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<document id="D3B980C8E39F4E01ED331540684E4CF6" ID-DOI="10.5281/zenodo.13651136" ID-ISSN="1732-2421" ID-Zenodo-Dep="13651136" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723142747611" checkinUser="felipe" docAuthor="Malyshkina, Tatiana" docDate="2006" docId="444087E3B95EFF829E7E0A30799CFA45" docLanguage="en" docName="ActaPalaeontolPol.51.3.465-475.pdf" docOrigin="Acta Palaeontologica Polonica 51 (3)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Premontreia (Premontreia) uralica Malyshkina, 2006, sp. nov." docType="treatment" docVersion="3" lastPageNumber="471" masterDocId="B879FF9BB95AFF849E180E197C12FFEE" masterDocTitle="Late Eocene scyliorhinid sharks from the Trans-Urals, Russia" masterLastPageNumber="475" masterPageNumber="465" pageNumber="469" updateTime="1725391873372" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="28C8CC2F5CE615DA2B7E92CBC9C58B37">Late Eocene scyliorhinid sharks from the Trans-Urals, Russia</mods:title>
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<mods:name id="F1E64D786228CBA618AB034EC9EBCE14" type="personal">
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<mods:roleTerm id="CCC75AC109653A6B8E75DB2108DCF73B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8C0B0C196F48690952E8C7D219267C7D">Malyshkina, Tatiana</mods:namePart>
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<mods:typeOfResource id="6D21539AC0478D871C7E67F8AABD3FBD">text</mods:typeOfResource>
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<mods:title id="739BB26BA3FFDA610FF8CA2D96608481">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="ADD988426B73291F9CA70AFB8F46B7F9">
<mods:date id="6E7886BF9AFE3A5A1EE65982ED7EA976">2006</mods:date>
<mods:detail id="4E4C074E3F0C2D1820A2EDB2BA5A8167" type="volume">
<mods:number id="1C76683F5C92B5B1E5E60747D9BD1273">51</mods:number>
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<mods:classification id="BAA23378A867BB989CCE13992D7C28DB">journal article</mods:classification>
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<mods:identifier id="BE5B2C9FB63B65015C757EDE65C3EDD5" type="ISSN">1732-2421</mods:identifier>
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<treatment id="444087E3B95EFF829E7E0A30799CFA45" LSID="urn:lsid:plazi:treatment:444087E3B95EFF829E7E0A30799CFA45" httpUri="http://treatment.plazi.org/id/444087E3B95EFF829E7E0A30799CFA45" lastPageId="6" lastPageNumber="471" pageId="4" pageNumber="469">
<subSubSection id="84F3657EB95EFF809E7E0A307E85FBAB" box="[102,663,1065,1093]" pageId="4" pageNumber="469" type="nomenclature">
<paragraph id="CC5636F5B95EFF809E7E0A307E85FBAB" blockId="4.[102,663,1065,1125]" box="[102,663,1065,1093]" pageId="4" pageNumber="469">
<heading id="971E8199B95EFF809E7E0A307E85FBAB" box="[102,663,1065,1093]" fontSize="12" level="2" pageId="4" pageNumber="469" reason="9">
<taxonomicName id="0BE94D76B95EFF809E7E0A307E38FBAB" authorityName="Malyshkina" authorityYear="2006" box="[102,554,1065,1093]" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="469" phylum="Chordata" rank="species" species="uralica" status="sp. nov." subGenus="Premontreia">
<emphasis id="FE9DEAE7B95EFF809E7E0A307D1AFBAB" box="[102,264,1065,1093]" italics="true" pageId="4" pageNumber="469">Premontreia</emphasis>
(
<emphasis id="FE9DEAE7B95EFF809F020A307DAEFBAB" box="[282,444,1065,1093]" italics="true" pageId="4" pageNumber="469">Premontreia</emphasis>
)
<emphasis id="FE9DEAE7B95EFF809FD60A307E38FBAB" box="[462,554,1065,1093]" italics="true" pageId="4" pageNumber="469">uralica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AE579CB95EFF809C2B0A307E85FBAB" box="[563,663,1065,1093]" pageId="4" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="84F3657EB95EFF829E7E0A49799CFA45" lastPageId="6" lastPageNumber="471" pageId="4" pageNumber="469" type="description">
<paragraph id="CC5636F5B95EFF809E7E0A497D08FB8B" blockId="4.[102,663,1065,1125]" box="[102,282,1104,1125]" pageId="4" pageNumber="469">
<figureCitation id="54D22A70B95EFF809E7E0A497CBBFB8B" box="[102,169,1104,1125]" captionStart="Fig" captionStartId="3.[102,135,1514,1534]" captionTargetBox="[102,1485,226,1486]" captionTargetId="figure-215@3.[102,1486,226,1486]" captionTargetPageId="3" captionText="Fig. 2. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. Holotype. PIN 5088/01, anterior tooth; in labial (A1), lingual (A2), labially−occlusal (A3), distally−basal (A4), lingually−occlusal (A5), and basal (A6) views. B. Paratype. PIN 5088/02, anterior tooth; in labial (B1), distally−basal (B2), lingual (B3), labially−occlusal (B4), lingually−occlusal (B5), and basal (B6) views. SEM microphotographs." figureDoi="http://doi.org/10.5281/zenodo.13651142" httpUri="https://zenodo.org/record/13651142/files/figure.png" pageId="4" pageNumber="469">Figs. 2</figureCitation>
,
<figureCitation id="54D22A70B95EFF809EAB0A497CD2FB8B" box="[179,192,1104,1125]" captionStart="Fig" captionStartId="4.[102,135,954,974]" captionTargetBox="[102,1485,229,924]" captionTargetId="figure-456@4.[102,1486,229,924]" captionTargetPageId="4" captionText="Fig. 3. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. 8CΠ/Ku−87/21, anterior tooth; in labial (A1), lingual (A2), occlusal (A3), distal (A4), and basal (A5) views. B. Lateral tooth; in labial (B1), lingual (B2), distal (B3), and occlusal (B4) views. Digital photographs." figureDoi="http://doi.org/10.5281/zenodo.13651148" httpUri="https://zenodo.org/record/13651148/files/figure.png" pageId="4" pageNumber="469">3</figureCitation>
,
<tableCitation id="816B034EB95EFF809ED30A497D07FB8B" box="[203,277,1104,1125]" captionStart="Table 1" captionStartId="4.[102,157,1727,1748]" captionText="Table 1. Measurements of the types series of Premontreia (Premontreia) uralica sp. nov." pageId="4" pageNumber="469">Table 1</tableCitation>
.
</paragraph>
<paragraph id="CC5636F5B95EFF809E7E0A6E7E5FFB62" blockId="4.[102,775,1143,1687]" box="[102,589,1143,1164]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809E7E0A6E7D56FB62" box="[102,324,1143,1164]" italics="true" pageId="4" pageNumber="469">Derivation of the name</emphasis>
: From the Urals, Latinised.
</paragraph>
<paragraph id="CC5636F5B95EFF809E7E0A817E96FB27" blockId="4.[102,775,1143,1687]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809E7E0A817CF1FB43" box="[102,227,1176,1197]" italics="true" pageId="4" pageNumber="469">Type locality</emphasis>
:
<collectingRegion id="0E2DF817B95EFF809EE80A817D29FB43" box="[240,315,1176,1197]" country="Russia" name="Kurgan" pageId="4" pageNumber="469">Kurgan</collectingRegion>
. The clay pit of the
<collectingRegion id="0E2DF817B95EFF809FE70A817E55FB43" box="[511,583,1176,1197]" country="Russia" name="Kurgan" pageId="4" pageNumber="469">Kurgan</collectingRegion>
brick factory in the northern part of
<materialsCitation id="7C813CA8B95EFF809F1A0AAD7E92FB27" box="[258,640,1204,1225]" country="Russia" location="South Trans-Urals" pageId="4" pageNumber="469" specimenCount="1" stateProvince="Kurgan">
<collectingRegion id="0E2DF817B95EFF809F1A0AAD7D58FB27" box="[258,330,1204,1225]" country="Russia" name="Kurgan" pageId="4" pageNumber="469">Kurgan</collectingRegion>
city,
<location id="C936602EB95EFF809F980AAD7E21FB27" LSID="urn:lsid:plazi:treatment:444087E3B95EFF829E7E0A30799CFA45:C936602EB95EFF809F980AAD7E21FB27" box="[384,563,1204,1225]" country="Russia" name="South Trans-Urals" pageId="4" pageNumber="469" stateProvince="Kurgan">South Trans−Urals</location>
,
<collectingCountry id="B4FE7665B95EFF809C250AAD7E92FB27" box="[573,640,1204,1225]" name="Russia" pageId="4" pageNumber="469">Russia</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="CC5636F5B95EFF809E7E0ACF7C80FAE8" blockId="4.[102,775,1143,1687]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809E7E0ACF7CF1FB05" box="[102,227,1238,1259]" italics="true" pageId="4" pageNumber="469">Type horizon</emphasis>
: Upper part of the gray−green clays of the Tavda Formation.
</paragraph>
<paragraph id="CC5636F5B95EFF809E7E0B0A7C8CFAAA" blockId="4.[102,775,1143,1687]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809E7E0B0A7C99FAC6" box="[102,139,1299,1320]" italics="true" pageId="4" pageNumber="469">Age</emphasis>
: Priabonian,
<emphasis id="FE9DEAE7B95EFF809F0B0B0A7E66FAC6" box="[275,628,1299,1320]" italics="true" pageId="4" pageNumber="469">
Charlesdowinea
<taxonomicName id="0BE94D76B95EFF809FA20B0A7E00FAC6" box="[442,530,1299,1320]" class="Dinophyceae" family="Peridiniaceae" genus="Charlesdowniea" higherTaxonomySource="GBIF" kingdom="Chromista" order="Peridiniales" pageId="4" pageNumber="469" phylum="Myzozoa" rank="species" species="clathrata">clathrata</taxonomicName>
angulosa
</emphasis>
Dinoflagellate Zone.
</paragraph>
<paragraph id="CC5636F5B95EFF809E7E0B497EA8FA8B" blockId="4.[102,775,1143,1687]" box="[102,698,1360,1381]" pageId="4" pageNumber="469">
<materialsCitation id="7C813CA8B95EFF809E7E0B497EA8FA8B" box="[102,698,1360,1381]" collectionCode="PIN" pageId="4" pageNumber="469" specimenCode="PIN 5088" specimenCount="1" typeStatus="holotype">
<emphasis id="FE9DEAE7B95EFF809E7E0B497CACFA8B" box="[102,190,1360,1381]" italics="true" pageId="4" pageNumber="469">
<typeStatus id="13528857B95EFF809E7E0B497CACFA8B" box="[102,190,1360,1381]" pageId="4" pageNumber="469" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCode id="9C4F9E8EB95EFF809ED20B497D3BFA8B" box="[202,297,1360,1381]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="4" pageNumber="469">PIN 5088</specimenCode>
/01, lower anterior−lateral tooth (
<figureCitation id="54D22A70B95EFF809C790B497EBDFA8B" box="[609,687,1360,1381]" captionStart="Fig" captionStartId="3.[102,135,1514,1534]" captionTargetBox="[102,1485,226,1486]" captionTargetId="figure-215@3.[102,1486,226,1486]" captionTargetPageId="3" captionText="Fig. 2. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. Holotype. PIN 5088/01, anterior tooth; in labial (A1), lingual (A2), labially−occlusal (A3), distally−basal (A4), lingually−occlusal (A5), and basal (A6) views. B. Paratype. PIN 5088/02, anterior tooth; in labial (B1), distally−basal (B2), lingual (B3), labially−occlusal (B4), lingually−occlusal (B5), and basal (B6) views. SEM microphotographs." figureDoi="http://doi.org/10.5281/zenodo.13651142" httpUri="https://zenodo.org/record/13651142/files/figure.png" pageId="4" pageNumber="469">Fig. 2A</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="CC5636F5B95EFF809E7E0B6B7EAAFA68" blockId="4.[102,775,1143,1687]" box="[102,696,1393,1415]" pageId="4" pageNumber="469">
<materialsCitation id="7C813CA8B95EFF809E7E0B6B7EA7FA68" box="[102,693,1393,1415]" collectionCode="PIN" pageId="4" pageNumber="469" specimenCode="PIN 5088" specimenCount="1" typeStatus="paratype">
<emphasis id="FE9DEAE7B95EFF809E7E0B6B7CACFA69" box="[102,190,1394,1415]" italics="true" pageId="4" pageNumber="469">
<typeStatus id="13528857B95EFF809E7E0B6B7CACFA69" box="[102,190,1394,1415]" pageId="4" pageNumber="469" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCode id="9C4F9E8EB95EFF809ED20B687D3AFA68" box="[202,296,1393,1414]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="4" pageNumber="469">PIN 5088</specimenCode>
/02, upper anterior−lateral tooth (
<figureCitation id="54D22A70B95EFF809C790B687EBEFA68" box="[609,684,1393,1414]" captionStart="Fig" captionStartId="3.[102,135,1514,1534]" captionTargetBox="[102,1485,226,1486]" captionTargetId="figure-215@3.[102,1486,226,1486]" captionTargetPageId="3" captionText="Fig. 2. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. Holotype. PIN 5088/01, anterior tooth; in labial (A1), lingual (A2), labially−occlusal (A3), distally−basal (A4), lingually−occlusal (A5), and basal (A6) views. B. Paratype. PIN 5088/02, anterior tooth; in labial (B1), distally−basal (B2), lingual (B3), labially−occlusal (B4), lingually−occlusal (B5), and basal (B6) views. SEM microphotographs." figureDoi="http://doi.org/10.5281/zenodo.13651142" httpUri="https://zenodo.org/record/13651142/files/figure.png" pageId="4" pageNumber="469">Fig. 2B</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CC5636F5B95EFF809E7E0B8E7E62FA41" blockId="4.[102,775,1143,1687]" box="[102,624,1430,1455]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809E7E0B8E7CDAFA41" box="[102,200,1431,1455]" italics="true" pageId="4" pageNumber="469">Material</emphasis>
. —
<collectingRegion id="0E2DF817B95EFF809EEB0B8E7D58FA41" box="[243,330,1431,1455]" country="Russia" name="Kurgan" pageId="4" pageNumber="469">Kurgan</collectingRegion>
: 45 teeth, Derney: 8 teeth.
</paragraph>
<paragraph id="CC5636F5B95EFF809E7E0BA679BEFA8E" blockId="4.[102,775,1143,1687]" lastBlockId="4.[812,1486,1064,1377]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809E7E0BA67CC7FA38" box="[102,213,1471,1494]" italics="true" pageId="4" pageNumber="469">Diagnosis</emphasis>
.—
<taxonomicName id="0BE94D76B95EFF809EEF0BA67DC9FA39" box="[247,475,1470,1495]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809EEF0BA67D90FA38" box="[247,386,1471,1494]" italics="true" pageId="4" pageNumber="469">Premontreia</emphasis>
species
</taxonomicName>
only known from isolated teeth. The teeth are up to 6.0 mm high. The principal cusps have rectilinear or almost rectilinear lateral edges, without a “waist”. The crowns, which bulge labio−lingually, are vertical in anterior teeth and slightly inclined in lateral teeth. The labial face of the crown is slightly convex, the lingual face is convex. The enameloid is smooth on the most of the teeth, sometimes with very short and barely distinct folds on the labial crown base. The cusplets are low, triangular in the labial view, closely set to the main cusp. The enameloid below the junction of cusp and cusplets is high, cutting edge is entire. The root is thick, protrudes over the crown, the basal face of the root is flat. The nutritive groove is wide and shallow. It divides the basal face of the root labio−lingually and appears lingually on the upper face of the root. There are number of small and some large foramina on the margino−lingual faces of the root. The central foramen is placed lingually in the nutritive groove.
</paragraph>
<paragraph id="CC5636F5B95EFF809D340B627FA4FA7A" blockId="4.[812,950,1403,1428]" box="[812,950,1403,1428]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809D340B627FA4FA7A" box="[812,950,1403,1428]" italics="true" pageId="4" pageNumber="469">Description</emphasis>
</paragraph>
<paragraph id="CC5636F5B95EFF809D340BB17FF5F86F" blockId="4.[812,1486,1448,1921]" pageId="4" pageNumber="469">
<emphasis id="FE9DEAE7B95EFF809D340BB17F81FA2E" box="[812,915,1448,1472]" italics="true" pageId="4" pageNumber="469">
<typeStatus id="13528857B95EFF809D340BB17F81FA2E" box="[812,915,1448,1472]" pageId="4" pageNumber="469" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="54D22A70B95EFF809DBC0BB17FEEFA2F" box="[932,1020,1448,1473]" captionStart="Fig" captionStartId="3.[102,135,1514,1534]" captionTargetBox="[102,1485,226,1486]" captionTargetId="figure-215@3.[102,1486,226,1486]" captionTargetPageId="3" captionText="Fig. 2. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. Holotype. PIN 5088/01, anterior tooth; in labial (A1), lingual (A2), labially−occlusal (A3), distally−basal (A4), lingually−occlusal (A5), and basal (A6) views. B. Paratype. PIN 5088/02, anterior tooth; in labial (B1), distally−basal (B2), lingual (B3), labially−occlusal (B4), lingually−occlusal (B5), and basal (B6) views. SEM microphotographs." figureDoi="http://doi.org/10.5281/zenodo.13651142" httpUri="https://zenodo.org/record/13651142/files/figure.png" pageId="4" pageNumber="469">Fig. 2A</figureCitation>
).—PIN 5088/01, a lower anterior−lateral tooth. The main cusp is triangular, slightly bent distally, with rectilinear mesial and distal edges and has a slight labio−lingual sigmoid twisting. The labial crown base has a series of short, centrally−placed, vertical folds. There is a pair of triangular cusplets, whose axis slightly laterally deviates from the main cusp; a second small cusplet is placed distally of the mesial cusplet. The root is thick, with a smooth contour, the branches are not well developed. The root protrudes behind the crown and forms the “pedestal” surrounding crown base. The nutritive groove is more wide than deep, slightly widened from the center of the basal face to the lingual side. There are some large nutritive foramina on the lingual face of the root and one large labially−situated central foramen in the nutritive groove.
