370 lines
37 KiB
XML
370 lines
37 KiB
XML
<document id="0C7E1C300C1080B66390AB89807DA256" ID-CLB-Dataset="298601" ID-DOI="10.3897/zse.100.120207" ID-GBIF-Dataset="6f7f44bf-fab1-47dd-aa0a-d5ad482b207e" ID-ZooBank="55DBF63A-85CF-42C0-8218-15F310FB177A" ID-publisher-id="120207" URI-arpha="123D4283-0C8F-5E2E-9E65-4D0700BDBA20" URI-zoobank="http://zoobank.org/55DBF63A-85CF-42C0-8218-15F310FB177A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1718641467983" checkinUser="pensoft" docAuthor="Villarreal, Osvaldo, Ahumada-C., Daniela & Navas-S., Gabriel R." docDate="2024" docId="F09894290F60556E9F8E8E43D7A1F2CE" docLanguage="en" docName="Zoosystematics and Evolution 100(3): 803-820" docOrigin="Zoosystematics and Evolution 100 (3)" docSource="https://zse.pensoft.net/article/120207/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Eutimesius aroa Villarreal" docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="820" masterDocId="123D42830C8F5E2E9E654D0700BDBA20" masterDocTitle="Exploring the diversity of Eutimesius Roewer, 1913: new species and records from Colombia and Venezuela (Opiliones, Gonyleptoidea, Stygnidae)" masterLastPageNumber="820" masterPageNumber="803" pageNumber="803" updateTime="1718651764718" updateUser="ExternalLinkService">
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<mods:title id="3B4D2901EB4F6712FD448F3690B5D7CC">Exploring the diversity of Eutimesius Roewer, 1913: new species and records from Colombia and Venezuela (Opiliones, Gonyleptoidea, Stygnidae)</mods:title>
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<mods:name id="E46CF17B68F543AC32859DE48C02579D" type="personal">
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<mods:namePart id="74C4C5B4CC4DD8928F4C9E9D2759B861">Villarreal, Osvaldo</mods:namePart>
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<mods:affiliation id="497ED3445B06EAC555BFB6291752CEEE">Centro de Ecología, Instituto Venezolano de Investigaciones Científicas (IVIC), km 11 carretera Panamericana, Altos de Pipe, edo. Miranda 1204 - A, Venezuela & Instituto y Museo del Instituto de Zoología Agrícola, Facultad de Agronomía, Universidad Central de Venezuela, Apartado 4579, Maracay 2101, Aragua, Venezuela & Departamento de Invertebrados, Museu Nacional / UFRJ, Quinta da Boa Vista, São Cristóvão, 20.940 - 040, Rio de Janeiro – RJ, Brazil</mods:affiliation>
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<mods:namePart id="514B76B61BE864EFE89DA553C4016BA2">Ahumada-C., Daniela</mods:namePart>
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<mods:nameIdentifier id="F41BFBD125BB967CA7680752BBCDF181" type="ORCID">0000-0002-4182-5143</mods:nameIdentifier>
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<mods:affiliation id="69E0E9509D1360BBA0625466AA1546A1">Grupo de Investigación Biología Descriptiva y Aplicada, Programa de Biología, Universidad de Cartagena, Zaragocilla Cra. 50 # 24 - 120, Cartagena de Indias, Bolívar, Colombia</mods:affiliation>
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<mods:namePart id="0F32B30AFE823243FB1EB5479FC76561">Navas-S., Gabriel R.</mods:namePart>
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<mods:nameIdentifier id="1985A0D830C6F7E76ACC3CF51D1E8794" type="ORCID">0000-0001-9554-6345</mods:nameIdentifier>
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<mods:affiliation id="E3810FDC9AF3775A28A75714D24DCD74">Grupo de Investigación Biología Descriptiva y Aplicada, Programa de Biología, Universidad de Cartagena, Zaragocilla Cra. 50 # 24 - 120, Cartagena de Indias, Bolívar, Colombia</mods:affiliation>
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<mods:typeOfResource id="8EA01090F69F1F7052697223605CA4A1">text</mods:typeOfResource>
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<mods:title id="5BAAA47CBA3D1B7E47C0A1D3E0DC714E">Zoosystematics and Evolution</mods:title>
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<mods:part id="5CFBB18AB9F916A618DB9C150509678D">
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<mods:date id="73EDE2104EF51401E0215C8572EA8021">2024</mods:date>
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<mods:number id="61774899B2DA93BE22A26B22D45A2DF5">2024-06-17</mods:number>
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<mods:number id="875D173FBEB271BF26D61E60194CFE76">100</mods:number>
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<mods:detail id="5E3DCA502597D0C14F9A0F9843D21677" type="issue">
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<mods:identifier id="C739E8B04814DFAFDCA24E0A11595973" type="CLB-Dataset">298601</mods:identifier>
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<mods:identifier id="A7BF17AE8CDFE2A07CF9FCEDA66EB9F8" type="DOI">10.3897/zse.100.120207</mods:identifier>
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<treatment id="F09894290F60556E9F8E8E43D7A1F2CE" ID-DOI="http://doi.org/10.5281/zenodo.11975103" ID-GBIF-Taxon="233216496" ID-Zenodo-Dep="11975103" ID-arpha="F0989429-0F60-556E-9F8E-8E43D7A1F2CE" ID-zoobank="https://zoobank.org/C957434C-5804-4610-88B7-8966D3A24FF7" LSID="urn:lsid:plazi:treatment:F09894290F60556E9F8E8E43D7A1F2CE" LSID-ZBK="urn:lsid:zoobank.org:act:C957434C-5804-4610-88B7-8966D3A24FF7" httpUri="http://treatment.plazi.org/id/F09894290F60556E9F8E8E43D7A1F2CE">
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||
<subSubSection id="D9EAB6FF6F1204733314E7B6415BA8EB" type="nomenclature">
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<paragraph id="C55455C4250E659BEA684B54472B00A9">
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<taxonomicName id="513D2398DC80805E754A50476CDBC3D3" authority="Villarreal" authorityName="Villarreal" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="aroa">
