treatments-xml/data/E1/38/7D/E1387D9950055EA1BB48A91CED368744.xml
2024-06-21 12:55:01 +02:00

263 lines
21 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/BDJ.7.e39140" ID-PMC="PMC6856199" ID-Pensoft-Pub="1314-2828-7-e39140" ID-Pensoft-UUID="D504D640D9DE54B9818CD192D3389EBC" ID-PubMed="31745388" ID-ZooBank="90B52445FA2F4B1A98A19219AA151CAC" ModsDocID="1314-2828-7-e39140" checkinTime="1573201115876" checkinUser="pensoft" docAuthor="Pham, Anh Van, Pham, Cuong The, Doan, Lan Duc, Ziegler, Thomas &amp; Nguyen, Truong Quang" docDate="2019" docId="E1387D9950055EA1BB48A91CED368744" docLanguage="en" docName="BiodivDatJour 7: e39140" docOrigin="Biodiversity Data Journal 7" docSource="http://dx.doi.org/10.3897/BDJ.7.e39140" docTitle="Megophrys parva Boulenger 1893" docType="treatment" docVersion="4" id="D504D640D9DE54B9818CD192D3389EBC" lastPageNumber="39140" masterDocId="D504D640D9DE54B9818CD192D3389EBC" masterDocTitle="New records of megophryids (Amphibia: Anura: Megophryidae) from Son La Province, Vietnam" masterLastPageNumber="39140" masterPageNumber="39140" pageNumber="39140" updateTime="1668125089560" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New records of megophryids (Amphibia: Anura: Megophryidae) from Son La Province, Vietnam</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pham, Anh Van</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pham, Cuong The</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Doan, Lan Duc</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ziegler, Thomas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nguyen, Truong Quang</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>7</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>39140</mods:start>
<mods:end>39140</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.7.e39140</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.7.e39140</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828-7-e39140</mods:identifier>
<mods:identifier type="ZooBank">90B52445FA2F4B1A98A19219AA151CAC</mods:identifier>
<mods:identifier type="Pensoft-UUID">D504D640D9DE54B9818CD192D3389EBC</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:E1387D9950055EA1BB48A91CED368744" httpUri="http://treatment.plazi.org/id/E1387D9950055EA1BB48A91CED368744" lastPageNumber="39140" pageId="0" pageNumber="39140">
<subSubSection pageId="0" pageNumber="39140" type="nomenclature">
<paragraph pageId="0" pageNumber="39140">
<taxonomicName LSID="E1387D99-5005-5EA1-BB48-A91CED368744" authority="Boulenger, 1893" authorityName="Boulenger" authorityYear="1893" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megophrys parva" order="Anura" pageId="0" pageNumber="39140" phylum="Chordata" rank="species" species="parva">Megophrys parva (Boulenger, 1893)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39140" type="materials_examined">
<paragraph pageId="0" pageNumber="39140">Materials</paragraph>
<paragraph pageId="0" pageNumber="39140">
<materialsCitation collectingDate="2014-06-19" collectionCode="Amphibians" country="Vietnam" latitude="21.396" location="near Huoi Pu Village" longLatPrecision="12" longitude="103.6425" pageId="0" pageNumber="39140" specimenCode="TBU PAE. 705" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="39140">Type status:</emphasis>
<typeStatus pageId="0" pageNumber="39140">Other material</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="39140">Occurrence:</emphasis>
catalogNumber:
<specimenCode pageId="0" pageNumber="39140">TBU PAE.705</specimenCode>
; individualCount:
<specimenCount pageId="0" pageNumber="39140">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="39140">male</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="39140">adult</specimenType>
;
<emphasis bold="true" pageId="0" pageNumber="39140">Taxon:</emphasis>
scientificNameID: Megophrysparva; scientificName: Megophrysparva; class: Amphibia; order: Anura; family: Megophryidae; genus: Megophrys; specificEpithet: parva; scientificNameAuthorship: (Boulenger, 1893);
