageNumber="287" refString="Nitzsch, C. L. (1818) Die Familien und Gattungen der Theierinsekten (Insecta epizoica); als ein Prodromus einer Naturgeschichte derselben. E. F. Germar's Magazin der Entomologie, 3, 261 - 318." type="journal article" year="1818">Nitzsch, 1818</bibRefCitation>
: 291
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(
<emphasis id="39FCEAEDFE94FEDDFE5562E0FDDDFF56" box="[438,532,224,245]" italics="true" pageId="286" pageNumber="287">in partim</emphasis>
).
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</paragraph>
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<treatmentCitationGroup id="2B9811D1FE94FEDDFF7462FDFDEDFE91" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFF7462FDFE14FEB0" authority="Neumann, 1906: 60" authorityName="Neumann" authorityPageNumber="60" authorityYear="1906" box="[151,477,253,275]" class="Insecta" family="Philopteridae" genus="Degeeriella" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFF7462FDFEC6FEB0" box="[151,271,253,275]" italics="true" pageId="286" pageNumber="287">Degeeriella</emphasis>
<treatmentCitation id="8A2910EEFE94FEDDFEF762FEFE14FEB0" author="Neumann" box="[276,477,253,275]" page="60" pageId="286" pageNumber="287" year="1906">
<bibRefCitation id="6F194B0EFE94FEDDFEF762FEFE73FEB0" author="Neumann" box="[276,442,253,275]" pageId="286" pageNumber="287" refString="Neumann, L. G. (1906) Notes sur les Mallophages. Bulletin de Socitee Zoologique de France, 31, 54 - 60." type="journal article" year="1906">Neumann, 1906</bibRefCitation>
: 60
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</taxonomicName>
(
<emphasis id="39FCEAEDFE94FEDDFE0F62FEFD80FEB0" box="[492,585,254,275]" italics="true" pageId="286" pageNumber="287">in partim</emphasis>
). 
<taxonomicName id="CC884D7CFE94FEDDFF74631CFE60FE91" authority="Keler, 1936 a: 257" authorityName="Keler" authorityPageNumber="257" authorityYear="1936" box="[151,425,284,306]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFF74631CFF27FE91" box="[151,238,284,306]" italics="true" pageId="286" pageNumber="287">Brueelia</emphasis>
<treatmentCitation id="8A2910EEFE94FEDDFF17631CFE60FE91" author="Keler" box="[244,425,284,306]" page="257" pageId="286" pageNumber="287" year="1936">
<bibRefCitation id="6F194B0EFE94FEDDFF17631CFEB3FE91" author="Keler" box="[244,378,284,306]" pageId="286" pageNumber="287" refString="Keler, S. von (1936) Uber einige Mallophagen aus Rossitten. Arbeiten in morphologische und taxonomische Entomologie von Berlin-Dahlem, 3, 256 - 264." type="journal article" year="1936" yearSuffix="a">Kéler, 1936a</bibRefCitation>
: 257
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</taxonomicName>
(
<emphasis id="39FCEAEDFE94FEDDFE5B631DFDDCFE91" box="[440,533,285,306]" italics="true" pageId="286" pageNumber="287">in partim</emphasis>
).
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<emphasis id="39FCEAEDFE94FEDDFF746359FECCFECC" bold="true" box="[151,261,345,367]" pageId="286" pageNumber="287">Type host.</emphasis>
<taxonomicName id="CC884D7CFE94FEDDFEEF635AFD92FECC" authority="Denny, 1842: 52" authorityName="Denny" authorityPageNumber="52" authorityYear="1842" box="[268,603,345,367]" class="Insecta" family="Philopteridae" genus="Nirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="apiastri">
<emphasis id="39FCEAEDFE94FEDDFEEF635AFE62FECC" box="[268,427,346,367]" italics="true" pageId="286" pageNumber="287">Nirmus apiastri</emphasis>
<bibRefCitation id="6F194B0EFE94FEDDFE52635AFDFFFECC" author="Denny" box="[433,566,345,367]" pageId="286" pageNumber="287" refString="Denny, H. (1842) Monographia anoplurorum Britanniae. Henry G. Bohn, London. xxvi + 262 pp." type="book" year="1842">Denny, 1842</bibRefCitation>
: 52
</taxonomicName>
, by original designation.
</paragraph>
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<subSubSection id="43926574FE94FEDDFF74639CFB11FB27" pageId="286" pageNumber="287" type="diagnosis">
<paragraph id="0B3736FFFE94FEDDFF74639CFB11FB27" blockId="286.[151,1437,412,2021]" pageId="286" pageNumber="287">
<emphasis id="39FCEAEDFE94FEDDFF74639CFED8FE16" bold="true" box="[151,273,412,437]" pageId="286" pageNumber="287">Diagnosis.</emphasis>
<taxonomicName id="CC884D7CFE94FEDDFEF8639CFE68FE16" authorityName="Conci" authorityYear="1941" box="[283,417,412,437]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFEF8639CFE68FE16" box="[283,417,412,437]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
is not particularly similar to any other genus treated here. 
<taxonomicName id="CC884D7CFE94FEDDFBB1639CFB11FE16" authorityName="Conci" authorityYear="1941" box="[1106,1240,412,437]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFBB1639CFB11FE16" box="[1106,1240,412,437]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
was placed near 
<taxonomicName id="CC884D7CFE94FEDDFF7463C1FEE9FE7B" authorityName="Eichler" authorityYear="1940" box="[151,288,449,472]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFF7463C1FEE9FE7B" box="[151,288,449,472]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
and 
<taxonomicName id="CC884D7CFE94FEDDFE8263C1FDCFFE7B" authorityName="Mey &amp; Barker" authorityYear="2014" box="[353,518,449,472]" class="Insecta" family="Philopteridae" genus="Motmotnirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFE8263C1FDCFFE7B" box="[353,518,449,472]" italics="true" pageId="286" pageNumber="287">Motmotnirmus</emphasis>
</taxonomicName>
in the phylogeny of Bush 
<emphasis id="39FCEAEDFE94FEDDFCAD63C1FC4DFE7B" box="[846,900,447,472]" italics="true" pageId="286" pageNumber="287">et al</emphasis>
. (2016), but this placement was not strongly supported. Morphologically, there are great differences among these three genera in many character sets; indeed, the male genitalia of 
<taxonomicName id="CC884D7CFE94FEDDFE6C6007FDDCFD83" authorityName="Conci" authorityYear="1941" box="[399,533,519,544]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFE6C6007FDDCFD83" box="[399,533,519,544]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
and 
<taxonomicName id="CC884D7CFE94FEDDFDB26009FD12FD83" authorityName="Eichler" authorityYear="1940" box="[593,731,521,544]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFDB26009FD12FD83" box="[593,731,521,544]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
are so different that they are hard to compare. The marginal carina is interrupted submedianly in all three genera. However, the carina of 
<taxonomicName id="CC884D7CFE94FEDDFBFA602DFB77FDE7" authorityName="Mey &amp; Barker" authorityYear="2014" box="[1049,1214,557,580]" class="Insecta" family="Philopteridae" genus="Motmotnirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFBFA602DFB77FDE7" box="[1049,1214,557,580]" italics="true" pageId="286" pageNumber="287">Motmotnirmus</emphasis>
</taxonomicName>
is interrupted only submedianly (
<figureCitation id="93B32A7AFE94FEDDFED6604FFE51FDCB" box="[309,408,591,616]" captionStart="FIGURES 504 – 508" captionStartId="303.[151,264,1934,1956]" captionTargetBox="[188,1401,208,1870]" captionTargetId="figure@303.[181,1405,193,1894]" captionTargetPageId="303" captionText="FIGURES 504 – 508. Motmotnirmus marginellus (Nitzsch [in Giebel], 1866) ex Momotus momota momota: 504, male head, dorsal and ventral views. 505, male genitalia, dorsal view. 506, male mesosome, ventral view. 507, male paramere, dorsal view. 508, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883428/files/figure.png" pageId="286" pageNumber="287">Fig. 504</figureCitation>
), whereas the carina of 
<taxonomicName id="CC884D7CFE94FEDDFD436051FCE0FDCB" authorityName="Eichler" authorityYear="1940" box="[672,809,593,616]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFD436051FCE0FDCB" box="[672,809,593,616]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
is reduced to a small remnant near the preantennal nodi (
<figureCitation id="93B32A7AFE94FEDDFF7C6074FEC1FD2F" box="[159,264,628,653]" captionStart="FIGURES 494 – 496" captionStartId="295.[151,264,1970,1992]" captionTargetBox="[184,1431,230,1925]" captionTargetId="figure@295.[151,1436,193,1949]" captionTargetPageId="295" captionText="FIGURES 494 – 496. Meropoecus balisong n. sp. ex Merops americanus: 494, male head, dorsal and ventral views. 495, male genitalia, ventral view. 496, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883420/files/figure.png" pageId="286" pageNumber="287">Figs 494</figureCitation>
, 
<figureCitation id="93B32A7AFE94FEDDFEF66074FE88FD2F" box="[277,321,628,652]" captionStart="FIGURES 499 – 501" captionStartId="298.[151,264,1935,1957]" captionTargetBox="[152,1433,210,1906]" captionTargetId="figure@298.[151,1436,193,1913]" captionTargetPageId="298" captionText="FIGURES 499 – 501. Meropoecus bartlowi n. sp. ex Merops ornatus: 499, male head, dorsal and ventral views. 500, male genitalia, ventral view. 501, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883424/files/figure.png" pageId="286" pageNumber="287">499</figureCitation>