</paragraph>
<caption id="9896667DB95EFF809E7E08A67D2AF91D" pageId="4" pageNumber="469" startId="4.[102,157,1727,1748]" targetType="table">
<paragraph id="CC5636F5B95EFF809E7E08A67D2AF91D" blockId="4.[102,775,1727,1779]" pageId="4" pageNumber="469">
Table 1. Measurements of the types series of
<taxonomicName id="0BE94D76B95EFF809C3108D97CF4F91D" authorityName="Malyshkina" authorityYear="2006" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="469" phylum="Chordata" rank="species" species="uralica" status="sp. nov." subGenus="Premontreia">
<emphasis id="FE9DEAE7B95EFF809C3108D97EB0F93B" box="[553,674,1728,1749]" italics="true" pageId="4" pageNumber="469">Premontreia</emphasis>
(
<emphasis id="FE9DEAE7B95EFF809CAB08D97C81F91D" italics="true" pageId="4" pageNumber="469">Premontreia</emphasis>
)
<emphasis id="FE9DEAE7B95EFF809EB908C77CF4F91D" box="[161,230,1758,1779]" italics="true" pageId="4" pageNumber="469">uralica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AE579CB95EFF809EF408C77D2AF91D" box="[236,312,1758,1779]" pageId="4" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="CC5636F5B95EFF809E7609077ECAF80C" pageId="4" pageNumber="469">
<table id="BEE9C455B95E007B9E7609087EF6F80C" box="[110,740,1809,2018]" gridcols="3" gridrows="6" pageId="4" pageNumber="469">
<tr id="72D934B7B95E007B9E7609087EF6F8AE" box="[110,740,1809,1856]" gridrow="0" pageId="4" pageNumber="469">
<th id="31085DCBB95E007B9E7609087DD6F8AE" box="[110,452,1809,1856]" gridcol="0" gridrow="0" pageId="4" pageNumber="469">Measurements</th>
<th id="31085DCBB95E007B9FEC09087E5FF8AE" box="[500,589,1809,1856]" gridcol="1" gridrow="0" pageId="4" pageNumber="469">Holotype (mm)</th>
<th id="31085DCBB95E007B9C8809087EF6F8AE" box="[656,740,1809,1856]" gridcol="2" gridrow="0" pageId="4" pageNumber="469">Paratype (mm)</th>
</tr>
<tr id="72D934B7B95E007B9E7609557EF6F88F" box="[110,740,1868,1889]" gridrow="1" pageId="4" pageNumber="469">
<th id="31085DCBB95E007B9E7609557DD6F88F" box="[110,452,1868,1889]" gridcol="0" gridrow="1" pageId="4" pageNumber="469">Tooth height</th>
<td id="31085DCBB95E007B9FEC09557E5FF88F" box="[500,589,1868,1889]" gridcol="1" gridrow="1" pageId="4" pageNumber="469">5.5</td>
<td id="31085DCBB95E007B9C8809557EF6F88F" box="[656,740,1868,1889]" gridcol="2" gridrow="1" pageId="4" pageNumber="469">4.7</td>
</tr>
<tr id="72D934B7B95E007B9E7609757EF6F86F" box="[110,740,1900,1921]" gridrow="2" pageId="4" pageNumber="469">
<th id="31085DCBB95E007B9E7609757DD6F86F" box="[110,452,1900,1921]" gridcol="0" gridrow="2" pageId="4" pageNumber="469">Tooth width</th>
<td id="31085DCBB95E007B9FEC09757E5FF86F" box="[500,589,1900,1921]" gridcol="1" gridrow="2" pageId="4" pageNumber="469">4.2</td>
<td id="31085DCBB95E007B9C8809757EF6F86F" box="[656,740,1900,1921]" gridcol="2" gridrow="2" pageId="4" pageNumber="469">3.6</td>
</tr>
<tr id="72D934B7B95E007B9E7609957EF6F84F" box="[110,740,1932,1953]" gridrow="3" pageId="4" pageNumber="469">
<th id="31085DCBB95E007B9E7609957DD6F84F" box="[110,452,1932,1953]" gridcol="0" gridrow="3" pageId="4" pageNumber="469">Crown base width</th>
<td id="31085DCBB95E007B9FEC09957E5FF84F" box="[500,589,1932,1953]" gridcol="1" gridrow="3" pageId="4" pageNumber="469">3.6</td>
<td id="31085DCBB95E007B9C8809957EF6F84F" box="[656,740,1932,1953]" gridcol="2" gridrow="3" pageId="4" pageNumber="469">3.8</td>
</tr>
<tr id="72D934B7B95E007B9E7609B47EF6F82C" box="[110,740,1965,1986]" gridrow="4" pageId="4" pageNumber="469">
<th id="31085DCBB95E007B9E7609B47DD6F82C" box="[110,452,1965,1986]" gridcol="0" gridrow="4" pageId="4" pageNumber="469">Cusp height</th>
<td id="31085DCBB95E007B9FEC09B47E5FF82C" box="[500,589,1965,1986]" gridcol="1" gridrow="4" pageId="4" pageNumber="469">4.25</td>
<td id="31085DCBB95E007B9C8809B47EF6F82C" box="[656,740,1965,1986]" gridcol="2" gridrow="4" pageId="4" pageNumber="469">0.75</td>
</tr>
<tr id="72D934B7B95E007B9E7609D47EF6F80C" box="[110,740,1997,2018]" gridrow="5" pageId="4" pageNumber="469">
<th id="31085DCBB95E007B9E7609D47DD6F80C" box="[110,452,1997,2018]" gridcol="0" gridrow="5" pageId="4" pageNumber="469">Height of the medial / distal cusplet</th>
<td id="31085DCBB95E007B9FEC09D47E5FF80C" box="[500,589,1997,2018]" gridcol="1" gridrow="5" pageId="4" pageNumber="469">1 / 0.7</td>
<td id="31085DCBB95E007B9C8809D47EF6F80C" box="[656,740,1997,2018]" gridcol="2" gridrow="5" pageId="4" pageNumber="469">0.7 / 0</td>
</tr>
</table>
</paragraph>
<paragraph id="CC5636F5B95EFF819D34098A7EE8F86C" blockId="4.[812,1485,1937,2026]" lastBlockId="5.[102,775,1609,2026]" lastPageId="5" lastPageNumber="470" pageId="4" pageNumber="469">
<materialsCitation id="7C813CA8B95EFF819D34098A7EE4F86C" collectionCode="PIN" lastPageId="5" lastPageNumber="470" pageId="4" pageNumber="469" specimenCode="PIN 5088" specimenCount="3" typeStatus="paratype">
<emphasis id="FE9DEAE7B95EFF809D34098A7F81F844" box="[812,915,1939,1962]" italics="true" pageId="4" pageNumber="469">
<typeStatus id="13528857B95EFF809D34098A7F81F844" box="[812,915,1939,1962]" pageId="4" pageNumber="469" type="paratype">Paratype</typeStatus>
</emphasis>
(
<figureCitation id="54D22A70B95EFF809DB009887810F844" box="[936,1026,1937,1962]" captionStart="Fig" captionStartId="3.[102,135,1514,1534]" captionTargetBox="[102,1485,226,1486]" captionTargetId="figure-215@3.[102,1486,226,1486]" captionTargetPageId="3" captionText="Fig. 2. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. Holotype. PIN 5088/01, anterior tooth; in labial (A1), lingual (A2), labially−occlusal (A3), distally−basal (A4), lingually−occlusal (A5), and basal (A6) views. B. Paratype. PIN 5088/02, anterior tooth; in labial (B1), distally−basal (B2), lingual (B3), labially−occlusal (B4), lingually−occlusal (B5), and basal (B6) views. SEM microphotographs." figureDoi="http://doi.org/10.5281/zenodo.13651142" httpUri="https://zenodo.org/record/13651142/files/figure.png" pageId="4" pageNumber="469">Fig. 2B</figureCitation>
).—
<specimenCode id="9C4F9E8EB95EFF809A34098878B1F844" box="[1068,1187,1937,1962]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="4" pageNumber="469">PIN 5088</specimenCode>
/02, upper anterior−lateral tooth. The main cusp is triangular, slightly bent distally, but upper third of the cusp is vertical. The main cusp has a weak labio−lingual sigmoid twisting. The mesial cusplet is less separated from the main cusp than that on the
<typeStatus id="13528857B95FFF819C6B08707ECAF96C" box="[627,728,1641,1666]" pageId="5" pageNumber="470" type="holotype">holotype</typeStatus>
, the tip is not sharp. The distal cusplet is reduced, there is just a smooth ledge. The root branches are more marked than in the
<typeStatus id="13528857B95FFF819E7E08D07CDEF90C" box="[102,204,1737,1762]" pageId="5" pageNumber="470" type="holotype">holotype</typeStatus>
: there are pronounced bulges, separated by the depression under the central part of the crown base (
<figureCitation id="54D22A70B95FFF819CB808F07EE4F8EC" box="[672,758,1769,1794]" captionStart="Fig" captionStartId="3.[102,135,1514,1534]" captionTargetBox="[102,1485,226,1486]" captionTargetId="figure-215@3.[102,1486,226,1486]" captionTargetPageId="3" captionText="Fig. 2. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. Holotype. PIN 5088/01, anterior tooth; in labial (A1), lingual (A2), labially−occlusal (A3), distally−basal (A4), lingually−occlusal (A5), and basal (A6) views. B. Paratype. PIN 5088/02, anterior tooth; in labial (B1), distally−basal (B2), lingual (B3), labially−occlusal (B4), lingually−occlusal (B5), and basal (B6) views. SEM microphotographs." figureDoi="http://doi.org/10.5281/zenodo.13651142" httpUri="https://zenodo.org/record/13651142/files/figure.png" pageId="5" pageNumber="470">Fig. 2B</figureCitation>
<figureCitation id="54D22A70B95FFF819CEE08ED7EEDF8EA" box="[758,767,1780,1796]" captionStart="Fig" captionStartId="1.[812,845,1193,1213]" captionTargetBox="[104,784,228,1287]" captionTargetId="graphics-387@1.[104,783,228,1286]" captionTargetPageId="1" captionText="Fig. 1. Geographical position of the localities where the studied material came from. A. The distribution of the Peri−Tethyan and West−Siberian waters in Priabonian times. B. Localities studied. C. Schematic local lithostratigaphic section." figureDoi="http://doi.org/10.5281/zenodo.13651140" httpUri="https://zenodo.org/record/13651140/files/figure.png" pageId="5" pageNumber="470">
<subScript id="506D34B0B95FFF819CEE08ED7EEDF8EA" attach="left" box="[758,767,1780,1796]" fontSize="7" pageId="5" pageNumber="470">1</subScript>
</figureCitation>
, B
<figureCitation id="54D22A70B95FFF819E60090D7C93F8CA" box="[120,129,1812,1828]" captionStart="Fig" captionStartId="5.[102,135,1459,1479]" captionTargetBox="[102,1485,230,1428]" captionTargetId="figure-285@5.[102,1486,230,1428]" captionTargetPageId="5" captionText="Fig. 4. Scyliorhinus zhelezkoi sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. Holotype. PIN 5088/03, lateral tooth; in labial (A1), lingual (A2), medial (A3), basal (A4), and lingually−occlusal (A5) views. B. Paratype. PIN 5088/04, anterior tooth; in labial (B1), labially−occlusal (B2), and lingually−mesial (B3) views; distal cusplets, enlargement (B4). SEM microphotographs." figureDoi="http://doi.org/10.5281/zenodo.13651152" httpUri="https://zenodo.org/record/13651152/files/figure.png" pageId="5" pageNumber="470">
<subScript id="506D34B0B95FFF819E60090D7C93F8CA" attach="left" box="[120,129,1812,1828]" fontSize="7" pageId="5" pageNumber="470">4</subScript>
</figureCitation>
). The nutritive groove is widened in the central part and almost closed on the central protuberance. There are some large and small nutritive foramina on the lingual surface of the root. The central foramen is large, and lingually situated
</materialsCitation>
.
</paragraph>
<caption id="9896667DB95FFF819E7E0BAA7F77FA11" ID-DOI="http://doi.org/10.5281/zenodo.13651152" ID-Zenodo-Dep="13651152" httpUri="https://zenodo.org/record/13651152/files/figure.png" pageId="5" pageNumber="470" startId="5.[102,135,1459,1479]" targetBox="[102,1485,230,1428]" targetPageId="5" targetType="figure">
<paragraph id="CC5636F5B95FFF819E7E0BAA7F77FA11" blockId="5.[102,1485,1459,1539]" pageId="5" pageNumber="470">
Fig. 4.
<taxonomicName id="0BE94D76B95FFF819EBD0BAA7D7CFA29" authorityName="Malyshkina" authorityYear="2006" box="[165,366,1459,1479]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="470" phylum="Chordata" rank="species" species="zhelezkoi" status="sp. nov.">
<emphasis id="FE9DEAE7B95FFF819EBD0BAA7D7CFA29" box="[165,366,1459,1479]" italics="true" pageId="5" pageNumber="470">Scyliorhinus zhelezkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AE579CB95FFF819F6A0BAD7DAEFA29" box="[370,444,1460,1479]" pageId="5" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
, Kurgan, Russia; Tavda Formation, Priabonian.
<emphasis id="FE9DEAE7B95FFF819D750BAA7F6FFA29" bold="true" box="[877,893,1459,1479]" pageId="5" pageNumber="470">A</emphasis>
. Holotype. PIN 5088/03, lateral tooth; in labial (A
<subScript id="506D34B0B95FFF819B5A0BA47958FA25" attach="right" box="[1346,1354,1469,1483]" fontSize="6" pageId="5" pageNumber="470">1</subScript>
), lingual (A
<subScript id="506D34B0B95FFF819BA00BA479D2FA25" attach="right" box="[1464,1472,1469,1483]" fontSize="6" pageId="5" pageNumber="470">2</subScript>
), medial (A
<subScript id="506D34B0B95FFF819EDC0BC07CDEFA09" attach="left" box="[196,204,1497,1511]" fontSize="6" pageId="5" pageNumber="470">3</subScript>
), basal (A
<subScript id="506D34B0B95FFF819F2A0BC07D28FA09" attach="right" box="[306,314,1497,1511]" fontSize="6" pageId="5" pageNumber="470">4</subScript>
), and lingually−occlusal (A
<subScript id="506D34B0B95FFF819C240BC07E56FA09" attach="left" box="[572,580,1497,1511]" fontSize="6" pageId="5" pageNumber="470">5</subScript>
) views.
<emphasis id="FE9DEAE7B95FFF819C800BD67EB5FA0C" bold="true" box="[664,679,1487,1506]" pageId="5" pageNumber="470">B</emphasis>
. Paratype. PIN 5088/04, anterior tooth; in labial (B
<subScript id="506D34B0B95FFF819A8B0BC07889FA09" attach="left" box="[1171,1179,1497,1511]" fontSize="6" pageId="5" pageNumber="470">1</subScript>
), labially−occlusal (B
<subScript id="506D34B0B95FFF819B7E0BC0797CFA09" attach="left" box="[1382,1390,1497,1511]" fontSize="6" pageId="5" pageNumber="470">2</subScript>
), and lingually−mesial (B
<subScript id="506D34B0B95FFF819EE50BEC7D17F9ED" attach="left" box="[253,261,1525,1539]" fontSize="6" pageId="5" pageNumber="470">3</subScript>
) views; distal cusplets, enlargement (B
<subScript id="506D34B0B95FFF819C720BEC7E60F9ED" attach="left" box="[618,626,1525,1539]" fontSize="6" pageId="5" pageNumber="470">4</subScript>
). SEM microphotographs.
</paragraph>
</caption>
<paragraph id="CC5636F5B95FFF819E7E098B784FF807" blockId="5.[102,775,1609,2026]" lastBlockId="5.[812,1485,1609,2026]" pageId="5" pageNumber="470">
<emphasis id="FE9DEAE7B95FFF819E7E098B7D18F844" box="[102,266,1938,1962]" italics="true" pageId="5" pageNumber="470">Anterior tooth</emphasis>
(
<figureCitation id="54D22A70B95FFF819F0509887D64F844" box="[285,374,1937,1962]" captionStart="Fig" captionStartId="4.[102,135,954,974]" captionTargetBox="[102,1485,229,924]" captionTargetId="figure-456@4.[102,1486,229,924]" captionTargetPageId="4" captionText="Fig. 3. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. 8CΠ/Ku−87/21, anterior tooth; in labial (A1), lingual (A2), occlusal (A3), distal (A4), and basal (A5) views. B. Lateral tooth; in labial (B1), lingual (B2), distal (B3), and occlusal (B4) views. Digital photographs." figureDoi="http://doi.org/10.5281/zenodo.13651148" httpUri="https://zenodo.org/record/13651148/files/figure.png" pageId="5" pageNumber="470">Fig. 3A</figureCitation>
).—IGG 8SP/Ku−87/21, shows the smooth enameloid on the both faces of the crown. There are one pair of the cusplets. The main cusp and distal cusplet are broken. The mesial cusplet is not high, acute−angled, slightly deviates laterally from the main cusp. The root is flat and thick, protrudes behind the crown and forms the “pedestal” round the crown. The basal face is flat. The root lobes are not developed, the root base in the labial view is straight. The nutritive groove is doubled (
<figureCitation id="54D22A70B95FFF819A4E08F578BDF8EB" box="[1110,1199,1772,1797]" captionStart="Fig" captionStartId="4.[102,135,954,974]" captionTargetBox="[102,1485,229,924]" captionTargetId="figure-456@4.[102,1486,229,924]" captionTargetPageId="4" captionText="Fig. 3. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. 8CΠ/Ku−87/21, anterior tooth; in labial (A1), lingual (A2), occlusal (A3), distal (A4), and basal (A5) views. B. Lateral tooth; in labial (B1), lingual (B2), distal (B3), and occlusal (B4) views. Digital photographs." figureDoi="http://doi.org/10.5281/zenodo.13651148" httpUri="https://zenodo.org/record/13651148/files/figure.png" pageId="5" pageNumber="470">Fig. 3A</figureCitation>
<figureCitation id="54D22A70B95FFF819AB708E178AAF8E6" box="[1199,1208,1784,1800]" captionStart="Fig" captionStartId="6.[102,135,1041,1061]" captionTargetBox="[102,1485,229,1009]" captionTargetId="figure-505@6.[102,1486,229,1009]" captionTargetPageId="6" captionText="Fig. 5. Scyliorhinus zhelezkoi sp. nov., Derney, Russia; Tavda Formation, Priabonian. IGG 8CΠ/D−03/01, lower lateral tooth; in labially−occlusal (A), lingual (B), occlusal (C), and lingually−occlusal (G) views; ribs on the labial side of the crown (D), lateral cusplet in the lingual view (E), ribs on the lingual side of the crown (F, H). SEM microphotographs." figureDoi="http://doi.org/10.5281/zenodo.13651154" httpUri="https://zenodo.org/record/13651154/files/figure.png" pageId="5" pageNumber="470">
<subScript id="506D34B0B95FFF819AB708E178AAF8E6" attach="left" box="[1199,1208,1784,1800]" fontSize="7" pageId="5" pageNumber="470">5</subScript>
</figureCitation>
), which is obviously an anomaly: there is a wide and shallow main nutritive groove displaced to the distal side of the root and another one almost equal to the main groove, but only the main groove extends on the labial and lingual faces of the root. There is a central foramen displaced lingually in the main groove and some smaller foramina in both grooves as well as on the lingual and labial faces of the root.
</paragraph>
<caption id="9896667DB95CFF829E7E0A087E3CFBB3" ID-DOI="http://doi.org/10.5281/zenodo.13651154" ID-Zenodo-Dep="13651154" httpUri="https://zenodo.org/record/13651154/files/figure.png" pageId="6" pageNumber="471" startId="6.[102,135,1041,1061]" targetBox="[102,1485,229,1009]" targetPageId="6" targetType="figure">
<paragraph id="CC5636F5B95CFF829E7E0A087E3CFBB3" blockId="6.[102,1485,1040,1117]" pageId="6" pageNumber="471">
Fig. 5.
<taxonomicName id="0BE94D76B95CFF829EBD0A087D7CFBCB" authorityName="Malyshkina" authorityYear="2006" box="[165,366,1041,1061]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="zhelezkoi" status="sp. nov.">
<emphasis id="FE9DEAE7B95CFF829EBD0A087D7CFBCB" box="[165,366,1041,1061]" italics="true" pageId="6" pageNumber="471">Scyliorhinus zhelezkoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AE579CB95CFF829F6B0A087DAEFBCA" box="[371,444,1041,1060]" pageId="6" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
, Derney, Russia; Tavda Formation, Priabonian. IGG 8CΠ/D−03/01, lower lateral tooth; in labially−occlusal (
<emphasis id="FE9DEAE7B95CFF829B930A097989FBCA" bold="true" box="[1419,1435,1040,1060]" pageId="6" pageNumber="471">A</emphasis>
), lingual (
<emphasis id="FE9DEAE7B95CFF829E820A347CBBFBAE" bold="true" box="[154,169,1069,1088]" pageId="6" pageNumber="471">B</emphasis>
), occlusal (
<emphasis id="FE9DEAE7B95CFF829F0A0A357D30FBAE" bold="true" box="[274,290,1068,1088]" pageId="6" pageNumber="471">C</emphasis>
), and lingually−occlusal (
<emphasis id="FE9DEAE7B95CFF829C120A357E0EFBAE" bold="true" box="[522,540,1068,1088]" pageId="6" pageNumber="471">G</emphasis>
) views; ribs on the labial side of the crown (
<emphasis id="FE9DEAE7B95CFF829DB50A347FAFFBAE" bold="true" box="[941,957,1069,1088]" pageId="6" pageNumber="471">D</emphasis>
), lateral cusplet in the lingual view (
<emphasis id="FE9DEAE7B95CFF829B100A347905FBAE" bold="true" box="[1288,1303,1069,1088]" pageId="6" pageNumber="471">E</emphasis>
), ribs on the lingual side of the crown (
<emphasis id="FE9DEAE7B95CFF829F080A507D0CFBB2" bold="true" box="[272,286,1097,1116]" pageId="6" pageNumber="471">F</emphasis>
,
<emphasis id="FE9DEAE7B95CFF829F310A507D29FBB2" bold="true" box="[297,315,1097,1116]" pageId="6" pageNumber="471">H</emphasis>
). SEM microphotographs.
</paragraph>
</caption>
<paragraph id="CC5636F5B95CFF829E7E0ABB7EE3F9F5" blockId="6.[102,776,1185,1564]" pageId="6" pageNumber="471">
<emphasis id="FE9DEAE7B95CFF829E7E0ABB7D33FB54" box="[102,289,1186,1210]" italics="true" pageId="6" pageNumber="471">The lateral tooth</emphasis>
(
<figureCitation id="54D22A70B95CFF829F290AB87D96FB54" box="[305,388,1185,1210]" captionStart="Fig" captionStartId="4.[102,135,954,974]" captionTargetBox="[102,1485,229,924]" captionTargetId="figure-456@4.[102,1486,229,924]" captionTargetPageId="4" captionText="Fig. 3. Premontreia uralica sp. nov., Kurgan, Russia; Tavda Formation, Priabonian. A. 8CΠ/Ku−87/21, anterior tooth; in labial (A1), lingual (A2), occlusal (A3), distal (A4), and basal (A5) views. B. Lateral tooth; in labial (B1), lingual (B2), distal (B3), and occlusal (B4) views. Digital photographs." figureDoi="http://doi.org/10.5281/zenodo.13651148" httpUri="https://zenodo.org/record/13651148/files/figure.png" pageId="6" pageNumber="471">Fig. 3B</figureCitation>
).—IGG 8SP/Ku−87/22, shows the smooth enameloid with the slight striation on the base of the labial face. The labial face of the crown is convex, the lingual face is very convex. The main cusp is bent distally and has a strong labial−lingual bend. There is one pair of the cusplets. The cusplets are small, not sharp, closely set to the main cusp. The root has a flat base. The contour of the root base in the labial view as well as the line of the root−crown connection is arched. The root lobes are not well developed. There is a well−developed nutritive groove, slightly widened in the central part of the basal face of the root. There are some foramina of the oval shape on the labial and lingual faces of the root.
</paragraph>
<paragraph id="CC5636F5B95CFF829E7E082B7D3CF804" blockId="6.[102,775,1585,2026]" pageId="6" pageNumber="471">
<emphasis id="FE9DEAE7B95CFF829E7E082B7D81F9A4" box="[102,403,1586,1610]" italics="true" pageId="6" pageNumber="471">Comparisons and remarks</emphasis>
.—The differences between
<taxonomicName id="0BE94D76B95CFF829CCD082A7D3AF984" authorityName="Malyshkina" authorityYear="2006" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="uralica" status="sp. nov.">
<emphasis id="FE9DEAE7B95CFF829CCD082A7D3AF984" italics="true" pageId="6" pageNumber="471">Premontreia uralica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AE579CB95CFF829F2C084B7D80F987" box="[308,402,1618,1641]" pageId="6" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="0BE94D76B95CFF829FCB084A7E7DF984" baseAuthorityName="Arambourg" baseAuthorityYear="1952" box="[467,623,1618,1642]" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="subulidens">
<emphasis id="FE9DEAE7B95CFF829FCB084A7E7DF984" box="[467,623,1618,1642]" italics="true" pageId="6" pageNumber="471">P. subulidens</emphasis>
</taxonomicName>
(
<bibRefCitation id="A8784B04B95CFF829C9B08487CC5F964" author="Arambourg, C." pageId="6" pageNumber="471" pagination="1 - 372" refId="ref7618" refString="Arambourg, C. 1952. Les vertebres fossiles des gisements de phosphates (Maroc - Algerie - Tunesie). Notes et Memoires. Service des Mines et de la Carte Geologique du Maroc 92: 1 - 372. Paris." type="journal article" year="1952">Arambourg 1952: 121</bibRefCitation>
, pl. 23: 5–19; Noubhani and Cappetta 1997: 55, pl. 21) are that the teeth of
<taxonomicName id="0BE94D76B95CFF829F62088A7DF9F944" authorityName="Malyshkina" authorityYear="2006" box="[378,491,1682,1706]" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="uralica">
<emphasis id="FE9DEAE7B95CFF829F62088A7DF9F944" box="[378,491,1682,1706]" italics="true" pageId="6" pageNumber="471">P. uralica</emphasis>
</taxonomicName>
have: (1) labio−lingually and medio−distally thicker crowns, so that the contour of the main cusp is a triangle with a wider base; (2) rectilinear or subrectilinear mesial and distal edges of the cusp; (3) distinctive enameloid ornamentation on the labial crown base; (4) shorter and less acute−angled cusplets; their axes more deviate from the cusp; (5) the root is more massive and high. The same differences apply to
<taxonomicName id="0BE94D76B95CFF829F87096A7E53F864" authorityName="Cappetta" authorityYear="1992" box="[415,577,1906,1930]" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="degremonti">
<emphasis id="FE9DEAE7B95CFF829F87096A7E53F864" box="[415,577,1906,1930]" italics="true" pageId="6" pageNumber="471">P. degremonti</emphasis>
</taxonomicName>
(
<bibRefCitation id="A8784B04B95CFF829C4D09687C81F844" author="Cappetta, H." pageId="6" pageNumber="471" pagination="639 - 646" refId="ref7779" refString="Cappetta, H. 1992. Carcharhiniformes nouveaux (Chondrichthyes, Neoselachii) de l'Ypresian du bassin de Paris. Geobios 25 (5): 639 - 646." type="journal article" year="1992">Cappetta 1992: 640</bibRefCitation>
, pl. 1), except for the cusplet characters and that
<taxonomicName id="0BE94D76B95CFF829CF6098A7CF7F824" authorityName="Cappetta" authorityYear="1992" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="degremonti">
<emphasis id="FE9DEAE7B95CFF829CF6098A7CF7F824" italics="true" pageId="6" pageNumber="471">P. degremonti</emphasis>
</taxonomicName>
has a more intensive striation near the base of its labial crown face.
</paragraph>
<paragraph id="CC5636F5B95CFF829D4A0ABB79A9FAF4" blockId="6.[812,1486,1185,1405]" pageId="6" pageNumber="471">
<taxonomicName id="0BE94D76B95CFF829D4A0ABB796CFB54" authority="(Casier, 1946)" baseAuthorityName="Casier" baseAuthorityYear="1946" box="[850,1406,1185,1210]" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="gilberti" subGenus="Oxyscyllium">
<emphasis id="FE9DEAE7B95CFF829D4A0ABB7FCFFB57" box="[850,989,1186,1209]" italics="true" pageId="6" pageNumber="471">Premontreia</emphasis>
(
<emphasis id="FE9DEAE7B95CFF829DF60ABB7867FB54" box="[1006,1141,1186,1210]" italics="true" pageId="6" pageNumber="471">Oxyscyllium</emphasis>
)
<emphasis id="FE9DEAE7B95CFF829A9F0ABB78C5FB54" box="[1159,1239,1186,1210]" italics="true" pageId="6" pageNumber="471">gilberti</emphasis>
(Casier, 1946)
</taxonomicName>
differs from the
<taxonomicName id="0BE94D76B95CFF829D880ADB7FEFFB34" authorityName="Malyshkina" authorityYear="2006" box="[912,1021,1218,1242]" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="uralica" status="sp. nov.">
<emphasis id="FE9DEAE7B95CFF829D880ADB7FEFFB34" box="[912,1021,1218,1242]" italics="true" pageId="6" pageNumber="471">P. uralica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5AE579CB95CFF829A1C0ADB7848FB37" box="[1028,1114,1218,1241]" pageId="6" pageNumber="471" rank="species">sp. nov.</taxonomicNameLabel>
by more intensive striation on the labial face of the crown and overhanging of the root crown labial face, as well as by narrower main cusp in the lower teeth.
</paragraph>
<paragraph id="CC5636F5B95CFF829D4A0B3C7FDCFA93" blockId="6.[812,1486,1185,1405]" pageId="6" pageNumber="471">
<taxonomicName id="0BE94D76B95CFF829D4A0B3C79B0FAD3" authority="(Lawley, 1876)" baseAuthorityName="Lawley" baseAuthorityYear="1876" box="[850,1442,1316,1341]" class="Elasmobranchii" family="Scyliorhinidae" genus="Premontreia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="471" phylum="Chordata" rank="species" species="dachiardi" subGenus="Oxyscyllium">
<emphasis id="FE9DEAE7B95CFF829D4A0B3C7FCFFAD2" box="[850,989,1317,1340]" italics="true" pageId="6" pageNumber="471">Premontreia</emphasis>
(
<emphasis id="FE9DEAE7B95CFF829DF50B3D7866FAD2" box="[1005,1140,1316,1340]" italics="true" pageId="6" pageNumber="471">Oxyscyllium</emphasis>
)
<emphasis id="FE9DEAE7B95CFF829A9D0B3D78E2FAD2" box="[1157,1264,1316,1340]" italics="true" pageId="6" pageNumber="471">dachiardi</emphasis>
(Lawley, 1876)
</taxonomicName>
has a smooth enameloid overhanging the root labially and a different root shape.
</paragraph>
<paragraph id="CC5636F5B95CFF829D340B8B799CFA45" blockId="6.[812,1422,1426,1451]" box="[812,1422,1426,1451]" pageId="6" pageNumber="471">
<emphasis id="FE9DEAE7B95CFF829D340B8B7FA7FA44" box="[812,949,1426,1450]" italics="true" pageId="6" pageNumber="471">Distribution</emphasis>
.—Priabonian of the West−Siberian Basin.
</paragraph>
</subSubSection>
</treatment>
</document>
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