treatments-xml/data/E1/15/87/E115879CFF9AFF86FF7EF944FE18F9B7.xml
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<mods:title id="DD4365D2668D6EF447E25BED45CC17F0">- On- two- new- species- of- Nannopus- Brady, - 1880 - (Copepoda: - Harpacticoida Nannopodidae) - from-intertidal-mudflats-of-the-Korean-west-coast- (Yellow-Sea)</mods:title>
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<mods:namePart id="1FDF03D324D44D9292C9BD9C687D3642">Vakati, Vinod</mods:namePart>
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<mods:affiliation id="03AE1EF0FE3A4C8A4AD4E78C944D6AA9">Laboratory of Conservation and Utilization of Fish Resources, College of Life Sciences, Neijiang Normal University, Neijiang 641100, China. vinodbio 88 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4492 - 9981 &amp; Department of Life Science, College of Natural Sciences, Hanyang University, Seoul 04763, Korea &amp; Fourth Institute of Oceanography, Ministry of Natural Resources, 26 New Century Avenue, Beihai, 536000, China</mods:affiliation>
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<mods:namePart id="BED7EF1D9C88B629A08A510A77392E43">Lee, Wonchoel</mods:namePart>
<mods:affiliation id="60BE6FF3E59A312799511B09C9C0A44D">Department of Life Science, College of Natural Sciences, Hanyang University, Seoul 04763, Korea</mods:affiliation>
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<treatment id="E115879CFF9AFF86FF7EF944FE18F9B7" ID-DOI="http://doi.org/10.5281/zenodo.5563935" ID-GBIF-Taxon="188623289" ID-Zenodo-Dep="5563935" LSID="urn:lsid:plazi:treatment:E115879CFF9AFF86FF7EF944FE18F9B7" httpUri="http://treatment.plazi.org/id/E115879CFF9AFF86FF7EF944FE18F9B7" lastPageId="8" lastPageNumber="537" pageId="2" pageNumber="531">
<subSubSection id="21A66501FF9AFF8CFF7EF944FEA8F978" pageId="2" pageNumber="531" type="nomenclature">
<paragraph id="6903368AFF9AFF8CFF7EF944FE53F927" blockId="2.[151,489,1699,1762]" box="[151,489,1699,1726]" pageId="2" pageNumber="531">
<heading id="324B81E6FF9AFF8CFF7EF944FE53F927" bold="true" box="[151,489,1699,1726]" fontSize="11" level="1" pageId="2" pageNumber="531" reason="1">
<emphasis id="5BC8EA98FF9AFF8CFF7EF944FE53F927" bold="true" box="[151,489,1699,1726]" pageId="2" pageNumber="531">
<taxonomicName id="AEBC4D09FF9AFF8CFF7EF944FE3EF924" authorityName="Vakati &amp; Lee" authorityYear="2021" box="[151,388,1699,1726]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="5BC8EA98FF9AFF8CFF7EF944FE3EF924" bold="true" box="[151,388,1699,1726]" italics="true" pageId="2" pageNumber="531">Nannopus robustus</emphasis>
</taxonomicName>
­sp.­nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="6903368AFF9AFF8CFF7EF920FEA8F978" blockId="2.[151,489,1699,1762]" box="[151,274,1735,1762]" pageId="2" pageNumber="531">
(
<figureCitation id="F1872A0FFF9AFF8CFF76F920FEB0F978" box="[159,266,1735,1762]" captionStart-0="FIGURE­4" captionStart-1="FIGURE­5" captionStartId-0="6.[151,256,1962,1986]" captionStartId-1="7.[151,256,1943,1967]" captionTargetBox-0="[196,1392,193,1926]" captionTargetBox-1="[193,1393,190,1914]" captionTargetId-0="figure-21@6.[182,1405,181,1938]" captionTargetId-1="figure-20@7.[183,1404,181,1919]" captionTargetPageId-1="7" captionText-0="FIGURE­4.­Nannopus robustus sp.­nov. (holotype ♂). A, habitus, dorsal; B, habitus, lateral." captionText-1="FIGURE­5.­Nannopus robustus sp.­nov. (holotype ♂). A, anal somite and caudal rami, dorsal (cylindrical shape arrowed); B, anal somite and caudal rami, ventral (cylindrical shape arrowed); C, anal somite and caudal ramus, lateral (cylindrical shape arrowed); D, P2 (recurved spine arrowed); E, P2 endopod." figureDoi-0="http://doi.org/10.5281/zenodo.5576344" figureDoi-1="http://doi.org/10.5281/zenodo.5576346" httpUri-0="https://zenodo.org/record/5576344/files/figure.png" httpUri-1="https://zenodo.org/record/5576346/files/figure.png" pageId="2" pageNumber="531">Figs. 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="21A66501FF9AFF8CFF7EF8E8FE18F8D7" pageId="2" pageNumber="531" type="description">
<paragraph id="6903368AFF9AFF8CFF7EF8E8FE18F8D7" blockId="2.[151,1436,1807,2013]" pageId="2" pageNumber="531">
<emphasis id="5BC8EA98FF9AFF8CFF7EF8E8FE87F8B3" bold="true" box="[151,317,1807,1833]" pageId="2" pageNumber="531">Type­ locality.</emphasis>