) and the marginal carina of 
<taxonomicName id="CC884D7CFE94FEDDFD6C6074FCDCFD2E" authorityName="Conci" authorityYear="1941" box="[655,789,628,653]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFD6C6074FCDCFD2E" box="[655,789,628,653]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
is interrupted submedianly and laterally, at least on the dorsal side (
<figureCitation id="93B32A7AFE94FEDDFEFE6097FEB6FD13" box="[285,383,663,688]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 481</figureCitation>
). In 
<taxonomicName id="CC884D7CFE94FEDDFE4C6099FD9DFD13" authorityName="Mey &amp; Barker" authorityYear="2014" box="[431,596,665,688]" class="Insecta" family="Philopteridae" genus="Motmotnirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFE4C6099FD9DFD13" box="[431,596,665,688]" italics="true" pageId="286" pageNumber="287">Motmotnirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFD806097FD08FD13" box="[611,705,663,688]" captionStart="FIGURES 504 – 508" captionStartId="303.[151,264,1934,1956]" captionTargetBox="[188,1401,208,1870]" captionTargetId="figure@303.[181,1405,193,1894]" captionTargetPageId="303" captionText="FIGURES 504 – 508. Motmotnirmus marginellus (Nitzsch [in Giebel], 1866) ex Momotus momota momota: 504, male head, dorsal and ventral views. 505, male genitalia, dorsal view. 506, male mesosome, ventral view. 507, male paramere, dorsal view. 508, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883428/files/figure.png" pageId="286" pageNumber="287">Fig. 504</figureCitation>
) and 
<taxonomicName id="CC884D7CFE94FEDDFD1C6097FC4CFD13" authorityName="Conci" authorityYear="1941" box="[767,901,663,688]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFD1C6097FC4CFD13" box="[767,901,663,688]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFC776097FC3BFD13" box="[916,1010,663,688]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 481</figureCitation>
) the dorsal preantennal suture reaches the 
<emphasis id="39FCEAEDFE94FEDDFF2760BCFF35FD76" box="[196,252,700,725]" italics="true" pageId="286" pageNumber="287">dsms</emphasis>
but not the 
<emphasis id="39FCEAEDFE94FEDDFE6C60BCFE7CFD76" box="[399,437,700,725]" italics="true" pageId="286" pageNumber="287">ads</emphasis>
, but in 
<taxonomicName id="CC884D7CFE94FEDDFDF460BDFD69FD77" authorityName="Eichler" authorityYear="1940" box="[535,672,701,724]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFDF460BDFD69FD77" box="[535,672,701,724]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
the suture reaches the 
<emphasis id="39FCEAEDFE94FEDDFC5160BCFC10FD76" box="[946,985,700,725]" italics="true" pageId="286" pageNumber="287">ads</emphasis>
as well (
<figureCitation id="93B32A7AFE94FEDDFBAB60BCFB78FD77" box="[1096,1201,700,725]" captionStart="FIGURES 494 – 496" captionStartId="295.[151,264,1970,1992]" captionTargetBox="[184,1431,230,1925]" captionTargetId="figure@295.[151,1436,193,1949]" captionTargetPageId="295" captionText="FIGURES 494 – 496. Meropoecus balisong n. sp. ex Merops americanus: 494, male head, dorsal and ventral views. 495, male genitalia, ventral view. 496, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883420/files/figure.png" pageId="286" pageNumber="287">Figs 494</figureCitation>
, 
<figureCitation id="93B32A7AFE94FEDDFB5C60BCFB24FD77" box="[1215,1261,700,724]" captionStart="FIGURES 499 – 501" captionStartId="298.[151,264,1935,1957]" captionTargetBox="[152,1433,210,1906]" captionTargetId="figure@298.[151,1436,193,1913]" captionTargetPageId="298" captionText="FIGURES 499 – 501. Meropoecus bartlowi n. sp. ex Merops ornatus: 499, male head, dorsal and ventral views. 500, male genitalia, ventral view. 501, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883424/files/figure.png" pageId="286" pageNumber="287">499</figureCitation>
). The 
<emphasis id="39FCEAEDFE94FEDDFAD960BCFAA6FD77" box="[1338,1391,700,724]" italics="true" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFAD960BCFAABFD77" box="[1338,1378,700,724]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
2
</emphasis>
are macrosetae in 
<taxonomicName id="CC884D7CFE94FEDDFED960E1FE16FD5B" authorityName="Mey &amp; Barker" authorityYear="2014" box="[314,479,737,760]" class="Insecta" family="Philopteridae" genus="Motmotnirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFED960E1FE16FD5B" box="[314,479,737,760]" italics="true" pageId="286" pageNumber="287">Motmotnirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFE0C60DFFD9BFD5B" box="[495,594,735,760]" captionStart="FIGURES 504 – 508" captionStartId="303.[151,264,1934,1956]" captionTargetBox="[188,1401,208,1870]" captionTargetId="figure@303.[181,1405,193,1894]" captionTargetPageId="303" captionText="FIGURES 504 – 508. Motmotnirmus marginellus (Nitzsch [in Giebel], 1866) ex Momotus momota momota: 504, male head, dorsal and ventral views. 505, male genitalia, dorsal view. 506, male mesosome, ventral view. 507, male paramere, dorsal view. 508, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883428/files/figure.png" pageId="286" pageNumber="287">Fig. 504</figureCitation>
), and 
<emphasis id="39FCEAEDFE94FEDDFD7560E0FD02FD5B" box="[662,715,736,760]" italics="true" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFD7560E0FD77FD5B" box="[662,702,736,760]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
2
</emphasis>
is longer than 
<emphasis id="39FCEAEDFE94FEDDFC9760E0FC60FD5B" box="[884,937,736,760]" italics="true" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFC9760E0FC57FD5B" box="[884,926,736,760]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
1
</emphasis>
and 
<emphasis id="39FCEAEDFE94FEDDFC0260E0FBFBFD5B" box="[993,1074,736,760]" italics="true" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFC0260E0FBCEFD5B" box="[993,1031,736,760]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
<quantity id="CC709B1AFE94FEDDFBE460E0FBFBFD5B" box="[1031,1074,736,760]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="286" pageNumber="287" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4–5</quantity>
</emphasis>
in 
<taxonomicName id="CC884D7CFE94FEDDFBB560E1FB16FD5B" authorityName="Eichler" authorityYear="1940" box="[1110,1247,737,760]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFBB560E1FB16FD5B" box="[1110,1247,737,760]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFB1360DFFA9FFD5B" box="[1264,1366,735,760]" captionStart="FIGURES 494 – 496" captionStartId="295.[151,264,1970,1992]" captionTargetBox="[184,1431,230,1925]" captionTargetId="figure@295.[151,1436,193,1949]" captionTargetPageId="295" captionText="FIGURES 494 – 496. Meropoecus balisong n. sp. ex Merops americanus: 494, male head, dorsal and ventral views. 495, male genitalia, ventral view. 496, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883420/files/figure.png" pageId="286" pageNumber="287">Figs 494</figureCitation>
, 
<figureCitation id="93B32A7AFE94FEDDFA8160E0FA59FD5B" box="[1378,1424,736,760]" captionStart="FIGURES 499 – 501" captionStartId="298.[151,264,1935,1957]" captionTargetBox="[152,1433,210,1906]" captionTargetId="figure@298.[151,1436,193,1913]" captionTargetPageId="298" captionText="FIGURES 499 – 501. Meropoecus bartlowi n. sp. ex Merops ornatus: 499, male head, dorsal and ventral views. 500, male genitalia, ventral view. 501, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883424/files/figure.png" pageId="286" pageNumber="287">499</figureCitation>
), but in 
<taxonomicName id="CC884D7CFE94FEDDFF3C6104FEACFCBE" authorityName="Conci" authorityYear="1941" box="[223,357,772,797]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFF3C6104FEACFCBE" box="[223,357,772,797]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
all 
<taxonomicName id="CC884D7CFE94FEDDFE726105FE71FCBF" box="[401,440,773,796]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">
<emphasis id="39FCEAEDFE94FEDDFE726105FE71FCBF" box="[401,440,773,796]" italics="true" pageId="286" pageNumber="287">mts</emphasis>
</taxonomicName>
except 
<emphasis id="39FCEAEDFE94FEDDFDF36104FD8CFCBF" box="[528,581,772,796]" italics="true" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFDF36104FDFEFCBF" box="[528,567,772,796]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
are of equal length (
<figureCitation id="93B32A7AFE94FEDDFCCC6104FC5BFCBF" box="[815,914,772,797]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 481</figureCitation>
). Gonopore is terminal in 
<taxonomicName id="CC884D7CFE94FEDDFB596105FA96FCBF" authorityName="Mey &amp; Barker" authorityYear="2014" box="[1210,1375,773,796]" class="Insecta" family="Philopteridae" genus="Motmotnirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFB596105FA96FCBF" box="[1210,1375,773,796]" italics="true" pageId="286" pageNumber="287">Motmotnirmus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFA8D6104FF0CFCE3" captionStart="FIGURES 504 – 508" captionStartId="303.[151,264,1934,1956]" captionTargetBox="[188,1401,208,1870]" captionTargetId="figure@303.[181,1405,193,1894]" captionTargetPageId="303" captionText="FIGURES 504 – 508. Motmotnirmus marginellus (Nitzsch [in Giebel], 1866) ex Momotus momota momota: 504, male head, dorsal and ventral views. 505, male genitalia, dorsal view. 506, male mesosome, ventral view. 507, male paramere, dorsal view. 508, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883428/files/figure.png" pageId="286" pageNumber="287">Fig. 506</figureCitation>
), but ventral in 
<taxonomicName id="CC884D7CFE94FEDDFE916127FE30FCE3" authorityName="Conci" authorityYear="1941" box="[370,505,807,832]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFE916127FE30FCE3" box="[370,505,807,832]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFDE46127FD12FCE3" box="[519,731,807,832]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="286" pageNumber="287">Figs 484, 487, 490</figureCitation>
), and other characters of the male genitalia are hard to compare between these three genera; the male genitalia of 
<taxonomicName id="CC884D7CFE94FEDDFD3E614DFCAEFCC7" authorityName="Eichler" authorityYear="1940" box="[733,871,845,868]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFD3E614DFCAEFCC7" box="[733,871,845,868]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFC9A614CFC2BFCC6" box="[889,994,844,869]" captionStart="FIGURES 494 – 496" captionStartId="295.[151,264,1970,1992]" captionTargetBox="[184,1431,230,1925]" captionTargetId="figure@295.[151,1436,193,1949]" captionTargetPageId="295" captionText="FIGURES 494 – 496. Meropoecus balisong n. sp. ex Merops americanus: 494, male head, dorsal and ventral views. 495, male genitalia, ventral view. 496, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883420/files/figure.png" pageId="286" pageNumber="287">Figs 495</figureCitation>
, 
<figureCitation id="93B32A7AFE94FEDDFC12614CFBD4FCC6" box="[1009,1053,844,869]" captionStart="FIGURES 499 – 501" captionStartId="298.[151,264,1935,1957]" captionTargetBox="[152,1433,210,1906]" captionTargetId="figure@298.[151,1436,193,1913]" captionTargetPageId="298" captionText="FIGURES 499 – 501. Meropoecus bartlowi n. sp. ex Merops ornatus: 499, male head, dorsal and ventral views. 500, male genitalia, ventral view. 501, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883424/files/figure.png" pageId="286" pageNumber="287">500</figureCitation>
) are unique within the 
<taxonomicName id="CC884D7CFE94FEDDFAD1614CFA5BFCC6" authorityName="Keler" authorityYear="1936" box="[1330,1426,844,869]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFAD1614CFA5BFCC6" box="[1330,1426,844,869]" italics="true" pageId="286" pageNumber="287">Brueelia</emphasis>
</taxonomicName>
- complex, and most of the terminology introduced here to describe the male genitalia of the 
<taxonomicName id="CC884D7CFE94FEDDFB44616FFACEFC2B" authorityName="Keler" authorityYear="1936" box="[1191,1287,879,904]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFB44616FFACEFC2B" box="[1191,1287,879,904]" italics="true" pageId="286" pageNumber="287">Brueelia</emphasis>
</taxonomicName>
-complex are largely inapplicable to 
<taxonomicName id="CC884D7CFE94FEDDFE436195FDE0FC0F" authorityName="Eichler" authorityYear="1940" box="[416,553,917,940]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFE436195FDE0FC0F" box="[416,553,917,940]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
. The female subgenital plate and vulval margin of 
<taxonomicName id="CC884D7CFE94FEDDFB9E6194FACAFC0E" authorityName="Conci" authorityYear="1941" box="[1149,1283,916,941]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFB9E6194FACAFC0E" box="[1149,1283,916,941]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFAF66194FAB1FC0F" box="[1301,1400,916,941]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 482</figureCitation>
) is structurally very similar to that of 
<taxonomicName id="CC884D7CFE94FEDDFDDA61B9FD0BFC73" authorityName="Eichler" authorityYear="1940" box="[569,706,953,976]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFDDA61B9FD0BFC73" box="[569,706,953,976]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFD3461B7FC8AFC73" box="[727,835,951,976]" captionStart="FIGURES 494 – 496" captionStartId="295.[151,264,1970,1992]" captionTargetBox="[184,1431,230,1925]" captionTargetId="figure@295.[151,1436,193,1949]" captionTargetPageId="295" captionText="FIGURES 494 – 496. Meropoecus balisong n. sp. ex Merops americanus: 494, male head, dorsal and ventral views. 495, male genitalia, ventral view. 496, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883420/files/figure.png" pageId="286" pageNumber="287">Figs 496</figureCitation>
, 
<figureCitation id="93B32A7AFE94FEDDFCB761B7FC4AFC73" box="[852,899,951,976]" captionStart="FIGURES 499 – 501" captionStartId="298.[151,264,1935,1957]" captionTargetBox="[152,1433,210,1906]" captionTargetId="figure@298.[151,1436,193,1913]" captionTargetPageId="298" captionText="FIGURES 499 – 501. Meropoecus bartlowi n. sp. ex Merops ornatus: 499, male head, dorsal and ventral views. 500, male genitalia, ventral view. 501, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883424/files/figure.png" pageId="286" pageNumber="287">501</figureCitation>
): both genera have broad lateral submarginal extensions of the subgenital plate, and in both genera, there is a distinct median extension of the subgenital plate, that may approach, but not reach, the vulval margin. Additionally, in both 
<taxonomicName id="CC884D7CFE94FEDDFBF96601FB6AFBBB" authorityName="Eichler" authorityYear="1940" box="[1050,1187,1025,1048]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFBF96601FB6AFBBB" box="[1050,1187,1025,1048]" italics="true" pageId="286" pageNumber="287">Meropoecus</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFB5B61FFFAEAFBBB" box="[1208,1315,1023,1048]" captionStart="FIGURES 494 – 496" captionStartId="295.[151,264,1970,1992]" captionTargetBox="[184,1431,230,1925]" captionTargetId="figure@295.[151,1436,193,1949]" captionTargetPageId="295" captionText="FIGURES 494 – 496. Meropoecus balisong n. sp. ex Merops americanus: 494, male head, dorsal and ventral views. 495, male genitalia, ventral view. 496, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883420/files/figure.png" pageId="286" pageNumber="287">Figs 496</figureCitation>
, 
<figureCitation id="93B32A7AFE94FEDDFAD761FFFA96FBBB" box="[1332,1375,1023,1048]" captionStart="FIGURES 499 – 501" captionStartId="298.[151,264,1935,1957]" captionTargetBox="[152,1433,210,1906]" captionTargetId="figure@298.[151,1436,193,1913]" captionTargetPageId="298" captionText="FIGURES 499 – 501. Meropoecus bartlowi n. sp. ex Merops ornatus: 499, male head, dorsal and ventral views. 500, male genitalia, ventral view. 501, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883424/files/figure.png" pageId="286" pageNumber="287">500</figureCitation>
) and 
<taxonomicName id="CC884D7CFE94FEDDFF746624FED4FB9E" authorityName="Conci" authorityYear="1941" box="[151,285,1060,1085]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE94FEDDFF746624FED4FB9E" box="[151,285,1060,1085]" italics="true" pageId="286" pageNumber="287">Meropsiella</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE94FEDDFECF6624FE47FB9F" box="[300,398,1060,1085]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 482</figureCitation>
), the 
<emphasis id="39FCEAEDFE94FEDDFE296625FE25FB9F" box="[458,492,1061,1084]" italics="true" pageId="286" pageNumber="287">vss</emphasis>
and 
<emphasis id="39FCEAEDFE94FEDDFDC06625FD86FB9F" box="[547,591,1061,1084]" italics="true" pageId="286" pageNumber="287">vms</emphasis>
are largely mixed, and do not form separate sets. All these characters are in stark contrast to 
<emphasis id="39FCEAEDFE94FEDDFEB36649FE31FBC3" box="[336,504,1097,1120]" italics="true" pageId="286" pageNumber="287">Motmornirmus</emphasis>
(
<figureCitation id="93B32A7AFE94FEDDFDE46647FDA0FBC3" box="[519,617,1095,1120]" captionStart="FIGURES 504 – 508" captionStartId="303.[151,264,1934,1956]" captionTargetBox="[188,1401,208,1870]" captionTargetId="figure@303.[181,1405,193,1894]" captionTargetPageId="303" captionText="FIGURES 504 – 508. Motmotnirmus marginellus (Nitzsch [in Giebel], 1866) ex Momotus momota momota: 504, male head, dorsal and ventral views. 505, male genitalia, dorsal view. 506, male mesosome, ventral view. 507, male paramere, dorsal view. 508, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883428/files/figure.png" pageId="286" pageNumber="287">Fig. 508</figureCitation>
), in which the subgenital plate does reach the vulval margin, but does not have lateral submarginal extensions, and in which the 
<emphasis id="39FCEAEDFE94FEDDFD16666DFCDEFB27" box="[757,791,1133,1156]" italics="true" pageId="286" pageNumber="287">vss</emphasis>
and 
<emphasis id="39FCEAEDFE94FEDDFCAD666DFCB3FB27" box="[846,890,1133,1156]" italics="true" pageId="286" pageNumber="287">vms</emphasis>
are separated into distinct sets.
</paragraph>
</subSubSection>
<subSubSection id="43926574FE94FEDCFF24668FFB9DFEBF" lastPageId="287" lastPageNumber="288" pageId="286" pageNumber="287" type="description">
<paragraph id="0B3736FFFE94FEDDFF24668FFCA7FAFE" blockId="286.[151,1437,412,2021]" pageId="286" pageNumber="287">
<emphasis id="39FCEAEDFE94FEDDFF24668FFE2EFB04" bold="true" box="[199,487,1167,1192]" pageId="286" pageNumber="287">
Description. 
<emphasis id="39FCEAEDFE94FEDDFE86668FFE2EFB04" bold="true" box="[357,487,1167,1192]" italics="true" pageId="286" pageNumber="287">Both sexes</emphasis>
</emphasis>
. Head trapezoidal (
<figureCitation id="93B32A7AFE94FEDDFD35668FFCF7FB0B" box="[726,830,1167,1192]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 481</figureCitation>
). Marginal carina interrupted submedianly, but not completely interrupted laterally. Ventral carina continuous with marginal carina. Frons hyaline and continuous with dorsal preantennal suture that does not reach 
<emphasis id="39FCEAEDFE94FEDDFD7D66D7FD0CFB53" box="[670,709,1239,1264]" italics="true" pageId="286" pageNumber="287">ads</emphasis>
. Dorsal anterior plate continuous with main head plate. Ventral anterior plate absent. Head setae as in 
<figureCitation id="93B32A7AFE94FEDDFDAC66FCFD79FAB7" box="[591,688,1276,1301]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 481</figureCitation>
; 
<emphasis id="39FCEAEDFE94FEDDFD5E66FCFD2AFAB6" box="[701,739,1276,1301]" italics="true" pageId="286" pageNumber="287">ads</emphasis>
very short, and may be absent in some species; 
<emphasis id="39FCEAEDFE94FEDDFAED66FCFAF2FAB6" box="[1294,1339,1276,1301]" italics="true" pageId="286" pageNumber="287">mds</emphasis>
and 
<emphasis id="39FCEAEDFE94FEDDFA9666FDFA55FAB7" box="[1397,1436,1277,1300]" italics="true" pageId="286" pageNumber="287">pns</emphasis>
absent. Antennae monomorphic. Temporal carinae not visible; 
<emphasis id="39FCEAEDFE94FEDDFC966720FC63FA9B" box="[885,938,1312,1336]" italics="true" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFC966720FC55FA9B" box="[885,924,1312,1336]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
only macrosetae. Temporal margin bulges distinctly between 
<emphasis id="39FCEAEDFE94FEDDFE896744FE69FAFF" box="[362,416,1348,1372]" italics="true" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFE896744FE58FAFF" box="[362,401,1348,1372]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
and 
<emphasis id="39FCEAEDFE94FEDDFE366744FDC3FAFF" box="[469,522,1348,1372]" italics="true" pageId="286" pageNumber="287">
<taxonomicName id="CC884D7CFE94FEDDFE366744FE35FAFF" box="[469,508,1348,1372]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
4
</emphasis>
. Gular plate roughly triangular.
</paragraph>
<paragraph id="0B3736FFFE94FEDDFF246767FC45F9B3" blockId="286.[151,1437,412,2021]" pageId="286" pageNumber="287">
Prothorax (
<figureCitation id="93B32A7AFE94FEDDFEAB6767FE21FA23" box="[328,488,1383,1408]" captionStart="FIGURES 479 – 480" captionStartId="289.[151,264,1975,1997]" captionTargetBox="[197,1378,195,1935]" captionTargetId="figure@289.[185,1385,193,1949]" captionTargetPageId="289" captionText="FIGURES 479 – 480. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 479, male habitus, dorsal and ventral views. 480, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883412/files/figure.png" pageId="286" pageNumber="287">Figs 479–480</figureCitation>
) rounded rectangular; 
<emphasis id="39FCEAEDFE94FEDDFD126769FCEAFA23" box="[753,803,1385,1408]" italics="true" pageId="286" pageNumber="287">ppss</emphasis>
absent or minute. Proepimera with hammer- or hookshaped median ends. Pterothorax rounded trapezoidal; lateral margins convex; posterior margin more or less flat (
<taxonomicName id="CC884D7CFE94FEDDFF4367B1FE0BFA6B" authority="Denny, 1842: 133" authorityName="Denny" authorityPageNumber="133" authorityYear="1842" box="[160,450,1456,1480]" class="Aves" family="Sturnidae" genus="Sturnus" kingdom="Animalia" order="Passeriformes" pageId="286" pageNumber="287" phylum="Chordata" rank="species" species="contra">
<emphasis id="39FCEAEDFE94FEDDFF4367B1FF20FA6B" box="[160,233,1457,1480]" italics="true" pageId="286" pageNumber="287">contra</emphasis>
<bibRefCitation id="6F194B0EFE94FEDDFF1167B0FE4EFA6B" author="Denny" box="[242,391,1456,1480]" pageId="286" pageNumber="287" refString="Denny, H. (1842) Monographia anoplurorum Britanniae. Henry G. Bohn, London. xxvi + 262 pp." type="book" year="1842">Denny, 1842</bibRefCitation>
: 133
</taxonomicName>
); 
<emphasis id="39FCEAEDFE94FEDDFE3B67B1FDC2FA6B" box="[472,523,1457,1480]" italics="true" pageId="286" pageNumber="287">mms</emphasis>
moderately separated medianly. Meso- and metasterna not fused, setae only on postero-lateral corners of metasternum. Metepisterna with hammer-shaped median ends. Leg chaetotaxy as in 
<figureCitation id="93B32A7AFE94FEDDFA9367D4FF7FF9B3" captionStart="FIGURES 25" captionStartId="14.[151,264,1012,1034]" captionTargetBox="[173,1428,205,974]" captionTargetId="figure@14.[151,1436,193,991]" captionTargetPageId="14" captionText="FIGURES 25. Dorso-ventral views of coxae I – III, trochanters I – III, and femora I – III, showing terminology of the leg setae used here. Other parts of the thorax removed for clarity. Setae situated marginally are illustrated and named on both dorsal (lefthand) and ventral (right-hand) sides. Terminology for individual seta as used in the text obtained by combining the leg segment (larger font) and the setal name (smaller font). Abbreviations: a 1 – 5, anterior setae 1 – 5; cI – III, coxae I – III; d 1 – 3, dorsal setae 1 – 3; dm 1 – 4, distal marginal setae 1 – 4; fI – III, femur I – III; p 1 – 3, posterior setae 1 – 3; s 1 – 6, sensilla 1 – 6; tI – III, trochanter I – III; v 1 – 4, ventral setae 1 – 4." httpUri="https://zenodo.org/record/883171/files/figure.png" pageId="286" pageNumber="287">Fig. 25</figureCitation>
, except 
<emphasis id="39FCEAEDFE94FEDDFEF267F8FDE5F9B3" box="[273,556,1528,1552]" italics="true" pageId="286" pageNumber="287">fI-v4, fI-p2, fII-v2, fIII-v2</emphasis>
absent; 
<emphasis id="39FCEAEDFE94FEDDFD6A67F8FD0EF9B3" box="[649,711,1528,1552]" italics="true" pageId="286" pageNumber="287">tII-v1</emphasis>
and 
<emphasis id="39FCEAEDFE94FEDDFD1D67F8FC8CF9B3" box="[766,837,1528,1552]" italics="true" pageId="286" pageNumber="287">tIII-v1</emphasis>
short.
</paragraph>
<paragraph id="0B3736FFFE94FEDDFF24641CFAFBF893" blockId="286.[151,1437,412,2021]" pageId="286" pageNumber="287">
Abdomen (
<figureCitation id="93B32A7AFE94FEDDFEAE641CFE3AF997" box="[333,499,1564,1589]" captionStart="FIGURES 479 – 480" captionStartId="289.[151,264,1975,1997]" captionTargetBox="[197,1378,195,1935]" captionTargetId="figure@289.[185,1385,193,1949]" captionTargetPageId="289" captionText="FIGURES 479 – 480. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 479, male habitus, dorsal and ventral views. 480, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883412/files/figure.png" pageId="286" pageNumber="287">Figs 479–480</figureCitation>
) elongated oval in both sexes, longer in female. Tergopleurites rectangular; tergopleurites II–IX+X in male and tergopleurites II–VIII in female moderately divided medianly. Sternal plates rectangular, medianly continuous, do not approach pleurites. Pleural incrassations moderate. Ventral section of tergopleurites moderate to wide. Re-entrant heads prominent. Anterior end of pleurite II with rectangular extensions medianly. 
<collectingRegion id="C94CF81DFE94FEDDFE7264ACFE02F966" box="[401,459,1708,1733]" country="Maldives" name="Maale" pageId="286" pageNumber="287">Male</collectingRegion>
subgenital plate rounded triangular, reaching posterior margin of abdomen. Female subgenital plate roughly triangular, approaching but not reaching vulval margin (
<figureCitation id="93B32A7AFE94FEDDFBDE64CFFB6DF94B" box="[1085,1188,1743,1768]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 482</figureCitation>
). Lateral submarginal extensions present. Abdominal chaetotaxy as in 
<tableCitation id="460A0344FE94FEDDFD5564F4FCC5F8AF" box="[694,780,1780,1805]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II – VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 – 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="286" pageNumber="287">Table 2</tableCitation>
. Vulval margin with median bulge (
<figureCitation id="93B32A7AFE94FEDDFB4264F4FACAF8AF" box="[1185,1283,1780,1805]" captionStart="FIGURES 481 – 482" captionStartId="290.[151,264,1969,1991]" captionTargetBox="[185,1410,213,1919]" captionTargetId="figure@290.[166,1410,193,1926]" captionTargetPageId="290" captionText="FIGURES 481 – 482. Meropsiella apiastri (Denny, 1842) ex Merops apiaster: 481, male head, dorsal and ventral views. 482, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883414/files/figure.png" pageId="286" pageNumber="287">Fig. 482</figureCitation>
), 
<emphasis id="39FCEAEDFE94FEDDFAF464F5FAF0F8AF" box="[1303,1337,1781,1804]" italics="true" pageId="286" pageNumber="287">vss</emphasis>
and 
<emphasis id="39FCEAEDFE94FEDDFA9364F5FA55F8AF" box="[1392,1436,1781,1804]" italics="true" pageId="286" pageNumber="287">vms</emphasis>
not forming separate rows; 
<emphasis id="39FCEAEDFE94FEDDFE286519FE39F893" box="[459,496,1817,1840]" italics="true" pageId="286" pageNumber="287">vos</emphasis>
follow lateral margins of subgenital plate, not approaching vulval margin.
</paragraph>
<paragraph id="0B3736FFFE94FEDCFF24653CFB9DFEBF" blockId="286.[151,1437,412,2021]" lastBlockId="287.[151,1436,151,1005]" lastPageId="287" lastPageNumber="288" pageId="286" pageNumber="287">
<collectingRegion id="C94CF81DFE94FEDDFF24653CFEC8F8F6" box="[199,257,1852,1877]" country="Maldives" name="Maale" pageId="286" pageNumber="287">Male</collectingRegion>
genitalia (
<figureCitation id="93B32A7AFE94FEDDFE9B653CFDD9F8F7" box="[376,528,1852,1877]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="286" pageNumber="287">Fig. 483–491</figureCitation>
) variable. Basal apodeme trapezoidal (
<figureCitation id="93B32A7AFE94FEDDFC25653CFBE0F8F7" box="[966,1065,1852,1877]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="286" pageNumber="287">Fig. 483</figureCitation>
), rounded rectangular (
<figureCitation id="93B32A7AFE94FEDDFACE653CFA59F8F6" box="[1325,1424,1852,1877]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="286" pageNumber="287">Fig. 486</figureCitation>
), or pentagonal (
<figureCitation id="93B32A7AFE94FEDDFEA7655FFE61F8DB" box="[324,424,1887,1912]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="286" pageNumber="287">Fig. 489</figureCitation>
). Proximal mesosome blunt, either broad (e.g. 
<taxonomicName id="CC884D7CFE94FEDDFC20655FFB63F8DB" authorityName="Bush" authorityYear="2017" baseAuthorityName="Denny" baseAuthorityYear="1842" box="[963,1194,1887,1912]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="apiastri">
<emphasis id="39FCEAEDFE94FEDDFC20655FFB63F8DB" box="[963,1194,1887,1912]" italics="true" pageId="286" pageNumber="287">Meropsiella apiastri</emphasis>
</taxonomicName>
, 
<figureCitation id="93B32A7AFE94FEDDFB59655FFAD5F8DB" box="[1210,1308,1887,1912]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="286" pageNumber="287">Fig. 484</figureCitation>
) or narrow (e.g. 
<taxonomicName id="CC884D7CFE94FEDDFF326585FE49F83E" authorityName="Bush" authorityYear="2017" baseAuthorityName="Williams" baseAuthorityYear="1981" box="[209,384,1924,1949]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="286" pageNumber="287" phylum="Arthropoda" rank="species" species="bullockoda">
<emphasis id="39FCEAEDFE94FEDDFF326585FF3AF83F" box="[209,243,1925,1948]" italics="true" pageId="286" pageNumber="287">Ms</emphasis>
. 
<emphasis id="39FCEAEDFE94FEDDFEE76584FE49F83E" box="[260,384,1924,1949]" italics="true" pageId="286" pageNumber="287">bullockoda</emphasis>
</taxonomicName>
, 
<figureCitation id="93B32A7AFE94FEDDFE736584FE3AF83F" box="[400,499,1924,1949]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="286" pageNumber="287">Fig. 487</figureCitation>
). Proximal mesosome overlapping basal apodeme. Gonopore with broad marginal thickening that vary in shape (
<figureCitation id="93B32A7AFE94FEDDFDE265A8FD2EF863" box="[513,743,1960,1985]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="286" pageNumber="287">Figs 484, 487, 490</figureCitation>
); open only distally. Mesosomal lobes fused distally, often square-shaped, but variable between species; 2 
<emphasis id="39FCEAEDFE94FEDDFD4965CDFD2DF847" box="[682,740,1997,2020]" italics="true" pageId="286" pageNumber="287">ames</emphasis>
sensilla on each side lateral to gonopore; absent or not visible 
<taxonomicName id="CC884D7CFE95FEDCFF746297FEF2FF13" box="[151,315,151,176]" pageId="287" pageNumber="288" rank="subSpecies" subSpecies="apiastri">
in 
<emphasis id="39FCEAEDFE95FEDCFF576299FF1FFF13" box="[180,214,153,176]" italics="true" pageId="287" pageNumber="288">Ms</emphasis>
. 
<emphasis id="39FCEAEDFE95FEDCFF076299FEF2FF13" box="[228,315,153,176]" italics="true" pageId="287" pageNumber="288">apiastri</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE95FEDCFEA96297FE64FF13" box="[330,429,151,176]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="287" pageNumber="288">Fig. 484</figureCitation>
); 2 
<emphasis id="39FCEAEDFE95FEDCFE366299FDC6FF13" box="[469,527,153,176]" italics="true" pageId="287" pageNumber="288">pmes</emphasis>
sensilla on each side lateral or sublateral. Rugose nodi may be present as fringed or papillate sections of distal margins of mesosome (
<figureCitation id="93B32A7AFE95FEDCFD0462BCFC4EFF77" box="[743,903,188,213]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="287" pageNumber="288">Figs 487, 490</figureCitation>
). Parameral heads bifid, but variable (
<figureCitation id="93B32A7AFE95FEDCFAD162BCFF36FF5B" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="287" pageNumber="288">Figs 485, 488, 491</figureCitation>
). Parameral blades generally broad, somewhat elongated, but those of 
<taxonomicName id="CC884D7CFE95FEDCFBCA62E1FB10FF5B" authorityName="Bush" authorityYear="2017" baseAuthorityName="Williams" baseAuthorityYear="1981" box="[1065,1241,223,248]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="bullockoda">
<emphasis id="39FCEAEDFE95FEDCFBCA62E1FB82FF5B" box="[1065,1099,225,248]" italics="true" pageId="287" pageNumber="288">Ms</emphasis>
. 
<emphasis id="39FCEAEDFE95FEDCFBBF62DFFB10FF5B" box="[1116,1241,223,248]" italics="true" pageId="287" pageNumber="288">bullockoda</emphasis>
</taxonomicName>
(
<figureCitation id="93B32A7AFE95FEDCFB0A62DFFA85FF5B" box="[1257,1356,223,248]" captionStart="FIGURES 483 – 491" captionStartId="291.[151,264,1900,1922]" captionTargetBox="[214,1397,205,1860]" captionTargetId="figure@291.[172,1416,193,1879]" captionTargetPageId="291" captionText="FIGURES 483 – 491. Meropsiella spp. male genitalia. Meropsiella apiastri (Denny, 1842): 483, dorsal view. 484, mesosome, ventral view. 485, paramere, dorsal view. Meropsiella bullockoda (Williams, 1981) n. comb. 486: dorsal view. 487, mesosome, ventral view. 488, paramere, dorsal view. Meropsiella cf. erythropteri (Piaget, 1885) n. comb. ex Merops philippinicus: 489, dorsal view. 490, mesomere, ventral view. 491, paramere, dorsal view." httpUri="https://zenodo.org/record/883416/files/figure.png" pageId="287" pageNumber="288">Fig. 488</figureCitation>
) broad with irregular margins; 
<emphasis id="39FCEAEDFE95FEDCFE7C6304FE23FEBF" box="[415,490,260,284]" italics="true" pageId="287" pageNumber="288">pst1–2</emphasis>
both sensilla, central, not near distal end of parameres.
