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<document id="FB0DADF60154B5E25300BB0DCAB13334" ID-DOI="10.11646/zootaxa.5551.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="14421367" ID-ZooBank="15F036B2-4B1F-41AB-953A-9F096BD2AB5E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1734024468337" checkinUser="plazi" docAuthor="Cao, Wenqiang, Yang, Ding &amp; Dong, Hui" docDate="2024" docId="E136E71CAC704474FF0DF955FCBDA08F" docLanguage="en" docName="zootaxa.5551.2.6.pdf" docOrigin="Zootaxa 5551 (2)" docSource="https://doi.org/10.11646/zootaxa.5551.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Opalimosina spathulata Hayashi 2010" docType="treatment" docVersion="2" lastPageNumber="366" masterDocId="1D0F9F64AC7C4479FF9AFFB4FFEEA719" masterDocTitle="A review of Opalimosina Roháček, 1983 (Diptera, Sphaeroceridae) from China with new records and two new species" masterLastPageNumber="368" masterPageNumber="353" pageNumber="365" updateTime="1734025112797" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4A3C324DE5DE3A731FA80050D901E816">A review of Opalimosina Roháček, 1983 (Diptera, Sphaeroceridae) from China with new records and two new species</mods:title>
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<mods:namePart id="294D35274008C40CB63C6AC6AA12DD70">Cao, Wenqiang</mods:namePart>
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<mods:affiliation id="06B9BE9B5B0A3C4B27C34026AFB74EDE">Department of Entomology, College of Plant Protection, China Agricultural University, 2 Yuanmingyuan West Road, Beijing 100193, China</mods:affiliation>
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<mods:namePart id="D097CF40E4A0ACA8821B03E660CB4887">Yang, Ding</mods:namePart>
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<mods:affiliation id="CBA76E6DF95F6E9C879E8B0A430DAADD">Department of Entomology, College of Plant Protection, China Agricultural University, 2 Yuanmingyuan West Road, Beijing 100193, China</mods:affiliation>
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<mods:affiliation id="8417914FD8BD6C40D6C89D4C535F0CFE">Key Laboratory of Southern Subtropical Plant Diversity, Fairylake Botanical Garden, Shenzhen &amp; Chinese Academy of Sciences, Shenzhen 518004, China</mods:affiliation>
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<paragraph id="6920560AAC704475FF0DF955FDE1A1E2" blockId="12.[151,527,1761,1823]" box="[151,527,1761,1788]" pageId="12" pageNumber="365">
<heading id="3268E166AC704475FF0DF955FDE1A1E2" bold="true" box="[151,527,1761,1788]" fontSize="11" level="1" pageId="12" pageNumber="365" reason="1">
<emphasis id="5BEB8A18AC704475FF0DF955FDE1A1E2" bold="true" box="[151,527,1761,1788]" pageId="12" pageNumber="365">
9.
<taxonomicName id="AE9F2D89AC704475FF2FF955FDE1A1E2" authority="Hayashi, 2010" authorityName="Hayashi" authorityYear="2010" box="[181,527,1761,1788]" class="Insecta" family="Sphaeroceridae" genus="Opalimosina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="365" phylum="Arthropoda" rank="species" species="spathulata">
<emphasis id="5BEB8A18AC704475FF2FF955FEB9A1E5" bold="true" box="[181,343,1761,1788]" italics="true" pageId="12" pageNumber="365">O. spathulata</emphasis>
<bibRefCitation id="0D0E2BFBAC704475FEC5F955FDE1A1E2" author="Hayashi, T." box="[351,527,1761,1787]" pageId="12" pageNumber="365" pagination="309 - 319" refId="ref10082" refString="Hayashi, T. (2010) The genus Opalimosina Rohacek (Diptera, Sphaeroceridae) from Japan. Medical Entomology and Zoology, 61 (3), 309 - 319. https: // doi. org / 10.7601 / mez. 61.309." type="journal article" year="2010">Hayashi, 2010</bibRefCitation>
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</paragraph>
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<subSubSection id="21850581AC704475FF0DF8B1FEC0A006" box="[151,302,1797,1823]" pageId="12" pageNumber="365" type="description">
<paragraph id="6920560AAC704475FF0DF8B1FEC0A006" blockId="12.[151,527,1761,1823]" box="[151,302,1797,1823]" pageId="12" pageNumber="365">
(
<figureCitation id="F1A44A8FAC704475FF05F8B1FF13A006" box="[159,253,1797,1823]" captionStart="FIGURES 12. 12" captionStartId="13.[151,264,658,682]" captionTargetBox="[354,1234,237,634]" captionTargetId="figure-49@13.[354,1234,237,634]" captionTargetPageId="13" captionText="FIGURES 12. 12AB. O. spathulata Hayashi, 2010 male habitus. 12A, lateral; 12B, dorsal. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14421402" httpUri="https://zenodo.org/record/14421402/files/figure.png" pageId="12" pageNumber="365">Figs. 12</figureCitation>
, 13)
</paragraph>
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<paragraph id="6920560AAC704475FF0DF8FEFB33A078" blockId="12.[151,1245,1866,1890]" box="[151,1245,1866,1890]" pageId="12" pageNumber="365">
<treatmentCitationGroup id="498F7124AC704475FF0DF8FEFB33A078" box="[151,1245,1866,1890]" pageId="12" pageNumber="365">
<treatmentCitation id="E83E701BAC704475FF0DF8FEFDB9A07B" author="Hayashi, T." box="[151,599,1866,1890]" page="317" pageId="12" pageNumber="365" year="2010">
<taxonomicName id="AE9F2D89AC704475FF0DF8FEFDB9A07B" authority="Hayashi, 2010: 317" authorityName="Hayashi" authorityPageNumber="317" authorityYear="2010" box="[151,599,1866,1890]" class="Insecta" family="Sphaeroceridae" genus="Opalimosina" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="365" phylum="Arthropoda" rank="species" species="spathulata">
<emphasis id="5BEB8A18AC704475FF0DF8FEFE65A07B" box="[151,395,1866,1890]" italics="true" pageId="12" pageNumber="365">Opalimosina spathulata</emphasis>
<bibRefCitation id="0D0E2BFBAC704475FE08F8FEFDB9A07B" author="Hayashi, T." box="[402,599,1866,1890]" pageId="12" pageNumber="365" pagination="309 - 319" refId="ref10082" refString="Hayashi, T. (2010) The genus Opalimosina Rohacek (Diptera, Sphaeroceridae) from Japan. Medical Entomology and Zoology, 61 (3), 309 - 319. https: // doi. org / 10.7601 / mez. 61.309." type="journal article" year="2010">Hayashi, 2010: 317</bibRefCitation>
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[bothsexes, illustr.].
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<typeStatus id="B624E8A8AC704475FCB1F8FEFCB3A07B" box="[811,861,1866,1890]" pageId="12" pageNumber="365">Type</typeStatus>
locality:
<collectingCountry id="1188169AAC704475FC27F8FEFC17A078" box="[957,1017,1866,1889]" name="Japan" pageId="12" pageNumber="365">Japan</collectingCountry>
,
<collectingRegion id="AB5B98E8AC704475FB9EF8FEFB8CA07B" box="[1028,1122,1866,1890]" country="Japan" name="Okinawa" pageId="12" pageNumber="365">Okinawa</collectingRegion>
,
<location id="6C4000D1AC704475FBF6F8FEFB37A078" LSID="urn:lsid:plazi:treatment:E136E71CAC704474FF0DF955FCBDA08F:6C4000D1AC704475FBF6F8FEFB37A078" box="[1132,1241,1866,1890]" country="Japan" name="Ishigaki Is" pageId="12" pageNumber="365" stateProvince="Okinawa">Ishigaki Is</location>
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.
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</paragraph>
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<paragraph id="6920560AAC704475FF0DF824FAA4A0B3" blockId="12.[151,1437,1936,2034]" box="[151,1354,1936,1962]" pageId="12" pageNumber="365">
<emphasis id="5BEB8A18AC704475FF0DF824FEC7A0B3" bold="true" box="[151,297,1936,1962]" pageId="12" pageNumber="365">Description.</emphasis>
Male (
<figureCitation id="F1A44A8FAC704475FEE2F824FDE3A0B3" box="[376,525,1936,1962]" captionStart="FIGURES 12. 12" captionStartId="13.[151,264,658,682]" captionTargetBox="[354,1234,237,634]" captionTargetId="figure-49@13.[354,1234,237,634]" captionTargetPageId="13" captionText="FIGURES 12. 12AB. O. spathulata Hayashi, 2010 male habitus. 12A, lateral; 12B, dorsal. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14421402" httpUri="https://zenodo.org/record/14421402/files/figure.png" pageId="12" pageNumber="365">Figs. 12AB</figureCitation>
). Body length
<quantity id="AE67FBEFAC704475FD2BF824FCE7A0B3" box="[689,777,1936,1962]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="365" unit="mm" value="1.2">1.2 mm</quantity>
, wing
<quantity id="AE67FBEFAC704475FCC9F824FC42A0B3" box="[851,940,1936,1962]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="365" unit="mm" value="1.1">1.1 mm</quantity>
. General color brown to dark brown.
</paragraph>
<paragraph id="6920560AAC704474FF5DF800FBF7A7CC" blockId="12.[151,1437,1936,2034]" lastBlockId="13.[151,1437,151,213]" lastPageId="13" lastPageNumber="366" pageId="12" pageNumber="365">Head generally dark brown, gena brown; postvertical bristles well developed; 4 interfrontal bristles, foremost one a little shorter; 2 fronto-orbital bristles; 7 fronto-orbital setulae forming a long row between interfrontal bristles and eye. Eye oval, its longest diameter about 2.3 times as long as smallest genal width. Antenna brown, postpedicel slighter brown; arista about 6.3 times as long as postpedicel and shortly ciliated.</paragraph>
<caption id="3DE00682AC714474FF0DFD26FB1DA5B0" ID-DOI="http://doi.org/10.5281/zenodo.14421402" ID-Zenodo-Dep="14421402" box="[151,1267,658,682]" httpUri="https://zenodo.org/record/14421402/files/figure.png" pageId="13" pageNumber="366" startId="13.[151,264,658,682]" targetBox="[354,1234,237,634]" targetPageId="13" targetType="figure">
<paragraph id="6920560AAC714474FF0DFD26FB1DA5B0" blockId="13.[151,1267,658,682]" box="[151,1267,658,682]" pageId="13" pageNumber="366">
<emphasis id="5BEB8A18AC714474FF0DFD26FE6AA5B3" bold="true" box="[151,388,658,682]" pageId="13" pageNumber="366">FIGURES 12. 12AB.</emphasis>
<taxonomicName id="AE9F2D89AC714474FE10FD26FD43A5B3" authority="Hayashi, 2010" authorityName="Hayashi" authorityYear="2010" box="[394,685,658,682]" class="Insecta" family="Sphaeroceridae" genus="Opalimosina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="366" phylum="Arthropoda" rank="species" species="spathulata">
<emphasis id="5BEB8A18AC714474FE10FD26FDFAA5B3" box="[394,532,658,682]" italics="true" pageId="13" pageNumber="366">O. spathulata</emphasis>
<bibRefCitation id="0D0E2BFBAC714474FD80FD26FD43A5B3" author="Hayashi, T." box="[538,685,658,682]" pageId="13" pageNumber="366" pagination="309 - 319" refId="ref10082" refString="Hayashi, T. (2010) The genus Opalimosina Rohacek (Diptera, Sphaeroceridae) from Japan. Medical Entomology and Zoology, 61 (3), 309 - 319. https: // doi. org / 10.7601 / mez. 61.309." type="journal article" year="2010">Hayashi, 2010</bibRefCitation>
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male habitus. 12A, lateral; 12B, dorsal. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="6920560AAC714474FF5DFD6CFE6FA342" blockId="13.[151,1437,728,1943]" pageId="13" pageNumber="366">
Thorax brown. Postpronotal lobe with 1 dorsolateral bristle, 23 small setulae along inner margin and 1 erect setula laterally; 2 postsutural dorsocentral bristles, anterior one short (but longer than dorsocentral setulae in front of it); about 6 rows of irregular acrostichal setulae in front of suture; prescutellar medial acrostichal setulae pair only slightly enlarged (shorter than anterior dorsocentral bristles). Scutellum roundly triangular, with a pair very long bristles, anterolateral seta shorter. Two katepisternal bristles, anterior one minute. Legs brown. Mid tibia ventrally with 1 anteroventral bristle at basal 1/2 and 1 strong apical bristle; hind tibia with 1 very long curved ventral preapical spur. Wing lightly infuscate, veins brownish; costa brown, with a pair of subequal costal bristles at base (outer one longer) and ending at R4+5; R2+3 sinuate, apex distinctly bent up to
<emphasis id="5BEB8A18AC714474FB89FC60FBC8A4F7" box="[1043,1062,980,1006]" italics="true" pageId="13" pageNumber="366">C</emphasis>
; R4+5 gently curved forward;
<emphasis id="5BEB8A18AC714474FA1AFC60FA7DA4F7" box="[1408,1427,980,1006]" italics="true" pageId="13" pageNumber="366">C</emphasis>
- index=0.9, second costal sector is subequal (slightly shorter) to third; outer corners of
<emphasis id="5BEB8A18AC714474FBCBFC4CFB9DA30B" box="[1105,1139,1016,1042]" italics="true" pageId="13" pageNumber="366">dm</emphasis>
rectangular and with short appendages of M1 and M4 beyond
<emphasis id="5BEB8A18AC714474FDB6FBA8FD82A32F" box="[556,620,1052,1078]" italics="true" pageId="13" pageNumber="366">dm-m</emphasis>
;
<emphasis id="5BEB8A18AC714474FDE6FBA9FD1CA32F" box="[636,754,1052,1079]" italics="true" pageId="13" pageNumber="366">r-mdm-m</emphasis>
:
<emphasis id="5BEB8A18AC714474FD63FBA8FCD7A32F" box="[761,825,1052,1078]" italics="true" pageId="13" pageNumber="366">dm-m</emphasis>
=2.1. Alula small, narrow, pointed. Halter with brown knob and paler stem.
</paragraph>
<paragraph id="6920560AAC714474FF5DFBD0FB08A3DF" blockId="13.[151,1437,728,1943]" pageId="13" pageNumber="366">Abdomen brown with dark brown apex. Tergites and sternites sparsely and shortly setulose. T1+2 short and simply sclerotized. S5 (Fig. 13D) posteromedially incised weakly, with two small characteristic patches posteriorly and a weakly sclerotized, semicircular area centrally; otherwise very sparsely and shortly setulose.</paragraph>
<paragraph id="6920560AAC714474FF5DFB64FE31A2FF" blockId="13.[151,1437,728,1943]" pageId="13" pageNumber="366">Genitalia: Epandrium (Figs. 13AB) sparsely and shortly setulose, posterior ventral part with rather dense setosity. Cercus fused with epandrium and projecting ventrally, with a few setulae and 1 long seta. Hypandrium short. Surstylus (Fig. 13C) flat; anterior part bifurcated and pointed, with a few short setae medioventrally; posterior corner blunt, with a few short setae and 2 robust bristles. Basiphallus with slender, apically truncated epiphallus. Distiphallus generally complex and well sclerotized; dorsal part narrows from base and bifurcates at tip (dorsally with small backward pointed structure at each side); ventral part links beneath bifurcated tip of dorsal part and anterodorsally connects with well sclerotized apex (closing medially). Postgonite curved, with a strongly dilated and rounded apex (Fig. 13E).</paragraph>
<paragraph id="6920560AAC714474FF5DFA44FCFAA113" blockId="13.[151,1437,728,1943]" box="[199,788,1520,1547]" pageId="13" pageNumber="366">Female. No specimens were found during this study.</paragraph>
</subSubSection>
<subSubSection id="21850581AC714474FF5DF9A0FDB6A16F" pageId="13" pageNumber="366" type="materials_examined">
<paragraph id="6920560AAC714474FF5DF9A0FDB6A16F" blockId="13.[151,1437,728,1943]" pageId="13" pageNumber="366">
<emphasis id="5BEB8A18AC714474FF5DF9A0FE5AA137" bold="true" box="[199,436,1556,1582]" pageId="13" pageNumber="366">Material examined:</emphasis>
<materialsCitation id="D9F75C57AC714474FE27F9A0FAC4A137" box="[445,1322,1556,1583]" collectingDate="2023-06-14" country="China" elevation="715" location="Badongjinsihou" municipality="Enshi" pageId="13" pageNumber="366" specimenCount="1" specimenCount-male="1" stateProvince="Hubei">
<collectingCountry id="1188169AAC714474FE27F9A0FDF4A137" box="[445,538,1556,1582]" name="China" pageId="13" pageNumber="366">CHINA</collectingCountry>
:
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<collectingRegion id="AB5B98E8AC714474FDBCF9A0FD81A137" box="[550,623,1556,1582]" country="China" name="Hubei" pageId="13" pageNumber="366">Hubei</collectingRegion>
</emphasis>
:
<specimenCount id="7F999D83AC714474FDE5F9A0FD4DA137" box="[639,675,1556,1582]" count="1" pageId="13" pageNumber="366" type="male">1♂</specimenCount>
,
<collectingMunicipality id="8944CC70AC714474FD2BF9A0FD1AA137" box="[689,756,1556,1582]" pageId="13" pageNumber="366">Enshi</collectingMunicipality>
,
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,
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, Jiuzhou Liu
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.
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, Yameng Wang
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, Yishen Xiao (all
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</paragraph>
</subSubSection>
<subSubSection id="21850581AC714474FF5DF934FCBDA08F" pageId="13" pageNumber="366" type="discussion">
<paragraph id="6920560AAC714474FF5DF934FCBDA08F" blockId="13.[151,1437,728,1943]" pageId="13" pageNumber="366">
<emphasis id="5BEB8A18AC714474FF5DF934FEBCA183" bold="true" box="[199,338,1664,1690]" pageId="13" pageNumber="366">Comments:</emphasis>
Details of male sternite 5 and epandrial chaetotaxy recorded here differ from the original description (
<bibRefCitation id="0D0E2BFBAC714474FF05F910FEA8A1A7" author="Hayashi, T." box="[159,326,1700,1726]" pageId="13" pageNumber="366" pagination="309 - 319" refId="ref10082" refString="Hayashi, T. (2010) The genus Opalimosina Rohacek (Diptera, Sphaeroceridae) from Japan. Medical Entomology and Zoology, 61 (3), 309 - 319. https: // doi. org / 10.7601 / mez. 61.309." type="journal article" year="2010">Hayashi, 2010</bibRefCitation>
), as we found S5 has a more weakly sclerotized, semicircular area centrally and the epandrium has dense bristles ventrally and a little more slender. Other details correspond with the original description. This species is similar to
<taxonomicName id="AE9F2D89AC714474FEBFF958FD66A01E" authority="(Rohacek, 1975)" baseAuthorityName="Rohacek" baseAuthorityYear="1975" box="[293,648,1772,1799]" class="Insecta" family="Sphaeroceridae" genus="Opalimosina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="366" phylum="Arthropoda" rank="species" species="calcarifera">
<emphasis id="5BEB8A18AC714474FEBFF958FE2AA01F" box="[293,452,1772,1798]" italics="true" pageId="13" pageNumber="366">O. calcarifera</emphasis>
(
<bibRefCitation id="0D0E2BFBAC714474FE4EF958FD6EA01E" author="Rohacek, J." box="[468,640,1772,1799]" pageId="13" pageNumber="366" pagination="115 - 126" refId="ref10282" refString="Rohacek, J. (1975) Three new species of the genus Limosina Macq. (Sphaeroceridae, Diptera) from Czechoslovakia. Scripta Facultatis Scientiarum Naturalium University J. E. Purkynianae Brunensis, Biologia, 2 (5), 115 - 126." type="journal article" year="1975">Roháček, 1975</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="AE9F2D89AC714474FD5AF958FC1DA01F" authority="Hayashi, 2009" authorityName="Hayashi" authorityYear="2009" box="[704,1011,1772,1798]" class="Insecta" family="Sphaeroceridae" genus="Opalimosina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="366" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="5BEB8A18AC714474FD5AF958FCABA01F" box="[704,837,1772,1798]" italics="true" pageId="13" pageNumber="366">O. australis</emphasis>
<bibRefCitation id="0D0E2BFBAC714474FCD7F958FC1DA01F" author="Hayashi, T." box="[845,1011,1772,1798]" pageId="13" pageNumber="366" pagination="221 - 224" refId="ref10029" refString="Hayashi, T. (2009) A new species of the genus Opalimosina Rohacek (Diptera, Sphaeroceridae) from the Oriental and Australasian Regions. Medical Entomology and Zoology, 60 (3), 221 - 224. https: // doi. org / 10.7601 / mez. 60.221" type="journal article" year="2009">Hayashi, 2009</bibRefCitation>
</taxonomicName>
, but it differs from both these species by its male fifth sternite lacking posteromedial incision, longer and stronger hind tibial spur, and distinctly curved and more dilated postgonite. In
<taxonomicName id="AE9F2D89AC714474FE6CF880FCBBA056" authority="(Rohacek, 1975)" baseAuthorityName="Rohacek" baseAuthorityYear="1975" box="[502,853,1844,1871]" class="Insecta" family="Sphaeroceridae" genus="Opalimosina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="366" phylum="Arthropoda" rank="species" species="calcarifera">
<emphasis id="5BEB8A18AC714474FE6CF880FD7DA057" box="[502,659,1844,1870]" italics="true" pageId="13" pageNumber="366">O. calcarifera</emphasis>
(
<bibRefCitation id="0D0E2BFBAC714474FD3BF880FCA3A056" author="Rohacek, J." box="[673,845,1844,1871]" pageId="13" pageNumber="366" pagination="115 - 126" refId="ref10282" refString="Rohacek, J. (1975) Three new species of the genus Limosina Macq. (Sphaeroceridae, Diptera) from Czechoslovakia. Scripta Facultatis Scientiarum Naturalium University J. E. Purkynianae Brunensis, Biologia, 2 (5), 115 - 126." type="journal article" year="1975">Roháček, 1975</bibRefCitation>
)
</taxonomicName>
, sternite 5 has a shallow incision, the hind tibial spur is shorter and the postgonite is straight. In
<taxonomicName id="AE9F2D89AC714474FDE3F8ECFC46A06B" authority="Hayashi, 2009" authorityName="Hayashi" authorityYear="2009" box="[633,936,1880,1906]" class="Insecta" family="Sphaeroceridae" genus="Opalimosina" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="366" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="5BEB8A18AC714474FDE3F8ECFD10A06B" box="[633,766,1880,1906]" italics="true" pageId="13" pageNumber="366">O. australis</emphasis>
<bibRefCitation id="0D0E2BFBAC714474FC9CF8ECFC46A06B" author="Hayashi, T." box="[774,936,1880,1906]" pageId="13" pageNumber="366" pagination="221 - 224" refId="ref10029" refString="Hayashi, T. (2009) A new species of the genus Opalimosina Rohacek (Diptera, Sphaeroceridae) from the Oriental and Australasian Regions. Medical Entomology and Zoology, 60 (3), 221 - 224. https: // doi. org / 10.7601 / mez. 60.221" type="journal article" year="2009">Hayashi, 2009</bibRefCitation>
</taxonomicName>
S5 has a deep and narrow incision, the hind tibial spur is shorter and the postgonite is only slightly curved.
</paragraph>
</subSubSection>
</treatment>
</document>