<emphasis bold="true" pageId="0" pageNumber="39140">Location:</emphasis>
country:
<collectingCountry pageId="0" pageNumber="39140">Vietnam</collectingCountry>
; countryCode: VN; stateProvince: Son La; county: Thuan Chau; municipality: Chieng Bom; locality:
<location LSID="urn:lsid:plazi:treatment:E1387D9950055EA1BB48A91CED368744:1018A1C836EF618CACCB6456A47407CA" country="Vietnam" latitude="21.396" longLatPrecision="12" longitude="103.6425" name="near Huoi Pu Village" pageId="0" pageNumber="39140">near Huoi Pu Village</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="39140">1290 m</elevation>
; verbatimLatitude:
<geoCoordinate degrees="21" direction="north" minutes="23.76" orientation="latitude" precision="9" value="21.396">21°23.76'N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate degrees="103" direction="east" minutes="38.55" orientation="longitude" precision="9" value="103.6425">103°38.55'E</geoCoordinate>
; verbatimCoordinateSystem: WGS84;
<emphasis bold="true" pageId="0" pageNumber="39140">Event:</emphasis>
eventDate:
<collectingDate pageId="0" pageNumber="39140" value="2014-06-19">June 19, 2014</collectingDate>
; eventRemarks: collected by A. V. Pham, H. V. Tu and C. K. P. D. Kham;
<emphasis bold="true" pageId="0" pageNumber="39140">Record Level:</emphasis>
language: en; collectionCode:
<collectionCode pageId="0" pageNumber="39140">Amphibians</collectionCode>
; basisOfRecord: Preserved Specimen
</materialsCitation>
<materialsCitation collectingDate="2014-06-19" collectionCode="Amphibians" country="Vietnam" latitude="21.396" location="near Huoi Pu Village" longLatPrecision="12" longitude="103.6425" pageId="0" pageNumber="39140" specimenCode="TBU PAE. 711" specimenCount="1" specimenCount-female="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="39140">Type status:</emphasis>
<typeStatus pageId="0" pageNumber="39140">Other material</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="39140">Occurrence:</emphasis>
catalogNumber:
<specimenCode pageId="0" pageNumber="39140">TBU PAE.711</specimenCode>
; individualCount:
<specimenCount pageId="0" pageNumber="39140">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="39140">female</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="39140">adult</specimenType>
;
<emphasis bold="true" pageId="0" pageNumber="39140">Taxon:</emphasis>
scientificNameID: Megophrysparva; scientificName: Megophrysparva; class: Amphibia; order: Anura; family: Megophryidae; genus: Megophrys; specificEpithet: parva; scientificNameAuthorship: (Boulenger, 1893);
<emphasis bold="true" pageId="0" pageNumber="39140">Location:</emphasis>
country:
<collectingCountry pageId="0" pageNumber="39140">Vietnam</collectingCountry>
; countryCode: VN; stateProvince: Son La; county: Thuan Chau; municipality: Chieng Bom; locality:
<location LSID="urn:lsid:plazi:treatment:E1387D9950055EA1BB48A91CED368744:CB4CDB6696A12C4DF09AB86E90C341FD" country="Vietnam" latitude="21.396" longLatPrecision="12" longitude="103.6425" name="near Huoi Pu Village" pageId="0" pageNumber="39140">near Huoi Pu Village</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="39140">1290 m</elevation>
; verbatimLatitude:
<geoCoordinate degrees="21" direction="north" minutes="23.76" orientation="latitude" precision="9" value="21.396">21°23.76'N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate degrees="103" direction="east" minutes="38.55" orientation="longitude" precision="9" value="103.6425">103°38.55'E</geoCoordinate>
; verbatimCoordinateSystem: WGS84;
<emphasis bold="true" pageId="0" pageNumber="39140">Event:</emphasis>
eventDate:
<collectingDate pageId="0" pageNumber="39140" value="2014-06-19">June 19, 2014</collectingDate>
; eventRemarks: collected by A. V. Pham, H. V. Tu and C. K. P. D. Kham;
<emphasis bold="true" pageId="0" pageNumber="39140">Record Level:</emphasis>
language: en; collectionCode:
<collectionCode pageId="0" pageNumber="39140">Amphibians</collectionCode>
; basisOfRecord: Preserved Specimen
</materialsCitation>
<materialsCitation collectingDate="2014-06-19" collectionCode="Amphibians" country="Vietnam" latitude="21.396" location="near Huoi Pu Village" longLatPrecision="12" longitude="103.6425" pageId="0" pageNumber="39140" specimenCode="TBU PAE. 712" specimenCount="1" specimenCount-female="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="39140">Type status:</emphasis>
<typeStatus pageId="0" pageNumber="39140">Other material</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="39140">Occurrence:</emphasis>
catalogNumber:
<specimenCode pageId="0" pageNumber="39140">TBU PAE.712</specimenCode>
; individualCount:
<specimenCount pageId="0" pageNumber="39140">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="39140">female</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="39140">adult</specimenType>
;
<emphasis bold="true" pageId="0" pageNumber="39140">Taxon:</emphasis>
scientificNameID: Megophrysparva; scientificName: Megophrysparva; class: Amphibia; order: Anura; family: Megophryidae; genus: Megophrys; specificEpithet: parva; scientificNameAuthorship: (Boulenger, 1893);
<emphasis bold="true" pageId="0" pageNumber="39140">Location:</emphasis>
country:
<collectingCountry pageId="0" pageNumber="39140">Vietnam</collectingCountry>
; countryCode: VN; stateProvince: Son La; county: Thuan Chau; municipality: Chieng Bom; locality:
<location LSID="urn:lsid:plazi:treatment:E1387D9950055EA1BB48A91CED368744:18BD1C9B0C0432C2E474C299844D050E" country="Vietnam" latitude="21.396" longLatPrecision="12" longitude="103.6425" name="near Huoi Pu Village" pageId="0" pageNumber="39140">near Huoi Pu Village</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="39140">1290 m</elevation>
; verbatimLatitude:
<geoCoordinate degrees="21" direction="north" minutes="23.76" orientation="latitude" precision="9" value="21.396">21°23.76'N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate degrees="103" direction="east" minutes="38.55" orientation="longitude" precision="9" value="103.6425">103°38.55'E</geoCoordinate>
; verbatimCoordinateSystem: WGS84;
<emphasis bold="true" pageId="0" pageNumber="39140">Event:</emphasis>
eventDate:
<collectingDate pageId="0" pageNumber="39140" value="2014-06-19">June 19, 2014</collectingDate>
; eventRemarks: collected by A. V. Pham, H. V. Tu and C. K. P. D. Kham;
<emphasis bold="true" pageId="0" pageNumber="39140">Record Level:</emphasis>
language: en; collectionCode:
<collectionCode pageId="0" pageNumber="39140">Amphibians</collectionCode>
; basisOfRecord: Preserved Specimen
</materialsCitation>
<materialsCitation collectingDate="2015-04-01" collectionCode="Amphibians" country="Vietnam" latitude="21.014833" location="near Pa Tet Village" longLatPrecision="12" longitude="103.63533" pageId="0" pageNumber="39140" specimenCode="TBU SC. 2015.06" specimenCount="1" specimenCount-male="1" typeStatus="Other material">
<emphasis bold="true" pageId="0" pageNumber="39140">Type status:</emphasis>
<typeStatus pageId="0" pageNumber="39140">Other material</typeStatus>
.
<emphasis bold="true" pageId="0" pageNumber="39140">Occurrence:</emphasis>
catalogNumber:
<specimenCode pageId="0" pageNumber="39140">TBU SC.2015.06</specimenCode>
; individualCount:
<specimenCount pageId="0" pageNumber="39140">1</specimenCount>
; sex:
<specimenType pageId="0" pageNumber="39140">male</specimenType>
; lifeStage:
<specimenType pageId="0" pageNumber="39140">adult</specimenType>
;
<emphasis bold="true" pageId="0" pageNumber="39140">Taxon:</emphasis>
scientificNameID: Megophrysparva; scientificName: Megophrysparva; class: Amphibia; order: Anura; family: Megophryidae; genus: Megophrys; specificEpithet: parva; scientificNameAuthorship: (Boulenger, 1893);
<emphasis bold="true" pageId="0" pageNumber="39140">Location:</emphasis>
country:
<collectingCountry pageId="0" pageNumber="39140">Vietnam</collectingCountry>
; countryCode: VN; stateProvince: Son La; county: Song Ma; municipality: Huoi Mot; locality:
<location LSID="urn:lsid:plazi:treatment:E1387D9950055EA1BB48A91CED368744:06508CB6EE8AFB34AF66F81650B2A84A" country="Vietnam" latitude="21.014833" longLatPrecision="12" longitude="103.63533" name="near Pa Tet Village" pageId="0" pageNumber="39140">near Pa Tet Village</location>
; verbatimElevation:
<elevation pageId="0" pageNumber="39140">1270 m</elevation>
; verbatimLatitude:
<geoCoordinate degrees="21" direction="north" minutes="00.89" orientation="latitude" precision="9" value="21.014833">21°00.89'N</geoCoordinate>
; verbatimLongitude:
<geoCoordinate degrees="103" direction="east" minutes="38.12" orientation="longitude" precision="9" value="103.63533">103°38.12'E</geoCoordinate>
; verbatimCoordinateSystem: WGS84;
<emphasis bold="true" pageId="0" pageNumber="39140">Event:</emphasis>
eventDate:
<collectingDate pageId="0" pageNumber="39140" value="2015-04-01">April 1, 2015</collectingDate>
; eventRemarks: collected by A. V. Pham and D. B. Giang;
<emphasis bold="true" pageId="0" pageNumber="39140">Record Level:</emphasis>
language: en; collectionCode:
<collectionCode pageId="0" pageNumber="39140">Amphibians</collectionCode>
; basisOfRecord: Preserved Specimen
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39140" type="description">
<paragraph pageId="0" pageNumber="39140">Description</paragraph>
<paragraph pageId="0" pageNumber="39140">
Morphological characters of the specimens (n = 4) from Son La Province agreed with the descriptions of
<bibRefCitation author="Boulenger, G A" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="39140" pagination="304 - 347" refId="B5305894" refString="Boulenger, G A, 1893. Concluding report on the reptiles and batrachians obtained in Burma by Signor L. Fea dealing with the collection made in Pegu and the Karin Hills in 1887-88. Annali del Museo Civico di Storia Naturale di Genova 2: 304 - 347" title="Concluding report on the reptiles and batrachians obtained in Burma by Signor L. Fea dealing with the collection made in Pegu and the Karin Hills in 1887 - 88" volume="2" year="1893">Boulenger (1893)</bibRefCitation>
,
<bibRefCitation author="Bourret, R" journalOrPublisher="Institut Oceanographique de l'Indochine, Hanoi" pageId="0" pageNumber="39140" refId="B5305904" refString="Bourret, R, 1942. Les Batraciens de l'Indochine. Institut Oceanographique de l'Indochine, Hanoi" title="Les Batraciens de l'Indochine" year="1942">Bourret (1942)</bibRefCitation>
, and
<bibRefCitation author="Fei, L" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="39140" refId="B5305923" refString="Fei, L, Hu, S-q, Ye, C-y, Huang, Y-z, 2009. Fauna Sinica. Amphibia. Vol. 2. Anura. Science Press, Beijing" title="Fauna Sinica. Amphibia. Vol. 2. Anura" year="2009">Fei et al. (2009)</bibRefCitation>
: SVL 44.5-46.7 mm in males (n = 2), 55.4-56.7 mm in females (n = 2); head longer than wide (males: HL 15.4-16.5 mm, HW 14.8-16.2 mm, HL/SVL 35%, HW/SVL 33-35%; females: HL 18.7 mm, HW 18.0-18.5 mm, HL/SVL 33-34%, HW/SVL 32-33%); snout obliquely truncate, projecting beyond upper jaw (SL 5.9-6.7 mm in males, 6.7-7.0 mm in females), longer than horizontal diameter of eye (EL 5.3-5.9 mm in males, 5.8 mm in females); loreal region concave; nostril lateral, slightly closer to eye than to tip of snout (NS 2.8-3.2 mm, EN 2.7-2.9 mm in males; NS 3.2-3.3 mm, EN 3.1-3.2 mm in females); interorbital space flat, broader than upper eyelid and narrower than internarial distance (IOD 4.6-4.7 mm, UEW 4.1-4.2 mm, IN 5.0-5.5 mm in males; IOD 5.0-5.2 mm, UEW 4.2-4.7 mm, IN 5.5-5.6 mm in females); tympanum distinct (TD 3.5-3.7 mm in males, 4.5 mm in females); vomerine teeth present; tongue
<normalizedToken originalValue="heartshaped">heart-shaped</normalizedToken>
, round posteriorly; vocal sacs present in males.
</paragraph>
<paragraph pageId="0" pageNumber="39140">Forelimb slender (FLL 27.0-28.9 mm in males, 31.0-34.1 mm in females); relative finger lengths I ≤ II &lt;IV &lt;III; fingers dermal fringe absent, free of webbing; tips of fingers round, not swollen; subarticular tubercles indistinct; palmar tubercles indistinct; nuptial pads absent in males.</paragraph>
<paragraph pageId="0" pageNumber="39140">Hindlimb slender, long (HLL 69.1-75.6 mm in males, 85.0-94.0 mm in females); tibia longer than thigh (TL 22.2-23.7 mm, FL 20.5-20.6 mm in males; TL 27.0-29.8 mm, FL 24.4-24.6 mm in females); relative toe lengths I &lt;II &lt;V &lt;III &lt;IV; tips of toes round, slightly swollen; webbing rudimentary; toes without dermal fringe; metatarsal tubercle indistinct; subarticular tubercles indistinct; tibiotarsal articulation reaching to anterior margin of eye.</paragraph>
<paragraph pageId="0" pageNumber="39140">Skin. Dorsal surface smooth; flanks with small glandular warts; supratympanic fold present, from posterior edge of eye to axilla; around cloaca with small tubercles; ventral surface smooth.</paragraph>
<paragraph pageId="0" pageNumber="39140">
Colouration in life. Dorsal surface light yellowish-brown; a darker brown triangular marking between eyes; upper lips with vertical dark brown bars; ventral surface whitish, a round white spot on each side of the chest present; ventral surface of limbs reddish (
<figureCitation captionStart="Figure 7" captionStartId="F5306246" captionText="Figure 7. Megophrys parva (TBU PAE. 705, adult male) from Son La Province, Vietnam. Photo by A. V. Pham." figureDoi="10.3897/BDJ.7.e39140.figure7" httpUri="https://binary.pensoft.net/fig/324975" pageId="0" pageNumber="39140" tableDoi="10.3897/BDJ.7.e39140.figure7">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39140" type="distribution">
<paragraph pageId="0" pageNumber="39140">Distribution</paragraph>
<paragraph pageId="0" pageNumber="39140">
In Vietnam, this species has been recorded from Lao Cai, Lai Chau, Dien Bien, and Ha Giang provinces (
<bibRefCitation author="Nguyen, S V" journalOrPublisher="Edition Chimaira, Frankfurt am Main" pageId="0" pageNumber="39140" refId="B5306039" refString="Nguyen, S V, Ho, C T, Nguyen, T Q, 2009. Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main" title="Herpetofauna of Vietnam" year="2009">Nguyen et al. 2009</bibRefCitation>
,
<bibRefCitation author="Orlov, N L" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="39140" pagination="206 - 218" refId="B5306090" refString="Orlov, N L, Poyarkov, N A J, Nguyen, T T, 2015. Taxonomic notes on Megophrys frogs (Megophryidae: Anura) of Vietnam, with description of a new species. Russian Journal of Herpetology 22: 206 - 218" title="Taxonomic notes on Megophrys frogs (Megophryidae: Anura) of Vietnam, with description of a new species" volume="22" year="2015">Orlov et al. 2015</bibRefCitation>
,
<bibRefCitation author="Luong, A M" journalOrPublisher="Herpetology Notes" pageId="0" pageNumber="39140" pagination="375 - 387" refId="B5306028" refString="Luong, A M, Nguyen, H Q, Le, D T, Nguyen, S H L, Nguyen, T Q, 2019. New records of amphibians (Anura: Megophryidae, Ranidae) from Dien Bien Province, Vietnam. Herpetology Notes 12: 375 - 387" title="New records of amphibians (Anura: Megophryidae, Ranidae) from Dien Bien Province, Vietnam" volume="12" year="2019">Luong et al. 2019</bibRefCitation>
). Our record in Son La Province is the southernmost record of this species in Vietnam. Elsewhere, this species is known from Nepal, India, Bangladesh, Myanmar, Thailand, China, and Laos (
<bibRefCitation author="Nguyen, S V" journalOrPublisher="Edition Chimaira, Frankfurt am Main" pageId="0" pageNumber="39140" refId="B5306039" refString="Nguyen, S V, Ho, C T, Nguyen, T Q, 2009. Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main" title="Herpetofauna of Vietnam" year="2009">Nguyen et al. 2009</bibRefCitation>
,
<bibRefCitation author="Fei, L" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="39140" refId="B5305932" refString="FrostD RAmphibian species of the worldhttp://research.amnh.org/herpetology/amphibia/index.php2019-07-30T00:00:00+03:00" year="2009">Frost 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="39140" type="ecology">
<paragraph pageId="0" pageNumber="39140">Ecology</paragraph>
<paragraph pageId="0" pageNumber="39140">The specimens were found on the edge of streams, between 19:00 and 22:00 h. The surrounding habitat was mixed secondary forest of medium hardwoods and shrubs.</paragraph>
</subSubSection>
</treatment>
</document>