</paragraph>
</subSubSection>
<subSubSection id="43926574FE95FEDCFF246327FB18FE2B" pageId="287" pageNumber="288" type="biology_ecology">
<paragraph id="0B3736FFFE95FEDCFF246327FED1FEC6" blockId="287.[151,1436,151,1005]" pageId="287" pageNumber="288">
<emphasis id="39FCEAEDFE95FEDCFF246327FE56FEE3" bold="true" box="[199,415,295,320]" pageId="287" pageNumber="288">Host distribution.</emphasis>
Species of this genus are only known from the genus 
<taxonomicName id="CC884D7CFE95FEDCFBCC6329FA8AFEE3" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[1071,1347,295,320]" class="Aves" family="Meropidae" genus="Merops" kingdom="Animalia" order="Coraciiformes" pageId="287" pageNumber="288" phylum="Chordata" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFBCC6329FB4AFEE3" box="[1071,1155,297,320]" italics="true" pageId="287" pageNumber="288">Merops</emphasis>
<bibRefCitation id="6F194B0EFE95FEDCFB6C6327FA8AFEE3" author="Linnaeus" box="[1167,1347,295,320]" pageId="287" pageNumber="288" refString="Linnaeus, C. von (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tenth Edition. Volume 1. Laurentii Salvii, Holmiae. iv + 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
, family 
<taxonomicName id="CC884D7CFE95FEDCFF74634CFEDDFEC6" box="[151,276,332,357]" class="Aves" family="Meropidae" kingdom="Animalia" order="Coraciiformes" pageId="287" pageNumber="288" phylum="Chordata" rank="family">Meropidae</taxonomicName>
.
</paragraph>
<paragraph id="0B3736FFFE95FEDCFF24636FFB18FE2B" blockId="287.[151,1436,151,1005]" box="[199,1233,367,392]" pageId="287" pageNumber="288">
<materialsCitation id="BBE03CA2FE95FEDCFF24636FFB18FE2B" ID-GBIF-Occurrence="1587279281" box="[199,1233,367,392]" country="Australia" location="South-East" pageId="287" pageNumber="288" specimenCount="1" stateProvince="South-East">
<emphasis id="39FCEAEDFE95FEDCFF24636FFE6CFE2B" bold="true" box="[199,421,367,392]" pageId="287" pageNumber="288">Geographic range.</emphasis>
Occurs throughout Africa and Europe to 
<collectingRegion id="C94CF81DFE95FEDCFC94636FFC3BFE2B" box="[887,1010,367,392]" country="Botswana" name="South East" pageId="287" pageNumber="288">South-East</collectingRegion>
Asia and 
<collectingCountry id="739F766FFE95FEDCFB80636FFB04FE2B" box="[1123,1229,367,392]" name="Australia" pageId="287" pageNumber="288">Australia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="43926574FE95FEDCFF246394FE98FB96" pageId="287" pageNumber="288" type="discussion">
<paragraph id="0B3736FFFE95FEDCFF246394FC9DFD53" blockId="287.[151,1436,151,1005]" pageId="287" pageNumber="288">
<emphasis id="39FCEAEDFE95FEDCFF246394FEF2FE0E" bold="true" box="[199,315,404,429]" pageId="287" pageNumber="288">Remarks.</emphasis>
<taxonomicName id="CC884D7CFE95FEDCFEA06394FE00FE0E" authorityName="Conci" authorityYear="1941" box="[323,457,404,429]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFEA06394FE00FE0E" box="[323,457,404,429]" italics="true" pageId="287" pageNumber="288">Meropsiella</emphasis>
</taxonomicName>
was erected by 
<bibRefCitation id="6F194B0EFE95FEDCFD606394FCA8FE0F" author="Conci" box="[643,865,404,429]" pageId="287" pageNumber="288" refString="Conci, C. (1941 b) Due nuovi generi e tre nuove specie di Mallofagi dei Meropes. Bollettino della Societa Entomologica Italiana, 73, 99 - 107." type="journal article" year="1941" yearSuffix="b">Conci (1941b: 104)</bibRefCitation>
for the slender lice parasitising 
<taxonomicName id="CC884D7CFE95FEDCFB336394FA84FE0E" box="[1232,1357,404,429]" class="Aves" family="Meropidae" kingdom="Animalia" order="Coraciiformes" pageId="287" pageNumber="288" phylum="Chordata" rank="family">Meropidae</taxonomicName>
. 
<bibRefCitation id="6F194B0EFE95FEDCFABA6394FF26FE73" author="Conci" pageId="287" pageNumber="288" refString="Conci, C. (1941 b) Due nuovi generi e tre nuove specie di Mallofagi dei Meropes. Bollettino della Societa Entomologica Italiana, 73, 99 - 107." type="journal article" year="1941" yearSuffix="b">Conci (1941b)</bibRefCitation>
stated that 
<taxonomicName id="CC884D7CFE95FEDCFE9A63B7FE36FE73" authorityName="Conci" authorityYear="1941" box="[377,511,439,464]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFE9A63B7FE36FE73" box="[377,511,439,464]" italics="true" pageId="287" pageNumber="288">Meropsiella</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="CC884D7CFE95FEDCFD7E63B7FD37FE73" authorityName="Keler" authorityYear="1936" box="[669,766,439,464]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFD7E63B7FD37FE73" box="[669,766,439,464]" italics="true" pageId="287" pageNumber="288">Brueelia</emphasis>
</taxonomicName>
, but distinguished by the almost perfectly triangular head shape, the distinct clypeal suture located at about 4/5ths of the length of the head, the presence of a long and a short seta on the frons, and the lack of setae on the posterior dorsal corners of the prothorax. He included only one species in the genus
<taxonomicName id="CC884D7CFE95FEDCFE986025FDC6FD9F" box="[379,527,549,573]" pageId="287" pageNumber="288" rank="subSpecies" subSpecies="apiastri">
, 
<emphasis id="39FCEAEDFE95FEDCFE646025FE60FD9F" box="[391,425,549,572]" italics="true" pageId="287" pageNumber="288">Ms</emphasis>
. 
<emphasis id="39FCEAEDFE95FEDCFE5B6025FDC6FD9F" box="[440,527,549,572]" italics="true" pageId="287" pageNumber="288">apiastri</emphasis>
</taxonomicName>
, but suspected that 
<taxonomicName id="CC884D7CFE95FEDCFD0C6025FC6FFD9E" authorityName="Bush" authorityYear="2017" baseAuthorityName="Piaget" baseAuthorityYear="1885" box="[751,934,548,573]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="erythropteri">
<emphasis id="39FCEAEDFE95FEDCFD0C6025FCD8FD9F" box="[751,785,549,572]" italics="true" pageId="287" pageNumber="288">Ms</emphasis>
. 
<emphasis id="39FCEAEDFE95FEDCFCC36024FC6FFD9E" box="[800,934,548,573]" italics="true" pageId="287" pageNumber="288">erythropteri</emphasis>
</taxonomicName>
, which he was unable to examine, would be included as well. 
<bibRefCitation id="6F194B0EFE95FEDCFEBF6047FD56FDC3" author="Hopkins" box="[348,671,583,608]" pageId="287" pageNumber="288" refString="Hopkins, G. H. &amp; Clay, T. (1952) A check list of the genera &amp; species of Mallophaga. British Museum (Natural History), London. 362 pp." type="book" year="1952">Hopkins &amp; Clay (1952: 225)</bibRefCitation>
considered 
<taxonomicName id="CC884D7CFE95FEDCFCC56047FC65FDC3" authorityName="Conci" authorityYear="1941" box="[806,940,583,608]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFCC56047FC65FDC3" box="[806,940,583,608]" italics="true" pageId="287" pageNumber="288">Meropsiella</emphasis>
</taxonomicName>
inseparable from 
<taxonomicName id="CC884D7CFE95FEDCFB9B6047FB11FDC3" authorityName="Keler" authorityYear="1936" box="[1144,1240,583,608]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFB9B6047FB11FDC3" box="[1144,1240,583,608]" italics="true" pageId="287" pageNumber="288">Brueelia</emphasis>
</taxonomicName>
, a view followed also by 
<bibRefCitation id="6F194B0EFE95FEDCFF0E606CFE78FD27" author="Price" box="[237,433,620,645]" pageId="287" pageNumber="288" refString="Price, R. D., Hellenthal, R. A. &amp; Palma, R. L. (2003) World checklist of chewing lice with host associations and keys to families and genera. In: Price, R. D., Hellenthal, R. A., Palma, R. L., Johnson, K. P. &amp; Clayton, D. H. The Chewing lice: world checklist and biological overview. Illinois Natural History Survey Special Publication 24. x + 501 pp." type="book" year="2003">
Price 
<emphasis id="39FCEAEDFE95FEDCFECE606DFE97FD26" box="[301,350,620,645]" italics="true" pageId="287" pageNumber="288">et al</emphasis>
. (2003
</bibRefCitation>
: 198). In the phylogeny of Bush 
<emphasis id="39FCEAEDFE95FEDCFCFD606DFC86FD26" box="[798,847,620,645]" italics="true" pageId="287" pageNumber="288">et al</emphasis>
. (2016), 
<taxonomicName id="CC884D7CFE95FEDCFC57606CFBF2FD26" authorityName="Conci" authorityYear="1941" box="[948,1083,620,645]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFC57606CFBF2FD26" box="[948,1083,620,645]" italics="true" pageId="287" pageNumber="288">Meropsiella</emphasis>
</taxonomicName>
was placed with strong support as a lineage sister to the main 
<taxonomicName id="CC884D7CFE95FEDCFDC5608FFD4FFD0B" authorityName="Keler" authorityYear="1936" box="[550,646,655,680]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFDC5608FFD4FFD0B" box="[550,646,655,680]" italics="true" pageId="287" pageNumber="288">Brueelia</emphasis>
</taxonomicName>
-complex, together with several other genera (
<emphasis id="39FCEAEDFE95FEDCFB22608FFEEEFD6F" italics="true" pageId="287" pageNumber="288">
<taxonomicName id="CC884D7CFE95FEDCFB22608FFA51FD0B" authority=", Couala" authorityName="Couala" box="[1217,1432,655,680]" class="Insecta" family="Philopteridae" genus="Buerelius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Psocodea" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">Buerelius, Couala</taxonomicName>
, 
<taxonomicName id="CC884D7CFE95FEDCFF7460B5FEEAFD6F" authorityName="Eichler" authorityYear="1940" box="[151,291,693,716]" class="Insecta" family="Philopteridae" genus="Meropoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">Meropoecus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="CC884D7CFE95FEDCFEBC60B5FDCDFD6F" authorityName="Mey &amp; Barker" authorityYear="2014" box="[351,516,693,716]" class="Insecta" family="Philopteridae" genus="Motmotnirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFEBC60B5FDCDFD6F" box="[351,516,693,716]" italics="true" pageId="287" pageNumber="288">Motmotnirmus</emphasis>
</taxonomicName>
) that parasitise non-passerines. 
<bibRefCitation id="6F194B0EFE95FEDCFC8D60B5FBAFFD6F" author="Mey" box="[878,1126,692,717]" pageId="287" pageNumber="288" refString="Mey, E. &amp; Barker, S. C. (2014) Eine neue auf den Feenvogeln (Irenidae) lebende Brueelia - Art (Insecta, Phthiraptera, Ischnocera, Philopteridae), nebst Anmerkungen zur Gattung Brueelia Keler, 1936 sensu lato. Rudolstadter naturhistorische Schriften, 19, 73 - 114." type="journal article" year="2014">Mey &amp; Barker (2014)</bibRefCitation>
suggested that 
<taxonomicName id="CC884D7CFE95FEDCFAF660B4FA52FD6E" authorityName="Conci" authorityYear="1941" box="[1301,1435,692,717]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFAF660B4FA52FD6E" box="[1301,1435,692,717]" italics="true" pageId="287" pageNumber="288">Meropsiella</emphasis>
</taxonomicName>
should be resurrected, but offered no reasons for their opinion.
</paragraph>
<paragraph id="0B3736FFFE95FEDCFF2460FDFBEEFC4E" blockId="287.[151,1436,151,1005]" pageId="287" pageNumber="288">
To date, the only revision of the 
<taxonomicName id="CC884D7CFE95FEDCFDCD60FCFD47FCB6" authorityName="Keler" authorityYear="1936" box="[558,654,764,789]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFDCD60FCFD47FCB6" box="[558,654,764,789]" italics="true" pageId="287" pageNumber="288">Brueelia</emphasis>
</taxonomicName>
-complex species on the bee-eaters is by 
<bibRefCitation id="6F194B0EFE95FEDCFBB660FCFAC7FCB7" author="Williams" box="[1109,1294,764,789]" pageId="287" pageNumber="288" refString="Williams, N. S. (1981) The Brueelia (Mallophaga: Philopteridae) of the Meropidae (Aves: Coraciiformes). Journal of the Kansas Entomological Society, 54, 510 - 518." type="journal article" year="1981">Williams (1981)</bibRefCitation>
, who treated five species, three of which were newly described in that paper. However, we recognise 
<taxonomicName id="CC884D7CFE95FEDCFB50611FFE8CFCFF" authority="Williams, 1981" authorityName="Williams" authorityYear="1981" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="bullockoda">
<emphasis id="39FCEAEDFE95FEDCFB50611FFA55FC9B" box="[1203,1436,799,824]" italics="true" pageId="287" pageNumber="288">Brueelia bullockoda</emphasis>
<bibRefCitation id="6F194B0EFE95FEDCFF746144FE8CFCFF" author="Williams" box="[151,325,836,861]" pageId="287" pageNumber="288" refString="Williams, N. S. (1981) The Brueelia (Mallophaga: Philopteridae) of the Meropidae (Aves: Coraciiformes). Journal of the Kansas Entomological Society, 54, 510 - 518." type="journal article" year="1981">Williams, 1981</bibRefCitation>
</taxonomicName>
as the only species belonging to 
<taxonomicName id="CC884D7CFE95FEDCFD2B6144FC87FCFE" authorityName="Conci" authorityYear="1941" box="[712,846,836,861]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFD2B6144FC87FCFE" box="[712,846,836,861]" italics="true" pageId="287" pageNumber="288">Meropsiella</emphasis>
</taxonomicName>
, because we place 
<taxonomicName id="CC884D7CFE95FEDCFBCC6145FEE7FCDC" authority="Williams, 1981 in Aporisticeras" authorityName="Bush" authorityYear="2017" baseAuthorityName="Williams" baseAuthorityYear="1981" class="Insecta" family="Philopteridae" genus="Aporisticeras" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="athertona" status="gen. nov.">
<emphasis id="39FCEAEDFE95FEDCFBCC6145FB0FFCFE" box="[1071,1222,836,861]" italics="true" pageId="287" pageNumber="288">
<collectingRegion id="C94CF81DFE95FEDCFBCC6145FB85FCFF" box="[1071,1100,837,860]" country="Turkey" name="Bolu" pageId="287" pageNumber="288">Br</collectingRegion>
. athertona
</emphasis>
<bibRefCitation id="6F194B0EFE95FEDCFB2C6144FAB4FCFF" author="Williams" box="[1231,1405,836,861]" pageId="287" pageNumber="288" refString="Williams, N. S. (1981) The Brueelia (Mallophaga: Philopteridae) of the Meropidae (Aves: Coraciiformes). Journal of the Kansas Entomological Society, 54, 510 - 518." type="journal article" year="1981">
Williams, 
<quantity id="CC709B1AFE95FEDCFAA66144FAB4FCFF" box="[1349,1405,836,860]" metricMagnitude="1" metricUnit="m" metricValue="5.03174" pageId="287" pageNumber="288" unit="in" value="1981.0">1981</quantity>
</bibRefCitation>
in 
<emphasis id="39FCEAEDFE95FEDCFF746168FEE7FCDC" box="[151,302,872,895]" italics="true" pageId="287" pageNumber="288">Aporisticeras</emphasis>
</taxonomicName>
<emphasis id="39FCEAEDFE95FEDCFEDA6167FE5DFC23" bold="true" box="[313,404,871,896]" pageId="287" pageNumber="288">
<taxonomicNameLabel id="22CF5796FE95FEDCFEDA6167FE47FC23" box="[313,398,871,896]" pageId="287" pageNumber="288" rank="genus">n. gen.</taxonomicNameLabel>
,
</emphasis>
and 
<taxonomicName id="CC884D7CFE95FEDCFE376167FC4AFC23" authority="Williams, 1981" authorityName="Williams" authorityYear="1981" box="[468,899,871,896]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="superciliosa">
<emphasis id="39FCEAEDFE95FEDCFE376167FD01FC23" box="[468,712,871,896]" italics="true" pageId="287" pageNumber="288">Brueelia superciliosa</emphasis>
<bibRefCitation id="6F194B0EFE95FEDCFD306167FC4AFC23" author="Williams" box="[723,899,871,896]" pageId="287" pageNumber="288" refString="Williams, N. S. (1981) The Brueelia (Mallophaga: Philopteridae) of the Meropidae (Aves: Coraciiformes). Journal of the Kansas Entomological Society, 54, 510 - 518." type="journal article" year="1981">Williams, 1981</bibRefCitation>
</taxonomicName>
is 
<bibRefCitation id="6F194B0EFE95FEDCFC4E6168FC70FCDC" author="Williams" box="[941,953,872,895]" pageId="287" pageNumber="288" refString="Williams, N. S. (1981) The Brueelia (Mallophaga: Philopteridae) of the Meropidae (Aves: Coraciiformes). Journal of the Kansas Entomological Society, 54, 510 - 518." type="journal article" year="1981" yearSuffix="a">a</bibRefCitation>
junior synonym of 
<taxonomicName id="CC884D7CFE95FEDCFB496167FEAEFC07" authority="(Denny, 1842: 48)" authorityPageNumber="48" baseAuthorityName="Denny" baseAuthorityYear="1842" class="Insecta" family="Philopteridae" genus="Alcedoffula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="alcedinis">
<emphasis id="39FCEAEDFE95FEDCFB496167FA55FC23" box="[1194,1436,871,896]" italics="true" pageId="287" pageNumber="288">Alcedoffula alcedinis</emphasis>
(
<bibRefCitation id="6F194B0EFE95FEDCFF7D618CFE96FC07" author="Denny" box="[158,351,908,932]" pageId="287" pageNumber="288" refString="Denny, H. (1842) Monographia anoplurorum Britanniae. Henry G. Bohn, London. xxvi + 262 pp." type="book" year="1842">Denny, 1842: 48</bibRefCitation>
)
</taxonomicName>
(see 
<bibRefCitation id="6F194B0EFE95FEDCFE45618CFD96FC06" author="Williams" box="[422,607,908,933]" pageId="287" pageNumber="288" refString="Williams, N. S. (1982 b) Status of Brueelia superciliosa (Mallophaga: Philopteridae). Journal of the Kansas Entomological Society, 55, 334." type="journal article" year="1982" yearSuffix="b">Williams 1982b</bibRefCitation>
). We have examined a large amount of lice from several species of beeeaters and found small but consistent differences between material we list as 
<taxonomicName id="CC884D7CFE95FEDCFBFF61B1FB1CFC6B" authorityName="Bush" authorityYear="2017" baseAuthorityName="Piaget" baseAuthorityYear="1885" box="[1052,1237,943,968]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="erythropteri">
<emphasis id="39FCEAEDFE95FEDCFBFF61B1FB1CFC6B" box="[1052,1237,943,968]" italics="true" pageId="287" pageNumber="288">Ms. erythropteri</emphasis>
</taxonomicName>
in Appendix III; populations of 
<taxonomicName id="CC884D7CFE95FEDCFEDC61D5FE3CFC4E" authorityName="Bush" authorityYear="2017" baseAuthorityName="Piaget" baseAuthorityYear="1885" box="[319,501,980,1005]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="erythropteri">
<emphasis id="39FCEAEDFE95FEDCFEDC61D5FE3CFC4E" box="[319,501,980,1005]" italics="true" pageId="287" pageNumber="288">Ms. erythropteri</emphasis>
</taxonomicName>
on different host species may be separate species.
</paragraph>
<paragraph id="0B3736FFFE95FEDCFF74661CFE98FB96" blockId="287.[151,1047,1052,1184]" box="[151,337,1052,1077]" pageId="287" pageNumber="288">Included species</paragraph>
</subSubSection>
<subSubSection id="43926574FE95FEDCFF74663FFCD8FB3C" pageId="287" pageNumber="288" type="reference_group">
<paragraph id="0B3736FFFE95FEDCFF74663FFCD8FB3C" blockId="287.[151,1047,1052,1184]" pageId="287" pageNumber="288">
<treatmentCitationGroup id="2B9811D1FE95FEDCFF74663FFCD8FB3C" pageId="287" pageNumber="288">
<emphasis id="39FCEAEDFE95FEDCFF74663FFD3AFBFB" bold="true" box="[151,755,1087,1112]" pageId="287" pageNumber="288">
<emphasis id="39FCEAEDFE95FEDCFF74663FFF6BFBFB" bold="true" box="[151,162,1087,1112]" italics="true" pageId="287" pageNumber="288">*</emphasis>
<taxonomicName id="CC884D7CFE95FEDCFF41663FFDADFBFB" authority="(Denny, 1842: 52)" authorityName="Bush" authorityPageNumber="52" authorityYear="2017" baseAuthorityName="Denny" baseAuthorityYear="1842" box="[162,612,1087,1112]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="apiastri">
<emphasis id="39FCEAEDFE95FEDCFF41663FFE45FBFB" bold="true" box="[162,396,1087,1112]" italics="true" pageId="287" pageNumber="288">Meropsiella apiastri</emphasis>
(
<treatmentCitation id="8A2910EEFE95FEDCFE79663FFD95FBFB" author="Denny" box="[410,604,1087,1112]" page="52" pageId="287" pageNumber="288" year="1842">
<bibRefCitation id="6F194B0EFE95FEDCFE79663FFD95FBFB" author="Denny" box="[410,604,1087,1112]" pageId="287" pageNumber="288" refString="Denny, H. (1842) Monographia anoplurorum Britanniae. Henry G. Bohn, London. xxvi + 262 pp." type="book" year="1842">Denny, 1842: 52</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
[in 
<taxonomicName id="CC884D7CFE95FEDCFD70663FFD23FBFB" authorityName="Nitzsch" authorityYear="1818" box="[659,746,1087,1112]" class="Insecta" family="Philopteridae" genus="Nirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFD70663FFD23FBFB" bold="true" box="[659,746,1087,1112]" italics="true" pageId="287" pageNumber="288">Nirmus</emphasis>
</taxonomicName>
]
</emphasis>
<emphasis id="39FCEAEDFE95FEDCFF746664FF6BFBDE" box="[151,162,1124,1149]" italics="true" pageId="287" pageNumber="288">*</emphasis>
<taxonomicName id="CC884D7CFE95FEDCFF416664FD64FBDE" authority="(Williams, 1981: 516)" authorityName="Bush" authorityPageNumber="516" authorityYear="2017" baseAuthorityName="Williams" baseAuthorityYear="1981" box="[162,685,1124,1149]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="bullockoda">
<emphasis id="39FCEAEDFE95FEDCFF416664FE66FBDE" box="[162,431,1124,1149]" italics="true" pageId="287" pageNumber="288">Meropsiella bullockoda</emphasis>
(
<treatmentCitation id="8A2910EEFE95FEDCFE5E6664FD6CFBDE" author="Williams" box="[445,677,1124,1149]" page="516" pageId="287" pageNumber="288" year="1981">
<bibRefCitation id="6F194B0EFE95FEDCFE5E6664FD6CFBDE" author="Williams" box="[445,677,1124,1149]" pageId="287" pageNumber="288" refString="Williams, N. S. (1981) The Brueelia (Mallophaga: Philopteridae) of the Meropidae (Aves: Coraciiformes). Journal of the Kansas Entomological Society, 54, 510 - 518." type="journal article" year="1981">Williams, 1981: 516</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
[in 
<taxonomicName id="CC884D7CFE95FEDCFD396664FCF3FBDE" authorityName="Keler" authorityYear="1936" box="[730,826,1124,1149]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFD396664FCF3FBDE" box="[730,826,1124,1149]" italics="true" pageId="287" pageNumber="288">Brueelia</emphasis>
</taxonomicName>
] 
<emphasis id="39FCEAEDFE95FEDCFCA96664FBDEFBDE" bold="true" box="[842,1047,1124,1149]" pageId="287" pageNumber="288">new combination</emphasis>
<emphasis id="39FCEAEDFE95FEDCFF746687FF6BFB03" box="[151,162,1159,1184]" italics="true" pageId="287" pageNumber="288">*</emphasis>
<taxonomicName id="CC884D7CFE95FEDCFF416687FD40FB03" authority="(Piaget, 1885: 28)" authorityName="Bush" authorityPageNumber="28" authorityYear="2017" baseAuthorityName="Piaget" baseAuthorityYear="1885" box="[162,649,1159,1184]" class="Insecta" family="Philopteridae" genus="Meropsiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="species" species="erythropteri">
<emphasis id="39FCEAEDFE95FEDCFF416687FE71FB03" box="[162,440,1159,1184]" italics="true" pageId="287" pageNumber="288">Meropsiella erythropteri</emphasis>
(
<treatmentCitation id="8A2910EEFE95FEDCFE246687FD48FB03" author="Piaget" box="[455,641,1159,1184]" page="28" pageId="287" pageNumber="288" year="1885">
<bibRefCitation id="6F194B0EFE95FEDCFE246687FD48FB03" author="Piaget" box="[455,641,1159,1184]" pageId="287" pageNumber="288" refString="Piaget, E. (1885) Les Pediculines. Essai Monographique. Supplement. E. J. Brill, Leide. xvi + 200 pp., 17 pls." type="book" year="1885">Piaget, 1885: 28</bibRefCitation>
</treatmentCitation>
)
</taxonomicName>
[in 
<taxonomicName id="CC884D7CFE95FEDCFD556689FCC0FB03" authorityName="Nitzsch" authorityYear="1818" box="[694,777,1161,1184]" class="Insecta" family="Philopteridae" genus="Nirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="287" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="39FCEAEDFE95FEDCFD556689FCC0FB03" box="[694,777,1161,1184]" italics="true" pageId="287" pageNumber="288">Nirmus</emphasis>
</taxonomicName>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>