223 lines
19 KiB
XML
223 lines
19 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6492992" ID-ISSN="1982-0224" ID-Zenodo-Dep="6492992" approvalRequired="1" approvalRequired_for_originalDoi="1" checkinTime="1630380785404" checkinUser="carolina" docAuthor="A, Pablo Lehmann, Mayer, Fernanda & Reis, Roberto E." docDate="2010" docId="663487D6FFA8FFD2FEC4F8F3FB86C85F" docLanguage="en" docName="NeotropIchthyol.2010.7.pdf" docOrigin="Neotropical Ichthyology 8 (1)" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Otocinclus arnoldi Regan 1909" docType="treatment" docVersion="2" lastPageNumber="60" masterDocId="9A0DFFAEFFABFFD1FFD5FFFDFFE9CC56" masterDocTitle="Re-validation of Otocinclus arnoldi Regan and reappraisal of Otocinclus phylogeny (Siluriformes: Loricariidae)" masterLastPageNumber="68" masterPageNumber="57" pageId="3" pageNumber="60" updateTime="1650995928801" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Re-validation of Otocinclus arnoldi Regan and reappraisal of Otocinclus phylogeny (Siluriformes: Loricariidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>A, Pablo Lehmann</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mayer, Fernanda</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reis, Roberto E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Neotropical Ichthyology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>8</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>57</mods:start>
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<mods:end>68</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ISSN">1982-0224</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6492992" ID-Zenodo-Dep="6492992" LSID="urn:lsid:plazi:treatment:663487D6FFA8FFD2FEC4F8F3FB86C85F" httpUri="http://treatment.plazi.org/id/663487D6FFA8FFD2FEC4F8F3FB86C85F" lastPageNumber="60" pageId="3" pageNumber="60">
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<subSubSection box="[273,627,1806,1829]" pageId="3" pageNumber="60" type="nomenclature">
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<paragraph blockId="3.[273,627,1806,1861]" box="[273,627,1806,1829]" pageId="3" pageNumber="60">
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<heading box="[273,627,1806,1829]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="60" reason="2">
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<taxonomicName authority="Regan, 1909" authorityName="Regan" authorityYear="1909" box="[273,627,1806,1829]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
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<emphasis bold="true" box="[273,627,1806,1829]" pageId="3" pageNumber="60">
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<emphasis bold="true" box="[273,480,1806,1829]" italics="true" pageId="3" pageNumber="60">Otocinclus arnoldi</emphasis>
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<bibRefCitation author="Regan, C" box="[487,627,1806,1829]" pageId="3" pageNumber="60" pagination="234 - 235" refId="ref11288" refString="Regan, C. T. 1909. Descriptions of three new freshwater fishes from South America presented to the British Museum by Herr J. Paul Arnold. Annals and Magazine of Natural History, 3: 234 - 235." type="journal article" year="1909">Regan, 1909</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[419,480,1838,1861]" pageId="3" pageNumber="60" type="description">
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<paragraph blockId="3.[273,627,1806,1861]" box="[419,480,1838,1861]" pageId="3" pageNumber="60">
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<figureCitation box="[419,480,1838,1861]" captionStart="Fig" captionStartId="6.[136,174,1777,1800]" captionTargetBox="[170,1417,176,1757]" captionTargetId="figure-52@6.[170,1417,176,1757]" captionTargetPageId="6" captionText="Fig.3. Color variation of Otocinclus arnoldi.(a) and(b): MCP 25234, SL= 43.8 and39.0 mm; (c): MCP 32565, SL = 31.9 mm; and (d): UFRGS 6838,SL =30.4mm." pageId="3" pageNumber="60">
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<emphasis bold="true" box="[419,480,1838,1861]" pageId="3" pageNumber="60">Fig. 3</emphasis>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="60" type="reference_group">
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<paragraph blockId="3.[129,771,1902,1957]" pageId="3" pageNumber="60">
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<treatmentCitationGroup pageId="3" pageNumber="60">
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<treatmentCitation author="Regan, C" box="[129,520,1902,1925]" page="234" pageId="3" pageNumber="60" year="1909">
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<taxonomicName authority="Regan, 1909: 234" authorityName="Regan" authorityPageNumber="234" authorityYear="1909" box="[129,520,1902,1925]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
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<emphasis box="[129,327,1902,1925]" italics="true" pageId="3" pageNumber="60">Otocinclus arnoldi</emphasis>
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<materialsCitation box="[332,520,1902,1925]" collectingDate="1908-12-05" collectionCode="BMNH" collectorName="La Plata" location="South America." pageId="3" pageNumber="60" specimenCount="1" typeStatus="holotype">
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<bibRefCitation author="Regan, C" box="[332,520,1902,1925]" pageId="3" pageNumber="60" pagination="234 - 235" refId="ref11288" refString="Regan, C. T. 1909. Descriptions of three new freshwater fishes from South America presented to the British Museum by Herr J. Paul Arnold. Annals and Magazine of Natural History, 3: 234 - 235." type="journal article" year="1909">Regan, 1909: 234</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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</treatmentCitation>
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. Type locality:
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<collectorName box="[678,766,1902,1925]" pageId="3" pageNumber="60">La Plata</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:663487D6FFA8FFD2FEC4F8F3FB86C85F:EB42601BFFA8FFD2FF4AF873FED5CBF3" box="[159,316,1934,1957]" name="South America." pageId="3" pageNumber="60">South America.</location>
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<typeStatus box="[318,416,1934,1957]" pageId="3" pageNumber="60">Holotype</typeStatus>
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: BMNH
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<date box="[505,614,1934,1957]" pageId="3" pageNumber="60" value="1908-12-05">
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<collectingDate box="[505,614,1934,1957]" pageId="3" pageNumber="60" value="1908-12-05">1908.12.5</collectingDate>
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</date>
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:13, 43 mm SL.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="60" type="diagnosis">
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<paragraph blockId="3.[809,1452,178,905]" pageId="3" pageNumber="60">
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<emphasis bold="true" box="[809,926,178,201]" pageId="3" pageNumber="60">Diagnosis.</emphasis>
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<taxonomicName authorityName="Regan" authorityYear="1909" box="[931,1131,178,201]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
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<emphasis box="[931,1131,178,201]" italics="true" pageId="3" pageNumber="60">Otocinclus arnoldi</emphasis>
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</taxonomicName>
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is distinguished from all other
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<taxonomicName box="[809,1027,210,233]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="undetermined">
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<emphasis box="[809,933,210,233]" italics="true" pageId="3" pageNumber="60">Otocinclus</emphasis>
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species
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</taxonomicName>
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except
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<taxonomicName authorityName="Axenrot & Kullander" authorityYear="2003" box="[1125,1260,210,233]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
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<emphasis box="[1125,1144,210,233]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1164,1260,210,233]" italics="true" pageId="3" pageNumber="60">mimulus</emphasis>
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</taxonomicName>
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, by having five branched rays in the pectoral-fin (
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<emphasis box="[1175,1198,242,265]" italics="true" pageId="3" pageNumber="60">vs</emphasis>
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. six branched pectoralfin rays), and except from
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<taxonomicName authorityName="Axenrot & Kullander" authorityYear="2003" box="[1104,1231,274,297]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
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<emphasis box="[1104,1123,274,297]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1138,1231,274,297]" italics="true" pageId="3" pageNumber="60">mimulus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Schaefer" authorityYear="1997" box="[1285,1417,274,297]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="xakriaba">
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<emphasis box="[1285,1304,274,297]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1319,1417,274,297]" italics="true" pageId="3" pageNumber="60">xakriaba</emphasis>
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</taxonomicName>
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by having the prootic involved in the contact with the hyomandibular articular condyle (
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<emphasis box="[1170,1192,338,361]" italics="true" pageId="3" pageNumber="60">vs</emphasis>
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. only compound pterotic contacting the hyomandibular articular condyle). It is distinguished from
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<taxonomicName authorityName="Axenrot & Kullander" authorityYear="2003" box="[1012,1132,402,425]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
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<emphasis box="[1012,1031,402,425]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1042,1132,402,425]" italics="true" pageId="3" pageNumber="60">mimulus</emphasis>
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</taxonomicName>
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by having a not elevated patch of enlarged odontodes on the postero-dorsal parietosupraoccipital tip and 15 caudal vertebrae (
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<emphasis box="[1302,1325,466,489]" italics="true" pageId="3" pageNumber="60">vs</emphasis>
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. having an elevated patch of enlarged odontodes on the parietosupraoccipital and typically having 16-17 caudal vertebrae - see
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<tableCitation box="[849,930,562,585]" captionStart="Table 2" captionStartId="5.[129,192,178,201]" captionText="Table 2. Frequency distribution of variable counts of Otocinclus species. Caudal vertebrae counted in clear and stained specimens only. *Five specimens counted and 13 from radiographs by S.O. Kulander; **Data from radiographs by S. O. Kulander." pageId="3" pageNumber="60">Table 2</tableCitation>
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). It is further distinguished (except for
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<taxonomicName authorityName="Steindachner" authorityYear="1877" box="[1346,1445,562,585]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="affinis">
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<emphasis box="[1346,1365,562,585]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1377,1445,562,585]" italics="true" pageId="3" pageNumber="60">affinis</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[809,948,594,617]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="hasemani">
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<emphasis box="[809,828,594,617]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[845,948,594,617]" italics="true" pageId="3" pageNumber="60">hasemani</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Schaefer" authorityYear="1997" box="[964,1096,594,617]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="xakriaba">
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<emphasis box="[964,983,594,617]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1000,1096,594,617]" italics="true" pageId="3" pageNumber="60">xakriaba</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Axenrot & Kullander" authorityYear="2003" box="[1113,1239,594,617]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
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<emphasis box="[1113,1132,594,617]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1148,1239,594,617]" italics="true" pageId="3" pageNumber="60">mimulus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Cope" authorityYear="1894" box="[1298,1406,594,617]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="flexilis">
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<emphasis box="[1298,1317,594,617]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1334,1406,594,617]" italics="true" pageId="3" pageNumber="60">flexilis</emphasis>
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</taxonomicName>
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) by possessing an iris operculum, and from
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<taxonomicName authorityName="Steindachner" authorityYear="1877" box="[1218,1314,626,649]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="affinis">
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<emphasis box="[1218,1237,626,649]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1248,1314,626,649]" italics="true" pageId="3" pageNumber="60">affinis</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Axenrot & Kullander" authorityYear="2003" box="[1325,1445,626,649]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
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<emphasis box="[1325,1344,626,649]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1356,1445,626,649]" italics="true" pageId="3" pageNumber="60">mimulus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName authorityName="Schaefer" authorityYear="1997" box="[855,984,658,681]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="xakriaba">
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<emphasis box="[855,874,658,681]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[888,984,658,681]" italics="true" pageId="3" pageNumber="60">xakriaba</emphasis>
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</taxonomicName>
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by having a not elevated patch of enlarged odontodes on the postero-dorsal parieto-supraoccipital tip. It is also distinguished from all remaining
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<taxonomicName box="[1252,1451,722,745]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="undetermined">
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<emphasis box="[1252,1368,722,745]" italics="true" pageId="3" pageNumber="60">Otocinclus</emphasis>
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species
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</taxonomicName>
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(except for
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<taxonomicName authorityName="Schaefer" authorityYear="1997" box="[927,1050,754,777]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="xakriaba">
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<emphasis box="[927,946,754,777]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[957,1050,754,777]" italics="true" pageId="3" pageNumber="60">xakriaba</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Cope" authorityYear="1894" box="[1062,1162,754,777]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="flexilis">
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<emphasis box="[1062,1081,754,777]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1092,1162,754,777]" italics="true" pageId="3" pageNumber="60">flexilis</emphasis>
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</taxonomicName>
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, and
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<taxonomicName authorityName="Axenrot & Kullander" authorityYear="2003" box="[1216,1335,754,777]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
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<emphasis box="[1216,1235,754,777]" italics="true" pageId="3" pageNumber="60">O</emphasis>
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.
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<emphasis box="[1246,1335,754,777]" italics="true" pageId="3" pageNumber="60">mimulus</emphasis>
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</taxonomicName>
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) by having a lateral trunk coloration composed of either a row of 3-6 distinct dark blotches or a distinct dark stripe extending from the compound pterotic to the base of the caudal fin, or a diffuse mixture of those two color patterns (
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<figureCitation box="[1285,1350,882,905]" captionStart="Fig" captionStartId="6.[136,174,1777,1800]" captionTargetBox="[170,1417,176,1757]" captionTargetId="figure-52@6.[170,1417,176,1757]" captionTargetPageId="6" captionText="Fig.3. Color variation of Otocinclus arnoldi.(a) and(b): MCP 25234, SL= 43.8 and39.0 mm; (c): MCP 32565, SL = 31.9 mm; and (d): UFRGS 6838,SL =30.4mm." pageId="3" pageNumber="60">Fig. 3</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="60" type="distribution">
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<paragraph blockId="3.[809,1451,946,1033]" pageId="3" pageNumber="60">
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<emphasis bold="true" box="[809,957,946,969]" pageId="3" pageNumber="60">Distribution.</emphasis>
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<taxonomicName authorityName="Regan" authorityYear="1909" box="[963,1165,946,969]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
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<emphasis box="[963,1165,946,969]" italics="true" pageId="3" pageNumber="60">Otocinclus arnoldi</emphasis>
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</taxonomicName>
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occurs in the tributaries of the lower río Paraná drainage, lower and middle rio
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<collectingCountry box="[1356,1447,978,1001]" name="Uruguay" pageId="3" pageNumber="60">Uruguai</collectingCountry>
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, and the río de La Plata (
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<figureCitation box="[1057,1121,1010,1033]" captionStart="Fig" captionStartId="8.[136,174,752,775]" captionTargetBox="[156,753,185,739]" captionTargetId="figure-1800@8.[144,771,174,745]" captionTargetPageId="8" captionText="Fig. 5. Geographic distribution of Otocinclus flexilis (triangles) and O. arnoldi (dots). Open dot represents paratypes of O. mimulus. One symbol may cover more than one lot or locality." pageId="3" pageNumber="60">Fig. 5</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |