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<mods:title id="3FA8D193151102D8463463E5B853455A">New and poorly known giant scale insects (Homoptera: Coccinea: Margarodidae s. lat.) from the Oriental region with taxonomic and nomenclatural notes on the subfamily Monophlebinae</mods:title>
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<taxonomicName id="05AB79C01C2CFFF0FDEDF8DCDE45F12E" ID-CoL="448LV" authority="Burmeister, 1835" authorityName="Burmeister" authorityYear="1835" box="[541,1047,1853,1880]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="51" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2CFFF0FDEDF8DCD916F121" bold="true" box="[541,836,1853,1880]" italics="true" pageId="1" pageNumber="51">Monophlebus atripennis</emphasis>
<bibRefCitation id="A63A7FB21C2CFFF0FCBCF8DCDE45F12E" author="Burmeister H." box="[844,1047,1853,1880]" pageId="1" pageNumber="51" refId="ref4233" refString="Burmeister H. 1835. Handbuch der Entomologie. Vol. 2. T. F. Inslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1835">Burmeister, 1835</bibRefCitation>
</taxonomicName>
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<paragraph id="C21402431C2CFFF3FF4DF89EDB21F731" blockId="1.[189,1117,1919,1946]" lastBlockId="2.[187,1399,265,327]" lastPageId="2" lastPageNumber="52" pageId="1" pageNumber="51">
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<treatmentCitation id="430A24521C2CFFF0FF4DF89ED94EF1EC" author="Gavrilov-Zimin I. A." box="[189,796,1919,1946]" page="159" pageId="1" pageNumber="51" year="2018">
<taxonomicName id="05AB79C01C2CFFF0FF4DF89ED94EF1EC" authority="Gavrilov-Zimin, 2018: 159" authorityName="Gavrilov-Zimin" authorityPageNumber="159" authorityYear="2018" box="[189,796,1919,1946]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="51" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="F0DFDE511C2CFFF0FF4DF89EDB81F1EF" box="[189,467,1919,1945]" italics="true" pageId="1" pageNumber="51">Monophlebus neglectus</emphasis>
<bibRefCitation id="A63A7FB21C2CFFF0FE2AF89ED94EF1EC" author="Gavrilov-Zimin I. A." box="[474,796,1919,1946]" pageId="1" pageNumber="51" pagination="1 - 260" refId="ref4551" refString="Gavrilov-Zimin I. A. 2018. Ontogenesis, morphology and higher classification of archaecococcids (Homoptera: Coccinea: Orthezioidea). Zoosystematica Rossica Supplementum 2: 1 - 260. https: // doi. org / 10.31610 / zsr / 2018. supl. 2.1" type="journal article" year="2018">Gavrilov-Zimin, 2018: 159</bibRefCitation>
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, figs 9.4.69.4.7.
<emphasis id="F0DFDE511C2CFFF0FC1CF89EDE0FF1EF" bold="true" box="[1004,1117,1919,1945]" pageId="1" pageNumber="51">Syn. nov.</emphasis>
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Remarks (the taxonomic notes published in
<bibRefCitation id="A63A7FB21C2FFFF3FD4FFEE8D9A0F752" author="Gavrilov-Zimin I. A." box="[703,1010,265,292]" pageId="2" pageNumber="52" pagination="1 - 260" refId="ref4551" refString="Gavrilov-Zimin I. A. 2018. Ontogenesis, morphology and higher classification of archaecococcids (Homoptera: Coccinea: Orthezioidea). Zoosystematica Rossica Supplementum 2: 1 - 260. https: // doi. org / 10.31610 / zsr / 2018. supl. 2.1" type="journal article" year="2018">Gavrilov-Zimin 2018: 153</bibRefCitation>
on this subject are partly reported here for clarity)
</paragraph>
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<paragraph id="C21402431C2FFFF3FF4DFEBAD877F243" blockId="2.[188,1399,347,1077]" pageId="2" pageNumber="52">
<taxonomicName id="05AB79C01C2FFFF3FF4DFEBADB0FF703" authorityName="Guerin-Meneville" authorityYear="1827" box="[189,349,347,373]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="F0DFDE511C2FFFF3FF4DFEBADB0FF703" box="[189,349,347,373]" italics="true" pageId="2" pageNumber="52">Monophlebus</emphasis>
</taxonomicName>
, the
<typeStatus id="1D10BCE11C2FFFF3FE65FEBADB9AF703" box="[405,456,347,373]" pageId="2" pageNumber="52">type</typeStatus>
genus of the tribe
<taxonomicName id="05AB79C01C2FFFF3FD6FFEBAD91AF703" authorityName="Signoret" authorityYear="1875" box="[671,840,347,373]" class="Insecta" family="Margarodidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="tribe" tribe="Monophlebini">Monophlebini</taxonomicName>
, was originally introduced in the coccidological literature by
<bibRefCitation id="A63A7FB21C2FFFF3FEA8FE9FD8D0F7EE" author="Guerin-Meneville F. E." box="[344,642,382,408]" pageId="2" pageNumber="52" refId="ref4635" refString="Guerin-Meneville F. E. 1827. Dictionnaire classique d'histoire naturelle. Tome 11. Rey et Gravier Libraires-Editeurs, Paris. https: // doi. org / 10.5962 / bhl. title. 33901" type="book" year="1827">Guérin-Méneville (1827)</bibRefCitation>
who did not include any species in this genus, but provided a diagnosis: “Genre de 1ordre des Hémiptères, section des Homoptères, famille des Gallinsectes, établi par Latreille (Familles Naturelles du Règne Animal) et dont il ne donne pas les caractères; il dit seulement quil diffère des Dorthésies et des Cochenilles, parce que les antennes sont moniliformes et composées denviron vingt-deux articles” (
<bibRefCitation id="A63A7FB21C2FFFF3FDC4FDEAD92DF453" author="Guerin-Meneville F. E." box="[564,895,523,549]" pageId="2" pageNumber="52" refId="ref4635" refString="Guerin-Meneville F. E. 1827. Dictionnaire classique d'histoire naturelle. Tome 11. Rey et Gravier Libraires-Editeurs, Paris. https: // doi. org / 10.5962 / bhl. title. 33901" type="book" year="1827">Guérin-Méneville 1827: 99</bibRefCitation>
). Obviously, Guérin-Méneville mentioned male characters, because most of
<taxonomicName id="05AB79C01C2FFFF3FDB8FDCFD952F43E" authorityName="Signoret" authorityYear="1875" box="[584,768,558,584]" class="Insecta" family="Margarodidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="subFamily" subFamily="Monophlebinae">Monophlebinae</taxonomicName>
males have “antennes …moniliformes” in contrast to all known females. The first species of
<taxonomicName id="05AB79C01C2FFFF3FD73FDB0D971F41D" authorityName="Guerin-Meneville" authorityYear="1827" box="[643,803,593,619]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="F0DFDE511C2FFFF3FD73FDB0D971F41D" box="[643,803,593,619]" italics="true" pageId="2" pageNumber="52">Monophlebus</emphasis>
</taxonomicName>
were described 8 years later by
<bibRefCitation id="A63A7FB21C2FFFF3FB68FDB0DF20F41A" author="Burmeister H." box="[1176,1394,593,620]" pageId="2" pageNumber="52" refId="ref4233" refString="Burmeister H. 1835. Handbuch der Entomologie. Vol. 2. T. F. Inslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1835">Burmeister (1835)</bibRefCitation>
, who provided very brief and incomplete descriptions of the males of
<taxonomicName id="05AB79C01C2FFFF3FBF2FD94DF25F4F9" authority="Burmeister, 1835" authorityName="Burmeister" authorityYear="1835" box="[1026,1399,628,655]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2FFFF3FBF2FD94DEF0F4F8" box="[1026,1186,629,654]" italics="true" pageId="2" pageNumber="52">M. atripennis</emphasis>
<bibRefCitation id="A63A7FB21C2FFFF3FB5AFD95DF25F4F9" author="Burmeister H." box="[1194,1399,628,655]" pageId="2" pageNumber="52" refId="ref4233" refString="Burmeister H. 1835. Handbuch der Entomologie. Vol. 2. T. F. Inslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1835">Burmeister, 1835</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="006FCCA11C2FFFF3FF0FFD79DB61F4C4" box="[255,307,664,690]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="52">Java</collectingRegion>
(
<collectingCountry id="BABC42D31C2FFFF3FEB2FD76DBEAF4C4" box="[322,440,663,690]" name="Indonesia" pageId="2" pageNumber="52">Indonesia</collectingCountry>
) and
<taxonomicName id="05AB79C01C2FFFF3FE06FD79D921F4C4" authority="Burmeister, 1835" authorityName="Burmeister" authorityYear="1835" box="[502,883,663,690]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="F0DFDE511C2FFFF3FE06FD79D8F3F4C4" box="[502,673,664,690]" italics="true" pageId="2" pageNumber="52">M. fuscipennis</emphasis>
<bibRefCitation id="A63A7FB21C2FFFF3FD58FD76D921F4C4" author="Burmeister H." box="[680,883,663,690]" pageId="2" pageNumber="52" refId="ref4233" refString="Burmeister H. 1835. Handbuch der Entomologie. Vol. 2. T. F. Inslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1835">Burmeister, 1835</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="BABC42D31C2FFFF3FC4AFD79DE7BF4C4" box="[954,1065,664,690]" name="Germany" pageId="2" pageNumber="52">Germany</collectingCountry>
. The
<typeStatus id="1D10BCE11C2FFFF3FB98FD79DEC9F4C4" box="[1128,1179,664,690]" pageId="2" pageNumber="52">type</typeStatus>
specimens of both these species were lost.
<bibRefCitation id="A63A7FB21C2FFFF3FE12FD5AD8A1F4A3" author="Cockerell T. D. A." box="[482,755,699,725]" pageId="2" pageNumber="52" pagination="232 - 233" refId="ref4306" refString="Cockerell T. D. A. 1902 a. A contribution to the classification of the Coccidae. The Entomologist 35: 232 - 233. https: // doi. org / 10.5962 / bhl. part. 19973" type="journal article" year="1902">Cockerell (1902a: 232</bibRefCitation>
;
<bibRefCitation id="A63A7FB21C2FFFF3FCF1FD5AD9D8F4A3" author="Cockerell T. D. A." box="[769,906,699,725]" pageId="2" pageNumber="52" pagination="317 - 319" refId="ref4347" refString="Cockerell T. D. A. 1902 b. What is Monophlebus Leach? The Entomologist 35: 317 - 319." type="journal article" year="1902">1902b: 317</bibRefCitation>
) designated and discussed
<taxonomicName id="05AB79C01C2FFFF3FB25FD5ADF24F4A2" authorityName="Burmeister" authorityYear="1835" box="[1237,1398,699,724]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2FFFF3FB25FD5ADF24F4A2" box="[1237,1398,699,724]" italics="true" pageId="2" pageNumber="52">M. atripennis</emphasis>
</taxonomicName>
as a
<typeStatus id="1D10BCE11C2FFFF3FF08FD3FDB79F48E" box="[248,299,734,760]" pageId="2" pageNumber="52">type</typeStatus>
species of genus
<taxonomicName id="05AB79C01C2FFFF3FDF7FD3FD8F5F48E" authorityName="Guerin-Meneville" authorityYear="1827" box="[519,679,734,760]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="F0DFDE511C2FFFF3FDF7FD3FD8F5F48E" box="[519,679,734,760]" italics="true" pageId="2" pageNumber="52">Monophlebus</emphasis>
</taxonomicName>
, in spite of the absence of any characters for identification of this species.
<bibRefCitation id="A63A7FB21C2FFFF3FE89FCE3D84FF56A" author="Reyne A." box="[377,541,769,796]" pageId="2" pageNumber="52" pagination="145 - 188" refId="ref4845" refString="Reyne A. 1965. Observations on some Indonesian scale insects. Tijdschrift voor Entomologie 108: 145 - 188. Available from https: // www. biodiversitylibrary. org / page / 28271610 # page / 189 / mode / 1 up [accessed 27 Mar. 2021]." type="journal article" year="1965">Reyne (1965)</bibRefCitation>
conducted an extensive study of
<taxonomicName id="05AB79C01C2FFFF3FC42FCE0DE38F56D" authorityName="Signoret" authorityYear="1875" box="[946,1130,769,795]" class="Insecta" family="Margarodidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="subFamily" subFamily="Monophlebinae">Monophlebinae</taxonomicName>
males, collected from Java, but was unable to identify
<taxonomicName id="05AB79C01C2FFFF3FDBAFCC4D8B9F548" authorityName="Burmeister" authorityYear="1835" box="[586,747,805,830]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2FFFF3FDBAFCC4D8B9F548" box="[586,747,805,830]" italics="true" pageId="2" pageNumber="52">M. atripennis</emphasis>
</taxonomicName>
. Moreover, scale insect systematics and identification is mainly based on female characters, whereas females of
<taxonomicName id="05AB79C01C2FFFF3FC76FCA9DE7AF517" authorityName="Burmeister" authorityYear="1835" box="[902,1064,840,865]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2FFFF3FC76FCA9DE7AF517" box="[902,1064,840,865]" italics="true" pageId="2" pageNumber="52">M. atripennis</emphasis>
</taxonomicName>
are unknown at all. So, an unambiguous identification of
<taxonomicName id="05AB79C01C2FFFF3FDD1FC8AD8ECF5F2" authorityName="Burmeister" authorityYear="1835" box="[545,702,875,900]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2FFFF3FDD1FC8AD8ECF5F2" box="[545,702,875,900]" italics="true" pageId="2" pageNumber="52">M. atripennis</emphasis>
</taxonomicName>
is not possible until now. The second species of Burmeister,
<taxonomicName id="05AB79C01C2FFFF3FF4CFC6EDB3AF5DE" authorityName="Burmeister" authorityYear="1835" box="[188,360,910,936]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="F0DFDE511C2FFFF3FF4CFC6EDB3AF5DE" box="[188,360,910,936]" italics="true" pageId="2" pageNumber="52">M. fuscipennis</emphasis>
</taxonomicName>
, was transferred by
<bibRefCitation id="A63A7FB21C2FFFF3FDABFC6FD971F5DE" author="Cockerell T. D. A." box="[603,803,910,936]" pageId="2" pageNumber="52" pagination="31 - 36" refId="ref4268" refString="Cockerell T. D. A. 1894. A check list of Nearctic Coccidae. The Canadian Entomologist 26: 31 - 36. https: // doi. org / 10.4039 / Ent 2631 - 2" type="journal article" year="1894">Cockerell (1894)</bibRefCitation>
to his genus
<taxonomicName id="05AB79C01C2FFFF3FC33FC6FDF7EF5DE" authority="Cockerell, 1894" authorityName="Cockerell" authorityYear="1894" box="[963,1324,910,936]" class="Insecta" family="Margarodidae" genus="Palaeococcus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="F0DFDE511C2FFFF3FC33FC6FDE34F5DE" box="[963,1126,910,936]" italics="true" pageId="2" pageNumber="52">Palaeococcus</emphasis>
<bibRefCitation id="A63A7FB21C2FFFF3FB9DFC6FDF7EF5DE" author="Cockerell T. D. A." box="[1133,1324,910,936]" pageId="2" pageNumber="52" pagination="31 - 36" refId="ref4268" refString="Cockerell T. D. A. 1894. A check list of Nearctic Coccidae. The Canadian Entomologist 26: 31 - 36. https: // doi. org / 10.4039 / Ent 2631 - 2" type="journal article" year="1894">Cockerell, 1894</bibRefCitation>
</taxonomicName>
. Later
<bibRefCitation id="A63A7FB21C2FFFF3FF4EFC50DB9FF5BD" author="Cockerell T. D. A." box="[190,461,945,971]" pageId="2" pageNumber="52" pagination="232 - 233" refId="ref4306" refString="Cockerell T. D. A. 1902 a. A contribution to the classification of the Coccidae. The Entomologist 35: 232 - 233. https: // doi. org / 10.5962 / bhl. part. 19973" type="journal article" year="1902">Cockerell (1902a: 233)</bibRefCitation>
designated
<taxonomicName id="05AB79C01C2FFFF3FDA9FC53D956F5BD" authorityName="Burmeister" authorityYear="1835" box="[601,772,945,971]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="F0DFDE511C2FFFF3FDA9FC53D956F5BD" box="[601,772,945,971]" italics="true" pageId="2" pageNumber="52">M. fuscipennis</emphasis>
</taxonomicName>
as the
<typeStatus id="1D10BCE11C2FFFF3FCA6FC53D9DBF5BA" box="[854,905,946,972]" pageId="2" pageNumber="52">type</typeStatus>
species of the genus. Several taxonomists (
<bibRefCitation id="A63A7FB21C2FFFF3FF35FC35DBD7F599" author="Westwood J. O." box="[197,389,980,1007]" pageId="2" pageNumber="52" pagination="21 - 22" refId="ref4965" refString="Westwood J. O. 1845. Plate VI. Illustrations of various species of Coccidae, belonging to the genus Monophlebus. Arcana Entomologica 1: 21 - 22. https: // doi. org / 10.5962 / bhl. title. 11771" type="journal article" year="1845">Westwood 1845</bibRefCitation>
;
<bibRefCitation id="A63A7FB21C2FFFF3FE60FC35D863F598" author="Hempel A." box="[400,561,980,1006]" pageId="2" pageNumber="52" pagination="329 - 377" refId="ref4671" refString="Hempel A. 1920. Descripcoes de coccidas novas e pouco conhecidas. Revista do Museu Paulista (Sao Paulo) 12: 329 - 377." type="journal article" year="1920">Hempel 1920</bibRefCitation>
;
<bibRefCitation id="A63A7FB21C2FFFF3FDCBFC35D8BFF598" author="Vayssiere P." box="[571,749,980,1006]" pageId="2" pageNumber="52" pagination="24 - 28" refId="ref4897" refString="Vayssiere P. 1932. Monophlebines et pseudococcines de l'Afrique du Nord [Hem. Coccidae]. Bulletin de la Societe entomologique de France 37: 24 - 28. Available from https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1932 _ num _ 37 _ 2 _ 14365 [accessed 27 Mar. 2021]." type="journal article" year="1932">Vayssière 1932</bibRefCitation>
) described additional species in the genus
<taxonomicName id="05AB79C01C2FFFF3FB27FC35DF25F598" authorityName="Guerin-Meneville" authorityYear="1827" box="[1239,1399,980,1006]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="F0DFDE511C2FFFF3FB27FC35DF25F598" box="[1239,1399,980,1006]" italics="true" pageId="2" pageNumber="52">Monophlebus</emphasis>
</taxonomicName>
from other regions of the world, but not from Java! All these descriptions are also incomplete or even based on male characters only.
</paragraph>
<paragraph id="C21402431C2FFFF3FF4DFBBCDE09F036" blockId="2.[187,1401,1116,1600]" pageId="2" pageNumber="52">
In order to resolve this very old problem,
<bibRefCitation id="A63A7FB21C2FFFF3FD6DFBBDD9B7F201" author="Gavrilov-Zimin I. A." box="[669,997,1116,1143]" pageId="2" pageNumber="52" pagination="1 - 260" refId="ref4551" refString="Gavrilov-Zimin I. A. 2018. Ontogenesis, morphology and higher classification of archaecococcids (Homoptera: Coccinea: Orthezioidea). Zoosystematica Rossica Supplementum 2: 1 - 260. https: // doi. org / 10.31610 / zsr / 2018. supl. 2.1" type="journal article" year="2018">Gavrilov-Zimin (2018: 153)</bibRefCitation>
suggested to use the only properly described species of
<taxonomicName id="05AB79C01C2FFFF3FE49FB9ED805F2EF" authorityName="Guerin-Meneville" authorityYear="1827" box="[441,599,1151,1177]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="F0DFDE511C2FFFF3FE49FB9ED805F2EF" box="[441,599,1151,1177]" italics="true" pageId="2" pageNumber="52">Monophlebus</emphasis>
</taxonomicName>
,
<taxonomicName id="05AB79C01C2FFFF3FD97FB61DE43F2EF" authority="Gavrilov-Zimin, 2018" authorityName="Gavrilov-Zimin" authorityYear="2018" box="[615,1041,1151,1177]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="F0DFDE511C2FFFF3FD97FB61D8ADF2EF" box="[615,767,1151,1177]" italics="true" pageId="2" pageNumber="52">M. neglectus</emphasis>
<bibRefCitation id="A63A7FB21C2FFFF3FCF8FB9EDE43F2EF" author="Gavrilov-Zimin I. A." box="[776,1041,1151,1177]" pageId="2" pageNumber="52" pagination="1 - 260" refId="ref4551" refString="Gavrilov-Zimin I. A. 2018. Ontogenesis, morphology and higher classification of archaecococcids (Homoptera: Coccinea: Orthezioidea). Zoosystematica Rossica Supplementum 2: 1 - 260. https: // doi. org / 10.31610 / zsr / 2018. supl. 2.1" type="journal article" year="2018">Gavrilov-Zimin, 2018</bibRefCitation>
</taxonomicName>
, collected in Java, as the new type species of the genus. However, the replacement of the type species could only be accomplished with the used of the plenary power by the International Commission on Zoological Nomenclature (Art. 81.1). Before the application to the ICZN, the type specimen for each involved species (the older type species and a newly proposed type species) have to be fixed. Since the type of
<taxonomicName id="05AB79C01C2FFFF3FC17FAEDDF56F350" authorityName="Burmeister" authorityYear="1835" box="[999,1284,1292,1318]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2FFFF3FC17FAEDDF56F350" box="[999,1284,1292,1318]" italics="true" pageId="2" pageNumber="52">Monophlebus atripennis</emphasis>
</taxonomicName>
is lost, an alternative solution was suggested by one of the ICZN commissioners (D.A. Dmitriev,
<collectingRegion id="006FCCA11C2FFFF3FB32FACEDF47F33F" box="[1218,1301,1327,1353]" country="United States of America" name="Illinois" pageId="2" pageNumber="52">Illinois</collectingRegion>
Natural History Survey,
<collectingCountry id="BABC42D31C2FFFF3FE70FAB2DBECF31B" box="[384,446,1363,1389]" name="United States of America" pageId="2" pageNumber="52">USA</collectingCountry>
, pers. com.). A
<typeStatus id="1D10BCE11C2FFFF3FD8BFAB2D88AF31B" box="[635,728,1363,1389]" pageId="2" pageNumber="52" type="neotype">neotype</typeStatus>
could be designated for
<taxonomicName id="05AB79C01C2FFFF3FBF3FAB3DF72F31A" authorityName="Burmeister" authorityYear="1835" box="[1027,1312,1362,1388]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2FFFF3FBF3FAB3DF72F31A" box="[1027,1312,1362,1388]" italics="true" pageId="2" pageNumber="52">Monophlebus atripennis</emphasis>
</taxonomicName>
, and if this type specimen is, at the same time, the
<typeStatus id="1D10BCE11C2FFFF3FD20FA97D965F3E6" box="[720,823,1398,1424]" pageId="2" pageNumber="52" type="holotype">holotype</typeStatus>
of
<taxonomicName id="05AB79C01C2FFFF3FC92FA97DE2BF3E6" authorityName="Gavrilov-Zimin" authorityYear="2018" box="[866,1145,1398,1424]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="F0DFDE511C2FFFF3FC92FA97DE2BF3E6" box="[866,1145,1398,1424]" italics="true" pageId="2" pageNumber="52">Monophlebus neglectus</emphasis>
</taxonomicName>
, this would establish the objective synonymy between two species names. Following this recommendation, I here designate the
<typeStatus id="1D10BCE11C2FFFF3FF1DFA5DDB06F3A0" box="[237,340,1468,1494]" pageId="2" pageNumber="52" type="holotype">holotype</typeStatus>
specimen of
<emphasis id="F0DFDE511C2FFFF3FE0BFA5DD95FF3A0" box="[507,781,1468,1494]" italics="true" pageId="2" pageNumber="52">
Monophebus
<taxonomicName id="05AB79C01C2FFFF3FD6EFA5DD95FF3A0" authorityName="Gavrilov-Zimin" authorityYear="2018" box="[670,781,1468,1494]" class="Insecta" family="Margarodidae" genus="Monophlebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="neglectus">neglectus</taxonomicName>
</emphasis>
also to be the
<typeStatus id="1D10BCE11C2FFFF3FC3AFA5CDE75F3A1" box="[970,1063,1469,1495]" pageId="2" pageNumber="52" type="neotype">neotype</typeStatus>
of
<taxonomicName id="05AB79C01C2FFFF3FBA5FA5DDF3DF3A0" box="[1109,1391,1468,1494]" class="Insecta" family="Margarodidae" genus="Monophebus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="52" phylum="Arthropoda" rank="species" species="atripennis">
<emphasis id="F0DFDE511C2FFFF3FBA5FA5DDF3DF3A0" box="[1109,1391,1468,1494]" italics="true" pageId="2" pageNumber="52">Monophebus atripennis</emphasis>
</taxonomicName>
, establishing the formal synonymy between the two names. The
<typeStatus id="1D10BCE11C2FFFF3FC30FA01DE4FF38C" box="[960,1053,1504,1530]" pageId="2" pageNumber="52" type="neotype">neotype</typeStatus>
label data are: adult female, K 1301-a,
<collectingCountry id="BABC42D31C2FFFF3FECAF9E3DBFDF06A" box="[314,431,1538,1564]" name="Indonesia" pageId="2" pageNumber="52">Indonesia</collectingCountry>
,
<collectingRegion id="006FCCA11C2FFFF3FE4BF9E2DBA0F06B" box="[443,498,1539,1565]" country="Indonesia" name="Jawa Timur" pageId="2" pageNumber="52">Java</collectingRegion>
, “Buitenzorg” [Bogor], 1907, host plant and collectors name unknown. The
<typeStatus id="1D10BCE11C2FFFF3FF4DF9C7DB48F036" box="[189,282,1574,1600]" pageId="2" pageNumber="52" type="neotype">neotype</typeStatus>
is deposited in the Zoological Institute, Russian Academy of Sciences.
</paragraph>
</subSubSection>
</treatment>
</document>