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<emphasis id="EC637718ECD1788AE878213050AAE70A" italics="true">Eutimesius aroa</emphasis>
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Villarreal
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</taxonomicName>
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& Ahumada-C. sp. nov.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="7D7D31B6B2A8B8B87539CD5F206DBA1E" type="description">
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||
<paragraph id="CBAF18A1294B488BCAE34B1A152C2A76">
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<figureCitation id="8733E6540C660B0F87C050CF834CD46E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eutimesius aroa sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105934); C, D. Female habitus, dorsal view (MIZA 0105936); C. Male habitus, lateral view (MIZA 0105934). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure4" httpUri="https://binary.pensoft.net/fig/1072632">Figs 4</figureCitation>
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,
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<figureCitation id="AE4E75AED9449A9423BD6A1BAF4D846D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutimesius aroa sp. nov., male (MIZA 0105934): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure5" httpUri="https://binary.pensoft.net/fig/1072633">5</figureCitation>
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,
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<figureCitation id="7FC1670951E52640633B00D914EFE4A9" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eutimesius spp., penis, distal portion in dorsal, ventral, and lateral view: A – C. E. aroa sp. nov., (MIZA 0105934); D – F. E. canoabo sp. nov., (MIZA 0105945); G – I. E. ephippiatus, (ICN-Ao - 1492); J – L. E. guaichia sp. nov., (MIZA 0105935); M – O. E. simoni, (ICN-Ao - 328). A – I. Unscaled." figureDoi="10.3897/zse.100.120207.figure14" httpUri="https://binary.pensoft.net/fig/1072642">14 A – C</figureCitation>
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,
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<figureCitation id="10EE05FFE969559203310476AF247FA6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map showing the distribution of Eutimesius." figureDoi="10.3897/zse.100.120207.figure2" httpUri="https://binary.pensoft.net/fig/1072630">2</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="SECID0EH3AE" type="type material">
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||
<paragraph id="17ADC78075F9010A69CCE044D6CCB24D">
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||
<heading id="2E71A24C08E8FF0CC5E8DBFA897C44A5" reason="title">Type material.</heading>
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||
</paragraph>
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||
<paragraph id="1224032ABBFB2640F5DAAE68107FB1AD">
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<materialsCitation id="EBE0685ABE04F09B94BC39D92900E18A" ID-GBIF-Occurrence="4879930307" collectingDate="2008-03-09" collectionCode="MIZA" collectorName="Villarreal O. & Escalona H. & Jayaro Y. & Viera E." country="Venezuela" elevation="1200" latitude="10.352" location="Las Cumaraguas" longLatPrecision="7" longitude="-68.8298" municipality="Road Cocorote - Aroa" specimenCode="MIZA 0105934" specimenCount="1" specimenCount-male="1" stateProvince="Yaracuy" typeStatus="holotype">
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||
<collectingCountry id="8C60660A5FEED0829A61C885BD349EC6" name="Venezuela">Venezuela</collectingCountry>
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•
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<specimenCount id="6215BA9DACBC6E94DBC01E1014B3C7AC" count="1" type="male">♂</specimenCount>
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||
<emphasis id="DF86F63C3818F66E4BFA48F94A80640A" bold="true" italics="true">
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||
<typeStatus id="2A3AC0B4C93DD754C76D0499E842F3A7" type="holotype">holotype</typeStatus>
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||
</emphasis>
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;
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<collectingRegion id="609864B38BC6A03E50B98C045F39041D" country="Venezuela" name="Yaracuy">Yaracuy</collectingRegion>
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,
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<collectingMunicipality id="A3C6FEAB183E827A6D5DB793046FFE13">Road Cocorote – Aroa</collectingMunicipality>
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, sector
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<location id="419E466375F8A38B421C69ECC989FB3D" LSID="urn:lsid:plazi:treatment:F09894290F60556E9F8E8E43D7A1F2CE:419E466375F8A38B421C69ECC989FB3D" country="Venezuela" latitude="10.352" longLatPrecision="7" longitude="-68.8298" municipality="Road Cocorote - Aroa" name="Las Cumaraguas" stateProvince="Yaracuy">Las Cumaraguas</location>
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; (
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0ET3AE" specific-use="{"type":"Point","coordinates":[-68.829800,10.352000]}">
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||
<geoCoordinate id="D83780BD17FA2307CAF74A303DF5A77A" degrees="10.3520" direction="north" orientation="latitude" precision="5" value="10.352">10.3520 ° N</geoCoordinate>
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,
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<geoCoordinate id="19ED90DF7DA40C0BCFE319ED86005888" degrees="68.8298" direction="west" orientation="longitude" precision="5" value="-68.8298">68.8298 ° W</geoCoordinate>