<materialsCitation id="D9D43CD7FF9AFF8CFEA5F8E8FE27F8D7" ID-GBIF-Occurrence="3382722302" country="South Korea" county="Yellow Sea" latitude="37.237946" location="Intertidal" longLatPrecision="1" longitude="126.47134" municipality="Ongjin-gun" pageId="2" pageNumber="531" specimenCount="1">
<location id="6C636051FF9AFF8CFEA5F8E8FE09F8B3" LSID="urn:lsid:plazi:treatment:E115879CFF9AFF86FF7EF944FE18F9B7:6C636051FF9AFF8CFEA5F8E8FE09F8B3" box="[332,435,1807,1833]" country="South Korea" county="Yellow Sea" latitude="37.237946" longLatPrecision="1" longitude="126.47134" municipality="Ongjin-gun" name="Intertidal" pageId="2" pageNumber="531">Intertidal</location>
mudflat, St-8—
<location id="6C636051FF9AFF8CFD9FF8E8FCE4F8B3" LSID="urn:lsid:plazi:treatment:E115879CFF9AFF86FF7EF944FE18F9B7:6C636051FF9AFF8CFD9FF8E8FCE4F8B3" box="[630,862,1807,1833]" country="South Korea" county="Yellow Sea" latitude="37.237946" longLatPrecision="1" longitude="126.47134" municipality="Ongjin-gun" name="Yeongheung-myeon" pageId="2" pageNumber="531">Yeongheung-myeon</location>
,
<collectingMunicipality id="8967ACF0FF9AFF8CFC98F8E8FC4FF8B3" box="[881,1013,1807,1833]" pageId="2" pageNumber="531">Ongjin-gun</collectingMunicipality>
,
<collectingCountry id="11AB761AFF9AFF8CFBE1F8F7FBF5F8B0" box="[1032,1103,1808,1834]" name="South Korea" pageId="2" pageNumber="531">Korea</collectingCountry>
,
<collectingCounty id="80624E06FF9AFF8CFB88F8E8FB51F8B3" box="[1121,1259,1807,1833]" pageId="2" pageNumber="531">Yellow Sea</collectingCounty>
,
<geoCoordinate id="0C88504DFF9AFF8CFB17F8E8FA26F8B3" box="[1278,1436,1807,1833]" degrees="37" direction="north" minutes="14" orientation="latitude" pageId="2" pageNumber="531" precision="1" seconds="16.6" value="37.237946">37°1416.6”N</geoCoordinate>
<geoCoordinate id="0C88504DFF9AFF8CFF7EF8D4FEFAF8D7" box="[151,320,1843,1869]" degrees="126" direction="east" minutes="28" orientation="longitude" pageId="2" pageNumber="531" precision="1" seconds="16.8" value="126.47134">126°2816.8”E</geoCoordinate>
(
<figureCitation id="F1872A0FFF9AFF8CFEB9F8D4FE2BF8D7" box="[336,401,1843,1869]" captionStart="FIGURE­1" captionStartId="3.[151,256,1943,1967]" captionTargetBox="[192,1395,181,1919]" captionTargetId="figure-20@3.[192,1395,181,1919]" captionText="FIGURE­1. Map showing the sampling localities and distribution of all valid Korean species identified and described so far. The black line between Stations 6a and 6b represents a man-made road with no gap for water entry from either side. Stations 1 to 15 refer to the study sites in Vakati et al. (2019), station 16 refers to the study site in Kim et al. (2017)." figureDoi="http://doi.org/10.5281/zenodo.5576335" httpUri="https://zenodo.org/record/5576335/files/figure.png" pageId="2" pageNumber="531">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="21A66501FF9AFF8CFF2EF8B0FDADF80F" pageId="2" pageNumber="531" type="materials_examined">
<paragraph id="6903368AFF9AFF8CFF2EF8B0FDADF80F" blockId="2.[151,1436,1807,2013]" pageId="2" pageNumber="531">
<emphasis id="5BC8EA98FF9AFF8CFF2EF8B0FEC3F8EB" bold="true" box="[199,377,1879,1905]" pageId="2" pageNumber="531">Type­ material.</emphasis>
<typeStatus id="B6078828FF9AFF8CFE6AF8B0FE4AF8EB" box="[387,496,1879,1905]" pageId="2" pageNumber="531" type="holotype">Holotype</typeStatus>
:
<specimenCount id="7FBAFD03FF9AFF8CFE17F8B0FDF6F8EB" box="[510,588,1879,1905]" pageId="2" pageNumber="531" type="male">1 male</specimenCount>
adult in 70% ethanol (NIBRIV0000860821). Samples were collected by Vinod Vakati,
<date id="1D02104AFF9AFF8CFED1F89CFDAEF80F" box="[312,532,1915,1941]" pageId="2" pageNumber="531" value="2013-11-20">20 November 2013</date>
.
</paragraph>
</subSubSection>
<subSubSection id="21A66501FF9AFF8CFF2EF878FD8BF847" pageId="2" pageNumber="531" type="etymology">
<paragraph id="6903368AFF9AFF8CFF2EF878FD8BF847" blockId="2.[151,1436,1807,2013]" pageId="2" pageNumber="531">
<emphasis id="5BC8EA98FF9AFF8CFF2EF878FEF4F823" bold="true" box="[199,334,1951,1977]" pageId="2" pageNumber="531">Etymology.</emphasis>
The species epithet is derived from the Latin “robustus”, referring to the very strong and sturdy distal-most apical spine of P2 exp-2.
</paragraph>
</subSubSection>
<caption id="3DC36602FF9BFF8DFF7EF870FB7BF862" ID-DOI="http://doi.org/10.5281/zenodo.5576335" ID-Zenodo-Dep="5576335" httpUri="https://zenodo.org/record/5576335/files/figure.png" pageId="3" pageNumber="532" startId="3.[151,256,1943,1967]" targetBox="[192,1395,181,1919]" targetPageId="3">
<paragraph id="6903368AFF9BFF8DFF7EF870FB7BF862" blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="532">
<emphasis id="5BC8EA98FF9BFF8DFF7EF870FEAEF835" bold="true" box="[151,276,1943,1967]" pageId="3" pageNumber="532">FIGURE­1.</emphasis>
Map showing the sampling localities and distribution of all valid Korean species identified and described so far. The black line between Stations 6a and 6b represents a man-made road with no gap for water entry from either side. Stations 1 to 15 refer to the study sites in
<bibRefCitation id="0D2D4B7BFF9BFF8DFE38F807FD2CF862" author="Vakati, V. &amp; Eyun, S. &amp; Lee, W." box="[465,662,2016,2040]" pageId="3" pageNumber="532" pagination="366 - 379" refId="ref13822" refString="Vakati, V., Eyun, S. &amp; Lee, W. (2019) Unraveling the intricate biodiversity of the benthic harpacticoid genus Nannopus (Copepoda, Harpacticoida, Nannopodidae) in Korean waters. Molecular Phylogenetics and Evolution, 130, 366 - 379. https: // doi. org / 10.1016 / j. ympev. 2018.10.004" type="journal article" year="2019">
Vakati
<emphasis id="5BC8EA98FF9BFF8DFDF1F807FDF1F862" box="[536,587,2016,2040]" italics="true" pageId="3" pageNumber="532">et al.</emphasis>
(2019)
</bibRefCitation>
, station 16 refers to the study site in
<bibRefCitation id="0D2D4B7BFF9BFF8DFBE3F807FB07F862" author="Kim, J. G. &amp; Choi, H. K. &amp; Yoon, S. M." box="[1034,1213,2016,2040]" pageId="3" pageNumber="532" pagination="1351 - 1365" refId="ref13550" refString="Kim, J. G., Choi, H. K. &amp; Yoon, S. M. (2017) A reappraisal of the genera Nannopus Brady, 1880 and Ilyophilus Lilljeborg, 1902 (Copepoda, Harpacticoida, Nannopodidae) with a description of N. parvipilis sp. nov. from South Korea. Crustaceana, 90, 1351 - 1365. https: // doi. org / 10.1163 / 15685403 - 00003700" type="journal article" year="2017">
Kim
<emphasis id="5BC8EA98FF9BFF8DFBD7F807FBCBF862" box="[1086,1137,2016,2040]" italics="true" pageId="3" pageNumber="532">et al.</emphasis>
(2017)
</bibRefCitation>
.
</paragraph>
</caption>
<caption id="3DC36602FF9CFF8AFF7EF85AFBE7F84C" ID-DOI="http://doi.org/10.5281/zenodo.5576338" ID-Zenodo-Dep="5576338" box="[151,1117,1981,2006]" httpUri="https://zenodo.org/record/5576338/files/figure.png" pageId="4" pageNumber="533" startId="4.[151,256,1981,2005]" targetBox="[304,1289,194,1941]" targetPageId="4">
<paragraph id="6903368AFF9CFF8AFF7EF85AFBE7F84C" blockId="4.[151,1117,1981,2006]" box="[151,1117,1981,2006]" pageId="4" pageNumber="533">
<emphasis id="5BC8EA98FF9CFF8AFF7EF85AFEA3F84C" bold="true" box="[151,281,1981,2006]" pageId="4" pageNumber="533">FIGURE­2.­</emphasis>
<taxonomicName id="AEBC4D09FF9CFF8AFEF0F859FE4DF84C" authorityName="Vakati &amp; Lee" authorityYear="2021" box="[281,503,1982,2006]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="5BC8EA98FF9CFF8AFEF0F859FE4DF84C" box="[281,503,1982,2006]" italics="true" pageId="4" pageNumber="533">Nannopus cylindricus</emphasis>
</taxonomicName>
<emphasis id="5BC8EA98FF9CFF8AFE14F859FDF4F84C" bold="true" box="[509,590,1982,2006]" pageId="4" pageNumber="533">sp.­nov.</emphasis>
(
<typeStatus id="B6078828FF9CFF8AFDB2F859FD0FF84C" box="[603,693,1982,2006]" pageId="4" pageNumber="533" type="holotype">holotype</typeStatus>
<specimenCount id="7FBAFD03FF9CFF8AFD55F859FD74F84C" box="[700,718,1982,2006]" pageId="4" pageNumber="533" type="female"></specimenCount>
). A, habitus, dorsal; B, habitus, lateral.
</paragraph>
</caption>
<caption id="3DC36602FF9DFF8BFF7EF870FF40F862" ID-DOI="http://doi.org/10.5281/zenodo.5576340" ID-Zenodo-Dep="5576340" httpUri="https://zenodo.org/record/5576340/files/figure.png" pageId="5" pageNumber="534" startId="5.[151,256,1943,1967]" targetBox="[189,1401,192,1911]" targetPageId="5">
<paragraph id="6903368AFF9DFF8BFF7EF870FF40F862" blockId="5.[151,1437,1943,2040]" pageId="5" pageNumber="534">
<emphasis id="5BC8EA98FF9DFF8BFF7EF870FEA0F82A" bold="true" box="[151,282,1943,1968]" pageId="5" pageNumber="534">FIGURE­3.­</emphasis>
<taxonomicName id="AEBC4D09FF9DFF8BFEF2F87FFE40F82A" authorityName="Vakati &amp; Lee" authorityYear="2021" box="[283,506,1944,1968]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="5" pageNumber="534" phylum="Arthropoda" rank="species" species="cylindricus">
<emphasis id="5BC8EA98FF9DFF8BFEF2F87FFE40F82A" box="[283,506,1944,1968]" italics="true" pageId="5" pageNumber="534">Nannopus cylindricus</emphasis>
</taxonomicName>
<emphasis id="5BC8EA98FF9DFF8BFDE8F87FFDE9F82A" bold="true" box="[513,595,1944,1968]" pageId="5" pageNumber="534">sp.­nov.</emphasis>
(
<typeStatus id="B6078828FF9DFF8BFD88F87FFD01F82A" box="[609,699,1944,1968]" pageId="5" pageNumber="534" type="holotype">holotype</typeStatus>
<specimenCount id="7FBAFD03FF9DFF8BFD2AF87FFD6FF82A" box="[707,725,1944,1968]" pageId="5" pageNumber="534" type="female"></specimenCount>
). A, anal somite and caudal rami, dorsal (cylindrical shape arrowed); B, anal somite and caudal rami, ventral (cylindrical shape arrowed); C, anal somite and caudal ramus, lateral (cylindrical shape arrowed).
</paragraph>
</caption>
<caption id="3DC36602FF9EFF88FF7EF84DFBF9F858" ID-DOI="http://doi.org/10.5281/zenodo.5576344" ID-Zenodo-Dep="5576344" box="[151,1091,1962,1986]" httpUri="https://zenodo.org/record/5576344/files/figure.png" pageId="6" pageNumber="535" startId="6.[151,256,1962,1986]" targetBox="[196,1392,193,1926]" targetPageId="6">
<paragraph id="6903368AFF9EFF88FF7EF84DFBF9F858" blockId="6.[151,1091,1962,1986]" box="[151,1091,1962,1986]" pageId="6" pageNumber="535">
<emphasis id="5BC8EA98FF9EFF88FF7EF84DFEA3F858" bold="true" box="[151,281,1962,1986]" pageId="6" pageNumber="535">FIGURE­4.­</emphasis>
<taxonomicName id="AEBC4D09FF9EFF88FEF0F84CFE66F858" authorityName="Vakati &amp; Lee" authorityYear="2021" box="[281,476,1962,1986]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="5BC8EA98FF9EFF88FEF0F84CFE66F858" box="[281,476,1962,1986]" italics="true" pageId="6" pageNumber="535">Nannopus robustus</emphasis>
</taxonomicName>
<emphasis id="5BC8EA98FF9EFF88FE0AF84DFD8EF858" bold="true" box="[483,564,1962,1986]" pageId="6" pageNumber="535">sp.­nov.</emphasis>
(
<typeStatus id="B6078828FF9EFF88FDA8F84DFD21F858" box="[577,667,1962,1986]" pageId="6" pageNumber="535" type="holotype">holotype</typeStatus>
<specimenCount id="7FBAFD03FF9EFF88FD4BF84DFD0DF858" box="[674,695,1962,1986]" pageId="6" pageNumber="535" type="male"></specimenCount>
). A, habitus, dorsal; B, habitus, lateral.
</paragraph>
</caption>
<caption id="3DC36602FF9FFF89FF7EF870FD52F862" ID-DOI="http://doi.org/10.5281/zenodo.5576346" ID-Zenodo-Dep="5576346" httpUri="https://zenodo.org/record/5576346/files/figure.png" pageId="7" pageNumber="536" startId="7.[151,256,1943,1967]" targetBox="[193,1393,190,1914]" targetPageId="7">
<paragraph id="6903368AFF9FFF89FF7EF870FD52F862" blockId="7.[151,1437,1943,2040]" pageId="7" pageNumber="536">
<emphasis id="5BC8EA98FF9FFF89FF7EF870FEA0F82A" bold="true" box="[151,282,1943,1968]" pageId="7" pageNumber="536">FIGURE­5.­</emphasis>
<taxonomicName id="AEBC4D09FF9FFF89FEF2F87FFE65F82A" authorityName="Vakati &amp; Lee" authorityYear="2021" box="[283,479,1944,1968]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="5BC8EA98FF9FFF89FEF2F87FFE65F82A" box="[283,479,1944,1968]" italics="true" pageId="7" pageNumber="536">Nannopus robustus</emphasis>
</taxonomicName>
<emphasis id="5BC8EA98FF9FFF89FE0FF87FFD8DF82A" bold="true" box="[486,567,1944,1968]" pageId="7" pageNumber="536">sp.­nov.</emphasis>
(
<typeStatus id="B6078828FF9FFF89FDACF87FFD25F82A" box="[581,671,1944,1968]" pageId="7" pageNumber="536" type="holotype">holotype</typeStatus>
<specimenCount id="7FBAFD03FF9FFF89FD4EF87FFD06F82A" box="[679,700,1944,1968]" pageId="7" pageNumber="536" type="male"></specimenCount>
). A, anal somite and caudal rami, dorsal (cylindrical shape arrowed); B, anal somite and caudal rami, ventral (cylindrical shape arrowed); C, anal somite and caudal ramus, lateral (cylindrical shape arrowed); D, P2 (recurved spine arrowed); E, P2 endopod.
</paragraph>
</caption>
<subSubSection id="21A66501FF90FF86FF2EFF71FE18F9B7" pageId="8" pageNumber="537" type="etymology">
<paragraph id="6903368AFF90FF86FF2EFF71FD35FE13" blockId="8.[151,1437,150,1581]" pageId="8" pageNumber="537">
<emphasis id="5BC8EA98FF90FF86FF2EFF71FD7BFF2B" bold="true" box="[199,705,150,177]" pageId="8" pageNumber="537">
Description­ of­ male­(based­ on­
<typeStatus id="B6078828FF90FF86FDAAFF71FD17FF2A" box="[579,685,150,176]" pageId="8" pageNumber="537" type="holotype">holotype</typeStatus>
).­
</emphasis>
Body fusiform (
<figureCitation id="F1872A0FFF90FF86FC96FF70FC41FF2B" box="[895,1019,151,177]" captionStart="FIGURE­4" captionStartId="6.[151,256,1962,1986]" captionTargetBox="[196,1392,193,1926]" captionTargetId="figure-21@6.[182,1405,181,1938]" captionText="FIGURE­4.­Nannopus robustus sp.­nov. (holotype ♂). A, habitus, dorsal; B, habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.5576344" httpUri="https://zenodo.org/record/5576344/files/figure.png" pageId="8" pageNumber="537">Fig. 4AB</figureCitation>
) narrow, total body length 461 μm, measured from tip of rostrum to posterior margin of caudal rami. Maximum width 138 μm, measured at posterior margin of cephalothorax. Body slightly slender, tapering posteriorly. Urosome/prosome length ratio 0.71 (
<figureCitation id="F1872A0FFF90FF86FAD5FF38FA2EFF63" box="[1340,1428,223,249]" captionStart="FIGURE­4" captionStartId="6.[151,256,1962,1986]" captionTargetBox="[196,1392,193,1926]" captionTargetId="figure-21@6.[182,1405,181,1938]" captionText="FIGURE­4.­Nannopus robustus sp.­nov. (holotype ♂). A, habitus, dorsal; B, habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.5576344" httpUri="https://zenodo.org/record/5576344/files/figure.png" pageId="8" pageNumber="537">Fig. 4A</figureCitation>
) and body length/width ratio about 3.3. Body ornamentation as in
<taxonomicName id="AEBC4D09FF90FF86FC9CFEE4FC48FE86" box="[885,1010,259,284]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="5BC8EA98FF90FF86FC9CFEE4FC48FE86" box="[885,1010,259,284]" italics="true" pageId="8" pageNumber="537">N. serratus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0D2D4B7BFF90FF86FBC5FEE4FA94FE87" author="Vakati, V. &amp; Lee, W." box="[1068,1326,259,285]" pageId="8" pageNumber="537" pagination="1 - 66" refId="ref13958" refString="Vakati, V. &amp; Lee, W. (2017) Five new species of the genus Nannopus (Copepoda: Harpacticoida: Nannopodidae) from intertidal mudflats of the Korean West Coast (Yellow Sea). Zootaxa, 4360 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4360.1.1" type="journal article" year="2017">Vakati &amp; Lee 2017: 24</bibRefCitation>
, 27, Figs. 17AF, 19AD). Each urosomite with rows of slim setules ventrally as in
<taxonomicName id="AEBC4D09FF90FF86FC32FEC0FBE3FEDA" box="[987,1113,295,320]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="5BC8EA98FF90FF86FC32FEC0FBE3FEDA" box="[987,1113,295,320]" italics="true" pageId="8" pageNumber="537">N. serratus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0D2D4B7BFF90FF86FB7DFEC0FA2DFEDB" author="Vakati, V. &amp; Lee, W." box="[1172,1431,295,321]" pageId="8" pageNumber="537" pagination="1 - 66" refId="ref13958" refString="Vakati, V. &amp; Lee, W. (2017) Five new species of the genus Nannopus (Copepoda: Harpacticoida: Nannopodidae) from intertidal mudflats of the Korean West Coast (Yellow Sea). Zootaxa, 4360 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4360.1.1" type="journal article" year="2017">Vakati &amp; Lee 2017: 24</bibRefCitation>
, Fig. 17F). Cephalothorax (
<figureCitation id="F1872A0FFF90FF86FE2DFEACFDA7FEFF" box="[452,541,331,357]" captionStart="FIGURE­4" captionStartId="6.[151,256,1962,1986]" captionTargetBox="[196,1392,193,1926]" captionTargetId="figure-21@6.[182,1405,181,1938]" captionText="FIGURE­4.­Nannopus robustus sp.­nov. (holotype ♂). A, habitus, dorsal; B, habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.5576344" httpUri="https://zenodo.org/record/5576344/files/figure.png" pageId="8" pageNumber="537">Fig. 4A</figureCitation>
) bell-shaped, slightly narrow, comprising 36% of total body length, with a few paired sensilla and serrated posterior margin.
</paragraph>
<paragraph id="6903368AFF90FF86FF2EFE74FC1AFE34" blockId="8.[151,1437,150,1581]" box="[199,928,403,430]" pageId="8" pageNumber="537">
Rostrum as in
<taxonomicName id="AEBC4D09FF90FF86FE83FE74FE5CFE36" box="[362,486,403,428]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="5BC8EA98FF90FF86FE83FE74FE5CFE36" box="[362,486,403,428]" italics="true" pageId="8" pageNumber="537">N. serratus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0D2D4B7BFF90FF86FDC8FE74FC98FE37" author="Vakati, V. &amp; Lee, W." box="[545,802,403,429]" pageId="8" pageNumber="537" pagination="1 - 66" refId="ref13958" refString="Vakati, V. &amp; Lee, W. (2017) Five new species of the genus Nannopus (Copepoda: Harpacticoida: Nannopodidae) from intertidal mudflats of the Korean West Coast (Yellow Sea). Zootaxa, 4360 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4360.1.1" type="journal article" year="2017">Vakati &amp; Lee 2017: 23</bibRefCitation>
, Fig. 16C).
</paragraph>
<paragraph id="6903368AFF90FF86FF2EFE50FED3FD83" blockId="8.[151,1437,150,1581]" pageId="8" pageNumber="537">
Prosome (
<figureCitation id="F1872A0FFF90FF86FED5FE50FE03FE4B" box="[316,441,439,465]" captionStart="FIGURE­4" captionStartId="6.[151,256,1962,1986]" captionTargetBox="[196,1392,193,1926]" captionTargetId="figure-21@6.[182,1405,181,1938]" captionText="FIGURE­4.­Nannopus robustus sp.­nov. (holotype ♂). A, habitus, dorsal; B, habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.5576344" httpUri="https://zenodo.org/record/5576344/files/figure.png" pageId="8" pageNumber="537">Fig. 4AB</figureCitation>
) 4-segmented, comprising cephalothorax and three subequal free pedigerous somites. P2-bearing somite with two long medial sensilla dorsally, posterior margins of prosomites serrated, and with few sensilla as figured.
</paragraph>
<paragraph id="6903368AFF90FF86FF2EFDC4FDB9FD6B" blockId="8.[151,1437,150,1581]" pageId="8" pageNumber="537">
Urosome (
<figureCitation id="F1872A0FFF90FF86FED4FDC4FE0CFDA7" box="[317,438,547,573]" captionStart="FIGURE­4" captionStartId="6.[151,256,1962,1986]" captionTargetBox="[196,1392,193,1926]" captionTargetId="figure-21@6.[182,1405,181,1938]" captionText="FIGURE­4.­Nannopus robustus sp.­nov. (holotype ♂). A, habitus, dorsal; B, habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.5576344" httpUri="https://zenodo.org/record/5576344/files/figure.png" pageId="8" pageNumber="537">Fig. 4AB</figureCitation>
) 5-segmented, comprising P5-bearing somite, genital double-somite, two free abdominal somites, and anal somite. Posterior margins of urosomites serrated dorsally and ventrally. Posterior margin of genital double-somite and following somite with pair of strong sensilla ventrally as in
<taxonomicName id="AEBC4D09FF90FF86FBE3FD8CFB3CFD1E" box="[1034,1158,619,644]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="5BC8EA98FF90FF86FBE3FD8CFB3CFD1E" box="[1034,1158,619,644]" italics="true" pageId="8" pageNumber="537">N. serratus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0D2D4B7BFF90FF86FB29FD8CFF0FFD33" author="Vakati, V. &amp; Lee, W." pageId="8" pageNumber="537" pagination="1 - 66" refId="ref13958" refString="Vakati, V. &amp; Lee, W. (2017) Five new species of the genus Nannopus (Copepoda: Harpacticoida: Nannopodidae) from intertidal mudflats of the Korean West Coast (Yellow Sea). Zootaxa, 4360 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4360.1.1" type="journal article" year="2017">Vakati &amp; Lee 2017: 29</bibRefCitation>
, arrowed in Fig. 21B). Each somite with one transverse row of spinules along posterior margin ventrally. Genital double-somite as in
<taxonomicName id="AEBC4D09FF90FF86FE92FD54FE42FD56" box="[379,504,691,716]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="5BC8EA98FF90FF86FE92FD54FE42FD56" box="[379,504,691,716]" italics="true" pageId="8" pageNumber="537">N. serratus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0D2D4B7BFF90FF86FDDAFD54FC8FFD57" author="Vakati, V. &amp; Lee, W." box="[563,821,691,717]" pageId="8" pageNumber="537" pagination="1 - 66" refId="ref13958" refString="Vakati, V. &amp; Lee, W. (2017) Five new species of the genus Nannopus (Copepoda: Harpacticoida: Nannopodidae) from intertidal mudflats of the Korean West Coast (Yellow Sea). Zootaxa, 4360 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4360.1.1" type="journal article" year="2017">Vakati &amp; Lee 2017: 29</bibRefCitation>
, Fig. 21B), clearly distinct dorsally and ventrally, with serrated dorsal posterior margin.
</paragraph>
<paragraph id="6903368AFF90FF86FF2EFD1CFB3EFCA3" blockId="8.[151,1437,150,1581]" pageId="8" pageNumber="537">
Anal somite (
<figureCitation id="F1872A0FFF90FF86FEB7FD1CFE78FC8F" box="[350,450,763,789]" captionStart="FIGURE­4" captionStartId="6.[151,256,1962,1986]" captionTargetBox="[196,1392,193,1926]" captionTargetId="figure-21@6.[182,1405,181,1938]" captionText="FIGURE­4.­Nannopus robustus sp.­nov. (holotype ♂). A, habitus, dorsal; B, habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.5576344" httpUri="https://zenodo.org/record/5576344/files/figure.png" pageId="8" pageNumber="537">Figs. 4A</figureCitation>
, 5AC) almost as long as wide with unique pattern of surface ornamentation dorsally as in
<taxonomicName id="AEBC4D09FF90FF86FF5DFCF8FE8BFCA2" box="[180,305,799,824]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="5BC8EA98FF90FF86FF5DFCF8FE8BFCA2" box="[180,305,799,824]" italics="true" pageId="8" pageNumber="537">N. serratus</emphasis>
</taxonomicName>
, ventral surface with six spinules (see
<bibRefCitation id="0D2D4B7BFF90FF86FD0BFCF8FC59FCA3" author="Vakati, V. &amp; Lee, W." box="[738,995,799,825]" pageId="8" pageNumber="537" pagination="1 - 66" refId="ref13958" refString="Vakati, V. &amp; Lee, W. (2017) Five new species of the genus Nannopus (Copepoda: Harpacticoida: Nannopodidae) from intertidal mudflats of the Korean West Coast (Yellow Sea). Zootaxa, 4360 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4360.1.1" type="journal article" year="2017">Vakati &amp; Lee 2017: 29</bibRefCitation>
, Fig. 21CD).
</paragraph>
<paragraph id="6903368AFF90FF86FF2EFCA4FF54FBE7" blockId="8.[151,1437,150,1581]" pageId="8" pageNumber="537">
Caudal rami (
<figureCitation id="F1872A0FFF90FF86FE8EFCA4FE59FCC7" box="[359,483,834,861]" captionStart="FIGURE­5" captionStartId="7.[151,256,1943,1967]" captionTargetBox="[193,1393,190,1914]" captionTargetId="figure-20@7.[183,1404,181,1919]" captionText="FIGURE­5.­Nannopus robustus sp.­nov. (holotype ♂). A, anal somite and caudal rami, dorsal (cylindrical shape arrowed); B, anal somite and caudal rami, ventral (cylindrical shape arrowed); C, anal somite and caudal ramus, lateral (cylindrical shape arrowed); D, P2 (recurved spine arrowed); E, P2 endopod." figureDoi="http://doi.org/10.5281/zenodo.5576346" httpUri="https://zenodo.org/record/5576346/files/figure.png" pageId="8" pageNumber="537">Fig. 5AC</figureCitation>
) square (dorsally) or sub-cylindrical (ventrally), almost as long as wide (in dorsal view), almost twice as long as wide (in ventral view), and with row of spinules along posteroventral margins. Caudal ramus bearing seven setae: seta I pinnate, shorter than ramus width, and located anterolaterally; seta II pinnate, almost as long as ramus length, and inserted anterolaterally midway along outer margin; seta III pinnate, as long as ramus length, and inserted anterolaterally midway along outer margin; seta IV bipinnate, 0.5 times as long as seta V, located at outer posterior margin and wide at proximal region; seta V strongest, bipinnate with proximal part heavily cylindrical and smooth (arrowed in
<figureCitation id="F1872A0FFF90FF86FD97FBFCFD4DFBAF" box="[638,759,1050,1077]" captionStart="FIGURE­5" captionStartId="7.[151,256,1943,1967]" captionTargetBox="[193,1393,190,1914]" captionTargetId="figure-20@7.[183,1404,181,1919]" captionText="FIGURE­5.­Nannopus robustus sp.­nov. (holotype ♂). A, anal somite and caudal rami, dorsal (cylindrical shape arrowed); B, anal somite and caudal rami, ventral (cylindrical shape arrowed); C, anal somite and caudal ramus, lateral (cylindrical shape arrowed); D, P2 (recurved spine arrowed); E, P2 endopod." figureDoi="http://doi.org/10.5281/zenodo.5576346" httpUri="https://zenodo.org/record/5576346/files/figure.png" pageId="8" pageNumber="537">Fig. 5AC</figureCitation>
), and located at inner posterior margin; seta VI naked, small, and located at inner posterior corner; seta VII slender, flexible, triarticulate, pinnate, located midway along inner margin.
</paragraph>
<paragraph id="6903368AFF90FF86FF2EFB60FA20FB3B" blockId="8.[151,1437,150,1581]" box="[199,1434,1159,1185]" pageId="8" pageNumber="537">
Antennule, maxillule, maxilla, and maxilliped as in
<taxonomicName id="AEBC4D09FF90FF86FD17FB60FC00FB3B" authorityName="Vakati, Kihara &amp; Lee" authorityYear="2016" box="[766,954,1159,1185]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="ganghwaensis">
<emphasis id="5BC8EA98FF90FF86FD17FB60FC00FB3B" box="[766,954,1159,1185]" italics="true" pageId="8" pageNumber="537">N. ganghwaensis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0D2D4B7BFF90FF86FC1BFB60FB57FB3B" author="Vakati, V. &amp; Kihara, T. C. &amp; Lee, W." box="[1010,1261,1159,1185]" pageId="8" pageNumber="537" pagination="212 - 233" refId="ref13885" refString="Vakati, V., Kihara, T. C. &amp; Lee, W. (2016) A new species of the genus Nannopus (Copepoda, Harpacticoida, Nannopodidae) from the mudflat of Ganghwa Island, Korea. Proceedings of the biological Society of Washington, 129, 212 - 233. https: // doi. org / 10.2988 / 0006 - 324 X- 129. Q 3.212" type="journal article" year="2016">
Vakati
<emphasis id="5BC8EA98FF90FF86FBD6FB60FBCCFB3B" box="[1087,1142,1159,1185]" italics="true" pageId="8" pageNumber="537">et al.</emphasis>
2016: 218
</bibRefCitation>
,
<figureCitation id="F1872A0FFF90FF86FB11FB60FA34FB3B" box="[1272,1422,1159,1185]" captionStart="FIGURE­4" captionStartId="6.[151,256,1962,1986]" captionTargetBox="[196,1392,193,1926]" captionTargetId="figure-21@6.[182,1405,181,1938]" captionText="FIGURE­4.­Nannopus robustus sp.­nov. (holotype ♂). A, habitus, dorsal; B, habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.5576344" httpUri="https://zenodo.org/record/5576344/files/figure.png" pageId="8" pageNumber="537">Fig. 4A, DF</figureCitation>
).
</paragraph>
<paragraph id="6903368AFF90FF86FF2EFB4CFF67FB73" blockId="8.[151,1437,150,1581]" pageId="8" pageNumber="537">
Antenna, mandible, P1 P6 as in
<taxonomicName id="AEBC4D09FF90FF86FDA4FB4CFD76FB5E" box="[589,716,1195,1220]" class="Hexanauplia" family="Nannopodidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="serratus">
<emphasis id="5BC8EA98FF90FF86FDA4FB4CFD76FB5E" box="[589,716,1195,1220]" italics="true" pageId="8" pageNumber="537">N. serratus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="0D2D4B7BFF90FF86FCE5FB4CFBA2FB5F" author="Vakati, V. &amp; Lee, W." box="[780,1048,1195,1221]" pageId="8" pageNumber="537" pagination="1 - 66" refId="ref13958" refString="Vakati, V. &amp; Lee, W. (2017) Five new species of the genus Nannopus (Copepoda: Harpacticoida: Nannopodidae) from intertidal mudflats of the Korean West Coast (Yellow Sea). Zootaxa, 4360 (1), 1 - 66. https: // doi. org / 10.11646 / zootaxa. 4360.1.1" type="journal article" year="2017">Vakati &amp; Lee 2017: 23</bibRefCitation>
, 25, 30, Figs. 16DF, 18A, E, 22 BD).
</paragraph>
<paragraph id="6903368AFF90FF86FF2EFB14FDF3F993" blockId="8.[151,1437,150,1581]" pageId="8" pageNumber="537">
P2 (
<figureCitation id="F1872A0FFF90FF86FF1BFB14FEDDFA97" box="[242,359,1266,1293]" captionStart="FIGURE­5" captionStartId="7.[151,256,1943,1967]" captionTargetBox="[193,1393,190,1914]" captionTargetId="figure-20@7.[183,1404,181,1919]" captionText="FIGURE­5.­Nannopus robustus sp.­nov. (holotype ♂). A, anal somite and caudal rami, dorsal (cylindrical shape arrowed); B, anal somite and caudal rami, ventral (cylindrical shape arrowed); C, anal somite and caudal ramus, lateral (cylindrical shape arrowed); D, P2 (recurved spine arrowed); E, P2 endopod." figureDoi="http://doi.org/10.5281/zenodo.5576346" httpUri="https://zenodo.org/record/5576346/files/figure.png" pageId="8" pageNumber="537">Fig. 5DE</figureCitation>
) with smooth and short concave intercoxal sclerite (not illustrated). Praecoxa somewhat triangular and shorter than coxa. Coxa with one spinular row on anterior surface. Basis with one row of outer spinules on anterior surface, one row of distal spinules near insertion of exopod, and one row of distal spinules near insertion of endopod. Basal outer seta naked. Exopod 3-segmented, and all segments subequal in length. Each exopodal segment with robust outer spinules and without inner setules. Exp-2 without inner seta. Exp-1 to -2 with serrated outer spines, exp-3 outer spines smooth, and outer distal-most spine heavily robust (arrowed in
<figureCitation id="F1872A0FFF90FF86FB6AFA40FB61FA5B" box="[1155,1243,1446,1473]" captionStart="FIGURE­5" captionStartId="7.[151,256,1943,1967]" captionTargetBox="[193,1393,190,1914]" captionTargetId="figure-20@7.[183,1404,181,1919]" captionText="FIGURE­5.­Nannopus robustus sp.­nov. (holotype ♂). A, anal somite and caudal rami, dorsal (cylindrical shape arrowed); B, anal somite and caudal rami, ventral (cylindrical shape arrowed); C, anal somite and caudal ramus, lateral (cylindrical shape arrowed); D, P2 (recurved spine arrowed); E, P2 endopod." figureDoi="http://doi.org/10.5281/zenodo.5576346" httpUri="https://zenodo.org/record/5576346/files/figure.png" pageId="8" pageNumber="537">Fig. 5D</figureCitation>
), hook-shaped or recurved, and relatively longer. Endopod 2-segmented, both segments subequal in size, with one smooth outer distal spine and one pinnate inner distal seta.
</paragraph>
<paragraph id="6903368AFF90FF86FF2EF9F4FE18F9B7" blockId="8.[151,1437,150,1581]" box="[199,418,1555,1581]" pageId="8" pageNumber="537">
<emphasis id="5BC8EA98FF90FF86FF2EF9F4FE9FF9B7" bold="true" box="[199,293,1555,1581]" pageId="8" pageNumber="537">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>