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</named-content>
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||
);
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<elevation id="8427C488646312B49A301A4915997731" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
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<quantity id="AFCD195DB8E53F230C8E9A5D73D32D42" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1,200 m</quantity>
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a. s. l.
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</elevation>
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;
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<date id="B7CC70E8DDD26D882BEEB529414A48F2" value="2008-03-09">
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<collectingDate id="8FCAE817A4EC6D4EBAB62D36009B516C" value="2008-03-09">09 Mar. 2008</collectingDate>
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</date>
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; (
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<collectorName id="E099BBC289DCF9A8434841D74CDE90C7">Villarreal O.</collectorName>
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,
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<collectorName id="A68E8541C02C9FF92B4952698CF5F201">Escalona H.</collectorName>
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,
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<collectorName id="1F3338F925D86E10D0CBAEABA82237F9">Jayaro Y.</collectorName>
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,
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<collectorName id="BF9D056CC2503230064B0F5BD9858F2A">Viera E.</collectorName>
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leg.) (
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<specimenCode id="817A842BD5B495D71C494B8AEF596E4A" collectionCode="MIZA" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" type="Museum">
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||
<named-content id="C12084FB24B03BD9724AEC67CCE6808F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
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0105934
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</specimenCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="8EB9E5567AE601C63562053A83F9FF33" ID-GBIF-Occurrence="4879930309" collectingDate="2008-03-09" collectionCode="MIZA" collectorName="Villarreal O. & Escalona H. & Jayaro Y. & Viera E. & Paratypes" country="Venezuela" elevation="1200" latitude="10.352" location="Las Cumaraguas" longLatPrecision="7" longitude="-68.8298" municipality="Road Cocorote - Aroa" specimenCode="MIZA 0105936" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yaracuy" typeStatus="paratype">
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||
<emphasis id="FEAC16B8EC6FD843F3818478A5246F62" bold="true" italics="true">
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<typeStatus id="E7AB36AE5D28508CB694CEE0C0F4C1A2" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
•
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<specimenCount id="C293223F2E9EE5102B1DD6999C65354D" count="1" type="female">1 ♀</specimenCount>
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,
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||
<specimenCount id="EB8133E5E457939DDDC1E7185FD80D63" count="1" type="male">1 ♂</specimenCount>
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; same as the holotype; (
|
||
<specimenCode id="E581E78D9137DF4A4AA8F582B117CC17" collectionCode="MIZA" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" type="Museum">
|
||
<named-content id="FBFEF0CDA62B5CCA4E13776DBD3A4B71" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
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0105936
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||
</specimenCode>
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||
)
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||
</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="SECID0EF4AE" type="diagnosis">
|
||
<paragraph id="F5B8EEA008AEA504BB3530FD50C01B89">
|
||
<heading id="A690F24238608518A1DB581B94C485BA" reason="title">Diagnosis.</heading>
|
||
</paragraph>
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<paragraph id="56A6DFE177452010CDFBDB4C87E6E382">
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It is distinguishable from all other species in the genus by the pattern of dry white spots, occupying the anterolateral zone of the carapace, lateral areas of the II and III areas, and part of the lateral margins of the dorsal shield (Figs
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<figureCitation id="C114E78F5648ED892EC52E4F109F0EFB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eutimesius aroa sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105934); C, D. Female habitus, dorsal view (MIZA 0105936); C. Male habitus, lateral view (MIZA 0105934). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure4" httpUri="https://binary.pensoft.net/fig/1072632">4 A, C</figureCitation>
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||
,
|
||
<figureCitation id="C7439EE8702119459C6C742B9E105F82" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutimesius aroa sp. nov., male (MIZA 0105934): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure5" httpUri="https://binary.pensoft.net/fig/1072633">5 A, B</figureCitation>
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||
), and by the ornamentation of the male femur IV, with large tubercles on both complete ventral rows (Fig.
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<figureCitation id="2E3EA7E56B3D270348EB7074D1E75C9B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutimesius aroa sp. nov., male (MIZA 0105934): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure5" httpUri="https://binary.pensoft.net/fig/1072633">5 G</figureCitation>
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||
).
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</paragraph>
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<caption id="BF989242CD9A4C230625F15D2A9A2BC3" ID-DOI="10.3897/zse.100.120207.figure4" ID-arpha="566D4771-DA54-5299-9865-2B41A5EA12F5" httpUri="https://binary.pensoft.net/fig/1072632" startId="F4">
|
||
<paragraph id="B3F02C2435898B8F0312529274DBD1B9">
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||
<label id="E31CCA11759BA9597C3D811349C02DB9">Figure 4.</label>
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</paragraph>
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<paragraph id="54A76169BAEF71C3C50853B46DF4CB46">
|
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<taxonomicName id="0D5A5D761D1C9AEF2846BE303CB07FE1" authority="Villarreal & Ahumada-C. & Navas-S., 2024" authorityName="Villarreal & Ahumada-C. & Navas-S." authorityYear="2024" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="aroa" status="sp. nov.">
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<emphasis id="5C714AF4F955B483920D4E949D8A6753" italics="true">Eutimesius aroa</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="83CB18B1042550E35B1866E6DC16DCDE" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="46F1D20F0F67837F9D986D1607EF93AD" bold="true">A, B.</emphasis>
|
||
Male habitus, dorsal and lateral views (
|
||
<specimenCode id="824FE91ECC99A6B3800CC36EB6ACAFE9">
|
||
<named-content id="5D092E7D381EEDFAC2E61597D464ED83" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
|
||
0105934
|
||
</specimenCode>
|
||
);
|
||
<emphasis id="EE90AFD631E76BF7711AFF544979079D" bold="true">C, D.</emphasis>
|
||
Female habitus, dorsal view (
|
||
<specimenCode id="4501CC33FCE2847FE4A0E00E2843D337">
|
||
<named-content id="A014DA094DDC096399C20B1357FE2FAF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
|
||
0105936
|
||
</specimenCode>
|
||
);
|
||
<emphasis id="8D725B5BC2039F5E7EB0B9185128267B" bold="true">C.</emphasis>
|
||
Male habitus, lateral view (
|
||
<specimenCode id="EB0F6BC9C137F5AF39A24CF80947E289">
|
||
<named-content id="A1454189D364F43D629FB6CD8C33AE52" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
|
||
0105934
|
||
</specimenCode>
|
||
). Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6022F9F19A44734B613B87F28484ADC1" ID-DOI="10.3897/zse.100.120207.figure5" ID-arpha="A24B1000-7974-5BD4-9623-FB02917B5032" httpUri="https://binary.pensoft.net/fig/1072633" startId="F5">
|
||
<paragraph id="798176D0F6D4A81624FB71EC1F0E75F2">
|
||
<label id="A89C2F95324F208DAD7503E0B0F1E093">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="1D7B1A9A3641D43245A1A019649CED98">
|
||
<taxonomicName id="0242B80C924BBD610610CDD51DCC9F39" authority="Villarreal & Ahumada-C. & Navas-S., 2024" authorityName="Villarreal & Ahumada-C. & Navas-S." authorityYear="2024" class="Arachnida" family="Stygnidae" genus="Eutimesius" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="species" species="aroa" status="sp. nov.">
|
||
<emphasis id="A542E3C2CCB3947F72C37A6228F36058" italics="true">Eutimesius aroa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19DA791B7FAF7820B811D7A44B371F81" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male (
|
||
<specimenCode id="A98E4E5605858D0819154F3C6E99E8BB">
|
||
<named-content id="CB3BB0C08B9C93B4B7F94E845E7939C2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-del-instituto-de-zoologia-agricola-francisco-fernandez-yepez" xlink_title="Museo del Instituto de Zoología Agrícola “Francisco Fernández Yépez”, Maracay, Venezuela">MIZA</named-content>
|
||
0105934
|
||
</specimenCode>
|
||
):
|
||
<emphasis id="DE99B56B9A204674F72B987F72860630" bold="true">A.</emphasis>
|
||
Habitus, dorsal view;
|
||
<emphasis id="63E13F88011B0191124565925B64B7E1" bold="true">B.</emphasis>
|
||
Ditto, lateral view;
|
||
<emphasis id="27799B0E9CEF3ACFA66A5C970622CBBB" bold="true">C.</emphasis>
|
||
Right chelicera, frontal view;
|
||
<emphasis id="D2C1737270ACDF17DAC05038F1F0BE8F" bold="true">D.</emphasis>
|
||
Right pedipalp, tibia, and tarsus, ectal view;
|
||
<emphasis id="5E3AFB7E179744669E74D513FD2513C5" bold="true">E.</emphasis>
|
||
Ditto, mesal view;
|
||
<emphasis id="C7E8895891D7A4E540E30BC82F755114" bold="true">F.</emphasis>
|
||
Right leg IV, dorsal view;
|
||
<emphasis id="242A45B667C6551E7C1CB25F7DB956FB" bold="true">G.</emphasis>
|
||
Ditto, ventral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EQAAG" type="etymology">
|
||
<paragraph id="523F0E610D44F579DA24B27153A58068">
|
||
<heading id="EB71552AAAAAA93E8A9EBA120AE28B2A" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="B7C4044ECC99C29ABE3058140753B501">The species is named in reference to the Sierra de Aroa, the mountain range in which it inhabits. Aroa is a Chibcha word believed to have a meaning associated with the jaguar, or “ tigre. ” Noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EVAAG" type="description">
|
||
<paragraph id="D7A93453DCB882F9BF9A44DF344DA3B7">
|
||
<heading id="A85D38E7D79D854C6E3DBE33FD478C1D" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="D24C225A67901233308961ECFB2D3D9B">
|
||
<emphasis id="F7FFC727634B508C8CFE957367585311" bold="true">
|
||
Male.
|
||
<emphasis id="EBDA8E92573901132220896BB69682C0" bold="true" italics="true">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
<abbrev id="ABBRID0EABAG" xlink_title="dorsal scutum length">DSL</abbrev>
|
||
4.0;
|
||
<abbrev id="ABBRID0EEBAG" xlink_title="dorsal scutum width">DSW</abbrev>
|
||
3.5;
|
||
<abbrev id="ABBRID0EIBAG" xlink_title="maximum abdominal scutum length">AL</abbrev>
|
||
1.9;
|
||
<abbrev id="ABBRID0EMBAG" xlink_title="maximum abdominal scutum width">AW</abbrev>
|
||
2.7;
|
||
<abbrev id="ABBRID0EQBAG" xlink_title="interocular distance">IOD</abbrev>
|
||
2.0; pedipalp:
|
||
<abbrev id="ABBRID0EUBAG" xlink_title="pedipalpal coxa">CoPp</abbrev>
|
||
0.8, TrPp 1.1,
|
||
<abbrev id="ABBRID0EYBAG" xlink_title="pedipalpal femur">FePp</abbrev>
|
||
3.8,
|
||
<abbrev id="ABBRID0E3BAG" xlink_title="pedipalpal patella">PaPp</abbrev>
|
||
1.8,
|
||
<abbrev id="ABBRID0EACAG" xlink_title="pedipalpal tibia">TiPp</abbrev>
|
||
2.4,
|
||
<abbrev id="ABBRID0EECAG" xlink_title="pedipalpal tarsus">TaPp</abbrev>
|
||
2.2,
|
||
<abbrev id="ABBRID0EICAG" xlink_title="pedipalp claw">ClPp</abbrev>
|
||
1.8; total 13.9; leg IV:
|
||
<abbrev id="ABBRID0EMCAG" xlink_title="femur length">FeL</abbrev>
|
||
7.2,
|
||
<abbrev id="ABBRID0EQCAG" xlink_title="tibia length">TiL</abbrev>
|
||
3.9.
|
||
<emphasis id="DDCC40B79A3CE719543EBD5F7F9FE37E" bold="true" italics="true">Dorsum</emphasis>
|
||
(Figs
|
||
<figureCitation id="8CD76E29D565C5E10A6A25CB487F1CD0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eutimesius aroa sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105934); C, D. Female habitus, dorsal view (MIZA 0105936); C. Male habitus, lateral view (MIZA 0105934). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure4" httpUri="https://binary.pensoft.net/fig/1072632">4 A, C</figureCitation>
|
||
,
|
||
<figureCitation id="9472028CEC4C37BD76CC2A5B74D64712" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutimesius aroa sp. nov., male (MIZA 0105934): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure5" httpUri="https://binary.pensoft.net/fig/1072633">5 A, B</figureCitation>
|
||
).
|
||
<abbrev id="ABBRID0E6CAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
outline Epsilon
|
||
<typeStatus id="489701929807CCD24AF46CC63887A593">type</typeStatus>
|
||
. Anterior margin of
|
||
<abbrev id="ABBRID0EDDAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
with 1–2 anterolateral tubercles. Anteromedial process of the cheliceral sockets shorter than lateral processes. Eyes separated into two small smooth mounds, placed posteriorly on the carapace. Interocular region with small granules in the posterior zone and one central elevated mound that terminates in two short spines. Lateral margins smooth. Mesotergum divided into four areas, III – IV partially fused: I divided medially into two triangular halves, with one conspicuous tubercle on each side; II entire, with four conspicuous tubercles; III with a lateral pair of large tubercles; one pair of paramedian large spines with granulated base; IV with some granules and one row of four tubercles, two each side. Posterior margin and free tergites with a pair of paramedian acute granules.
|
||
<emphasis id="3B33295C8819CF0A9557F50B6EB083A2" bold="true" italics="true">Venter</emphasis>
|
||
(Figs
|
||
<figureCitation id="CD94051C809902944CCD681E7A35834D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eutimesius aroa sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105934); C, D. Female habitus, dorsal view (MIZA 0105936); C. Male habitus, lateral view (MIZA 0105934). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure4" httpUri="https://binary.pensoft.net/fig/1072632">4 C</figureCitation>
|
||
,
|
||
<figureCitation id="CD99EB97416F2F29A917703718F3E557" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutimesius aroa sp. nov., male (MIZA 0105934): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure5" httpUri="https://binary.pensoft.net/fig/1072633">5 B</figureCitation>
|
||
). Coxa I with four distal tubercles and a medial row of six tubercles; II with a median row of seven-eight tubercules and a posterior row of two-three small tubercles; III with six intercoxal tubercles, five distal, two anterior, six medial, five posterior tubercles; IV with eight intercoxal tubercles, six anterior tubercles, nine medial tubercles, and about 12 posterior tubercles not aligned. Genital operculum with scattered granules. Stigmatic area with two anterior tubercles, a medial row of four tubercles, and a posterior row of minute granules on the posterior border. Free sternites with a row of small granules.
|
||
<emphasis id="F41D1DFC486FC69A1846455ECA43FD75" bold="true" italics="true">Chelicerae</emphasis>
|
||
(Fig.
|
||
<figureCitation id="9493B9BC655E8BAF02E2A5470D49BDB7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutimesius aroa sp. nov., male (MIZA 0105934): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure5" httpUri="https://binary.pensoft.net/fig/1072633">5 A – C</figureCitation>
|
||
). Segment I smooth with well-defined bulla, with one or two ectoproximal tubercles and one ectodistal tubercle. Segment II swollen, fixed finger with a proximal wide laminar tooth, followed by two very small denticles and three decreasing medial teeth; mobile finger with one subproximal large truncated tooth, one medial pyramidal tooth, and two small subdistal teeth.
|
||
<emphasis id="58DE6533E064B84014B296656B085871" bold="true" italics="true">Pedipalps</emphasis>
|
||
(Fig.
|
||
<figureCitation id="A8E06C7917E646675E976A08E953B39D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutimesius aroa sp. nov., male (MIZA 0105934): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure5" httpUri="https://binary.pensoft.net/fig/1072633">5 D, E</figureCitation>
|
||
). Coxa with a group of eight ventral tubercles and three-four dorsal tubercles. Trochanter with a large ventral tubercle and two-three dorsal tubercles. Femur with a ventral row of six small tubercles and dorsally smooth. Patella smooth, distally swollen. Tibia dorsal smooth, ventrally with a row of minute granules; mesal IIiIi; ectal IiIiI. Tarsus dorsally smooth, ventrally with two rows of minute granules, mesal IiIiIi; ectal iIiiIii.
|
||
<emphasis id="C02C59D16850055A2C4EA2C7FAD2BDE5" bold="true" italics="true">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation id="436183A8ABD102FE26BDBD7ADA97C5E2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutimesius aroa sp. nov., male (MIZA 0105934): A. Habitus, dorsal view; B. Ditto, lateral view; C. Right chelicera, frontal view; D. Right pedipalp, tibia, and tarsus, ectal view; E. Ditto, mesal view; F. Right leg IV, dorsal view; G. Ditto, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure5" httpUri="https://binary.pensoft.net/fig/1072633">5 F – I</figureCitation>
|
||
). Coxae I – II with two dorsal tubercles; III – IV connected by four – five intercoxal tubercles; IV with one dorsodistal large tubercle and scattered small granules and tubercles. Trochanter I dorsally smooth, ventrally with three tubercles; II with two dorsal, one retrodistal, and three ventral tubercles; III with one prolateral, two-four dorsal, one retrodorsal large, and five ventral tubercles; IV with one prolateral, one dorsal, and some minute dorsal granules; one retrolateral large tubercle; and eight ventral tubercles. Femora I – II with longitudinal rows of minute granules; III and IV with the one proventral and one retroventral row of large and spaced apart tubercles; and with two dorsoapical tubercles. Patella III – IV each with one large proventral tubercle; IV with pro and retrodorsal distal large tubercles and with sparse minute granules. Tibia III, with the tubercles of the distal portion of the proventral row enlarged and all the retroventral row with large tubercles increasing in size distally; IV with similar pattern but more conspicuous. Basitarsus I slightly swollen. Tarsal process and scopula present. Tarsal claws III and IV opposites and pectinated. Tarsal counts: 7 (3) / 19 (
|
||
<abbrev id="ABBRID0EHEAG" xlink_title="broken">br</abbrev>
|
||
) - 22 (3) / 8 / 9.
|
||
<emphasis id="B718175A4770ADCD360D086F97DCF27D" bold="true" italics="true">Penis</emphasis>
|
||
(Fig.
|
||
<figureCitation id="692FA35394885B1F70A49807F4E1C0C6" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eutimesius spp., penis, distal portion in dorsal, ventral, and lateral view: A – C. E. aroa sp. nov., (MIZA 0105934); D – F. E. canoabo sp. nov., (MIZA 0105945); G – I. E. ephippiatus, (ICN-Ao - 1492); J – L. E. guaichia sp. nov., (MIZA 0105935); M – O. E. simoni, (ICN-Ao - 328). A – I. Unscaled." figureDoi="10.3897/zse.100.120207.figure14" httpUri="https://binary.pensoft.net/fig/1072642">14 A – C</figureCitation>
|
||
).
|
||
<taxonomicName id="952D2920C2D595A38FBAC4950AACD093" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Stygnidae" kingdom="Animalia" order="Opiliones" phylum="Arthropoda" rank="subFamily" subFamily="Heterostygninae">Heterostygninae</taxonomicName>
|
||
general pattern, as described in
|
||
<bibRefCitation id="5EF6CBAC1FF2E55649EC3653FE1A05C3" DOI="10.1590/1678-4766e2019008" author="Villarreal O & Sanchéz N & De Ascenção A" issue="1" journalOrPublisher="Série Zoologia" pagination="1-6" refId="B18" refString="Villarreal O, Sanchéz N, De Ascenção A (2019 b) New generic assignment to the harvestman Metaphareus punctatus (Opiliones: Stygnidae) and observations about it reproductive behavior. Iheringia. Série Zoologia 109 (1): 1–6. https://doi.org/10.1590/1678-4766e2019008" title="New generic assignment to the harvestman Metaphareus punctatus (Opiliones: Stygnidae) and observations about it reproductive behavior. Iheringia." volume="109">Villarreal et al. (2019 b</bibRefCitation>
|
||
). Truncus with the malleus swollen.
|
||
<emphasis id="99AA7C9A436BF005D8874C617117D33F" italics="true">Lamina parva</emphasis>
|
||
(
|
||
<abbrev id="ABBRID0E4EAG" xlink_title="lamina parva">LP</abbrev>
|
||
) with a shallow neck and a deep distal cleft.
|
||
<abbrev id="ABBRID0EBFAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- A 1 - A 2 located on the malleus, one pair laterally and one pair more ventrally;
|
||
<abbrev id="ABBRID0EFFAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- B pair ventrally located on the
|
||
<abbrev id="ABBRID0EJFAG" xlink_title="lamina parva">LP</abbrev>
|
||
; two pairs of
|
||
<abbrev id="ABBRID0ENFAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- C located medially on the
|
||
<abbrev id="ABBRID0ERFAG" xlink_title="lamina parva">LP</abbrev>
|
||
, dorsally to the neck;
|
||
<abbrev id="ABBRID0EVFAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- D 1 large, located slightly proximal to
|
||
<abbrev id="ABBRID0EZFAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- E;
|
||
<abbrev id="ABBRID0E4FAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- D 2 basally located, near the base of the gland;
|
||
<abbrev id="ABBRID0EBGAG" xlink_title="macrosetae of penis">MS</abbrev>
|
||
- E with only one pair of short setae. Gland globose, with short and dorsally curved stylus with small dorsal process.
|
||
<emphasis id="0D2FBDE72D4F80999918E711D89E44F1" bold="true" italics="true">Color</emphasis>
|
||
(Fig.
|
||
<figureCitation id="DEEFEC02A32F61F7FB79C65A12FA7411" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eutimesius aroa sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105934); C, D. Female habitus, dorsal view (MIZA 0105936); C. Male habitus, lateral view (MIZA 0105934). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure4" httpUri="https://binary.pensoft.net/fig/1072632">4 A, C</figureCitation>
|
||
).
|
||
<abbrev id="ABBRID0EMGAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
mottled on background brilliant yellow (83). Spots on the carapace moderate brown (58); area I, bases of tubercles of area II; tubercles of area III and margins of
|
||
<abbrev id="ABBRID0EQGAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
; free tergites and coxae I – IV dorsally dark grayish brown (62) and blackish red (21). Trochanters I – IV and chelicerae reticulated in the same colors as
|
||
<abbrev id="ABBRID0EUGAG" xlink_title="dorsal scutum">DS</abbrev>
|
||
. Femora I – II reticulated dark grayish olive (111) on background moderate yellow green (120); III and IV dark grayish olive (111).
|
||
</paragraph>
|
||
<paragraph id="A0F08958ADB891CFB1BB853BD1BCF8C6">
|
||
<emphasis id="863D7A10E941D7526A0BC842C0B62CB5" bold="true">
|
||
Female.
|
||
<emphasis id="53C21E0F9AFF8FB2C95251DC61C24A77" bold="true" italics="true">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dorsal scutum length 4.1; dorsal scutum width 3.4; abdominal scutum length 1.8; abdominal scutum width 2.8; interocular distance 1.7; pedipalp: coxa 0.6, trochanter 0.8, femur 3.9, patella 1.6, tibia 2.0, tarsus 2.2, claw 1.7; total 12.8; leg IV: femur 7.3, tibia 3.9.
|
||
<emphasis id="B401A0957E3FFFF16A180F778D4AEC71" bold="true" italics="true">Description</emphasis>
|
||
(Fig.
|
||
<figureCitation id="58E0496BA349E81CDD057CC3AA5688A6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eutimesius aroa sp. nov., A, B. Male habitus, dorsal and lateral views (MIZA 0105934); C, D. Female habitus, dorsal view (MIZA 0105936); C. Male habitus, lateral view (MIZA 0105934). Scale bars: 1 mm." figureDoi="10.3897/zse.100.120207.figure4" httpUri="https://binary.pensoft.net/fig/1072632">4 B</figureCitation>
|
||
). Similar to male, except by abdominal scutum Eta
|
||
<typeStatus id="397D76425B989678CDCF89A1B344979C">type</typeStatus>
|
||
; chelicerae not swollen; ornamentation of leg IV conspicuously less developed; basitarsus I not swollen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EGHAG" type="distribution">
|
||
<paragraph id="581773A66ED80B2852B89B5FEDD096BB">
|
||
<heading id="499D27ECA47CFFC39D2DBEBB95C5396A" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="A478FB9F7817DC6B745FDFDD7B354538">
|
||
<collectingCountry id="661DC6B58C7C88DA5A14735E3BEBAABF" name="Venezuela">Venezuela</collectingCountry>
|
||
,
|
||
<collectingRegion id="8E9C8EF6E0F46C40D8D72B1623A601D8" country="Venezuela" name="Yaracuy">Yaracuy</collectingRegion>
|
||
. Only known from the
|
||
<typeStatus id="EEDA31C02D9AF53EF312884BF059608A">type</typeStatus>
|
||
locality (Fig.
|
||
<figureCitation id="BC34CD413EC63A6E074C3578D275176B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Map showing the distribution of Eutimesius." figureDoi="10.3897/zse.100.120207.figure2" httpUri="https://binary.pensoft.net/fig/1072630">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |