treatments-xml/data/D2/3C/87/D23C87A0FFB32E17FEB2258CFE75BAE6.xml
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<document id="D81139D8B0D0B322FAD846559DBCB8B6" ID-CLB-Dataset="198285" ID-DOI="10.1080/00222933.2022.2138600" ID-GBIF-Dataset="f3ea9905-e60b-4102-9404-65b3dcd1583c" ID-ISSN="1464-5262" ID-Zenodo-Dep="7380764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669805273948" checkinUser="plazi" docAuthor="Tiwari, Shivam, Padate, Vinay P., Cubelio, Sherine S. &amp; Osawa, Masayuki" docDate="2022" docId="D23C87A0FFB32E17FEB2258CFE75BAE6" docLanguage="en" docName="JNATHIST.56.41-44.1819-1839.pdf" docOrigin="Journal of Natural History 56" docSource="http://dx.doi.org/10.1080/00222933.2022.2138600" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Munidopsis kadal Tiwari &amp; Padate &amp; Cubelio &amp; Osawa 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="1837" masterDocId="2E05FFD8FFBC2E05FF442110FFC2BF11" masterDocTitle="Squat lobsters of the genera Raymunida Macpherson &amp; Machordom, 2000 and Munidopsis Whiteaves, 1874 (Decapoda: Anomura: Galatheoidea) from the Indian Exclusive Economic Zone, with descriptions of three new species" masterLastPageNumber="1839" masterPageNumber="1819" pageNumber="1834" updateTime="1699412247313" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B7D5875EF771C87A21F3FA64876BC986">Squat lobsters of the genera Raymunida Macpherson &amp; Machordom, 2000 and Munidopsis Whiteaves, 1874 (Decapoda: Anomura: Galatheoidea) from the Indian Exclusive Economic Zone, with descriptions of three new species</mods:title>
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<mods:namePart id="53251A29CB61CE460FF8C61712E2358F">Tiwari, Shivam</mods:namePart>
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<mods:namePart id="5A409B5104442ADA8508A9FD1066F028">Padate, Vinay P.</mods:namePart>
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<mods:namePart id="75E9219BE43CCDF217E5A15299E0CCD9">Cubelio, Sherine S.</mods:namePart>
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<mods:namePart id="37CDB56EDD76B19301FF362CC07038CB">Osawa, Masayuki</mods:namePart>
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<mods:date id="B82D7442572BE3EFF46D126F842AD5DD">2022</mods:date>
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<heading id="016281DAFFB32E0AFEB2258CFCECBBA7" box="[502,814,1180,1206]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="1834" reason="5">
<emphasis id="68E1EAA4FFB32E0AFEB2258CFCECBBA7" box="[502,814,1180,1206]" italics="true" pageId="15" pageNumber="1834">
<taxonomicName id="9D954D35FFB32E0AFEB2258CFD0CBBA7" ID-CoL="032d2349-b012-4d94-aae1-7efaed36952b" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[502,718,1180,1206]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="1834" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFB32E0AFEB2258CFD0CBBA7" bold="true" box="[502,718,1180,1206]" italics="true" pageId="15" pageNumber="1834">Munidopsis kadal</emphasis>
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<taxonomicNameLabel id="73D257DFFFB32E0AFD92258CFCECBBA7" box="[726,814,1180,1206]" pageId="15" pageNumber="1834" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="128F653DFFB32E0AFD7325AFFBE3BA1C" pageId="15" pageNumber="1834" type="description">
<paragraph id="5A2A36B6FFB32E0AFD7325AFFD2EBBC8" blockId="15.[502,814,1180,1241]" box="[567,748,1215,1241]" pageId="15" pageNumber="1834">
(
<figureCitation id="C2AE2A33FFB32E0AFD7B25AFFD6EBBC8" box="[575,684,1215,1241]" captionStart="Figure 3" captionStartId="5.[160,225,1345,1368]" captionTargetBox="[163,1152,142,1314]" captionTargetId="figure-50@5.[163,1152,142,1314]" captionTargetPageId="5" captionText="Figure 3. Carapace, pleon and cephalic appendages, dorsal view. (a) Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/00141; (b) R. shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140; (c) R. vittata Macpherson, 2009, ♀ (PCL 9.0 mm, CW 7.7 mm), IO/SS/ANO/00142;(d) Munidopsis bengala sp. nov., holotype ♀ (PCL 8.0 mm, CW 7.7 mm), IO/ SS/ANO/00145; (e) M. kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380770" httpUri="https://zenodo.org/record/7380770/files/figure.png" pageId="15" pageNumber="1834">Figures 3</figureCitation>
(e), 3)
</paragraph>
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urn:lsid:zoobank.org:act:
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<emphasis id="68E1EAA4FFB32E0AFFE4242FFE86BA48" bold="true" box="[160,324,1343,1369]" italics="true" pageId="15" pageNumber="1834">Type material</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFB32E0AFFE42472FD45BAD0" blockId="15.[160,1156,1343,1473]" pageId="15" pageNumber="1834">
<materialsCitation id="EAFD3CEBFFB32E0AFFE42472FD45BAD0" ID-GBIF-Occurrence="3978782305" collectingDate="2015-06-17" collectionCode="CMLRE, IO" collectorName="N. Saravanane" country="India" elevation="1018" latitude="10.99" location="Arabian Sea" longLatPrecision="778" longitude="74.98" pageId="15" pageNumber="1834" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="holotype">
<emphasis id="68E1EAA4FFB32E0AFFE42472FED0BA6D" bold="true" box="[160,274,1378,1404]" italics="true" pageId="15" pageNumber="1834">
<typeStatus id="852E8814FFB32E0AFFE42472FECFBA6D" box="[160,269,1378,1404]" pageId="15" pageNumber="1834" type="holotype">Holotype</typeStatus>
.
</emphasis>
<emphasis id="68E1EAA4FFB32E0AFE652473FE9CBA6D" bold="true" box="[289,350,1379,1404]" pageId="15" pageNumber="1834">
<collectingCountry id="22827626FFB32E0AFE652473FE9CBA6D" box="[289,350,1379,1404]" name="India" pageId="15" pageNumber="1834">India</collectingCountry>
</emphasis>
<specimenCount id="4C93FD3FFFB32E0AFE3D2473FE4FBA6B" box="[377,397,1379,1402]" pageId="15" pageNumber="1834" type="male"></specimenCount>
(PCL
<quantity id="9D6D9B53FFB32E0AFE962473FDEEBA6D" box="[466,556,1379,1404]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="15" pageNumber="1834" unit="mm" value="5.2">5.2 mm</quantity>
, CW 5.0 mm); south-eastern
<location id="5F4A606DFFB32E0AFC3E2473FBC7BA6D" LSID="urn:lsid:plazi:treatment:D23C87A0FFB32E17FEB2258CFE75BAE6:5F4A606DFFB32E0AFC3E2473FBC7BA6D" box="[890,1029,1379,1404]" country="India" latitude="10.99" longLatPrecision="778" longitude="74.98" name="Arabian Sea" pageId="15" pageNumber="1834" stateProvince="Kerala">Arabian Sea</location>
, off
<collectingRegion id="9851F854FFB32E0AFB7D2473FB42BA6D" box="[1081,1152,1379,1404]" country="India" name="Kerala" pageId="15" pageNumber="1834">Kerala</collectingRegion>
; FORVSS stn. 341I06;
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,
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;
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<quantity id="9D6D9B53FFB32E0AFD152495FD65BA8F" box="[593,679,1413,1438]" metricMagnitude="3" metricUnit="m" metricValue="1.018" pageId="15" pageNumber="1834" unit="m" value="1018.0">1018 m</quantity>
depth
</elevation>
; naturalists dredge;
<date id="2E2B1076FFB32E0AFCA02495FB42BA8F" box="[996,1152,1413,1438]" pageId="15" pageNumber="1834" value="2015-06-17">
<collectingDate id="3E6FE99EFFB32E0AFCA02495FB42BA8F" box="[996,1152,1413,1438]" pageId="15" pageNumber="1834" value="2015-06-17">17 June 2015</collectingDate>
</date>
;
<collectorName id="F7605360FFB32E0AFFE424B8FEFDBAD0" box="[160,319,1448,1473]" pageId="15" pageNumber="1834">N. Saravanane</collectorName>
leg.;
<collectionCode id="3C84AE73FFB32E0AFE3824BDFE70BAAF" box="[380,434,1453,1470]" pageId="15" pageNumber="1834">CMLRE</collectionCode>
<collectionCode id="3C84AE73FFB32E0AFEFD24B8FE13BAD0" box="[441,465,1448,1473]" country="Portugal" name="Instituto de Oceanografia da Universidade de Lisboa" pageId="15" pageNumber="1834">IO</collectionCode>
/SS/ANO/00146.
</materialsCitation>
</paragraph>
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<emphasis id="68E1EAA4FFB32E0AFFE424E3FEF4B91C" bold="true" box="[160,310,1523,1549]" italics="true" pageId="15" pageNumber="1834">Type locality</emphasis>
</paragraph>
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<materialsCitation id="EAFD3CEBFFB32E0AFFE42706FC34B93E" ID-GBIF-Occurrence="3978782310" box="[160,1014,1558,1583]" country="India" elevation="1018" latitude="10.99" location="South-eastern Arabian Sea" longLatPrecision="778" longitude="74.98" pageId="15" pageNumber="1834" specimenCount="1" stateProvince="Kerala">
<collectingCountry id="22827626FFB32E0AFFE42706FF18B93E" box="[160,218,1558,1583]" name="India" pageId="15" pageNumber="1834">India</collectingCountry>
:
<location id="5F4A606DFFB32E0AFFA02706FDD3B93E" LSID="urn:lsid:plazi:treatment:D23C87A0FFB32E17FEB2258CFE75BAE6:5F4A606DFFB32E0AFFA02706FDD3B93E" box="[228,529,1558,1583]" country="India" latitude="10.99" longLatPrecision="778" longitude="74.98" name="South-eastern Arabian Sea" pageId="15" pageNumber="1834" stateProvince="Kerala">South-eastern Arabian Sea</location>
, off
<collectingRegion id="9851F854FFB32E0AFD052706FD4AB93E" box="[577,648,1558,1583]" country="India" name="Kerala" pageId="15" pageNumber="1834">Kerala</collectingRegion>
;
<geoCoordinate id="3FA15071FFB32E0AFDD72706FD2EB93E" box="[659,748,1558,1583]" degrees="10.99" direction="north" orientation="latitude" pageId="15" pageNumber="1834" precision="555" value="10.99">10.99°N</geoCoordinate>
,
<geoCoordinate id="3FA15071FFB32E0AFDB22706FC89B93E" box="[758,843,1558,1583]" degrees="74.98" direction="east" orientation="longitude" pageId="15" pageNumber="1834" precision="555" value="74.98">74.98°E</geoCoordinate>
;
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<quantity id="9D6D9B53FFB32E0AFC112706FC6BB93E" box="[853,937,1558,1583]" metricMagnitude="3" metricUnit="m" metricValue="1.018" pageId="15" pageNumber="1834" unit="m" value="1018.0">1018 m</quantity>
depth
</elevation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="128F653DFFB32E0AFFE42771FEF8B9D1" pageId="15" pageNumber="1834" type="etymology">
<paragraph id="5A2A36B6FFB32E0AFFE42771FEDFB96A" blockId="15.[160,1155,1633,1728]" box="[160,285,1633,1659]" pageId="15" pageNumber="1834">
<emphasis id="68E1EAA4FFB32E0AFFE42771FEDFB96A" bold="true" box="[160,285,1633,1659]" italics="true" pageId="15" pageNumber="1834">Etymology</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFB32E0AFFE42795FEF8B9D1" blockId="15.[160,1155,1633,1728]" pageId="15" pageNumber="1834">The species name is derived from the Malayalam word for sea. The name is used as noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="128F653DFFAC2E15FFE4219EFC57BD9C" pageId="16" pageNumber="1835" type="diagnosis">
<paragraph id="5A2A36B6FFAC2E15FFE4219EFED6BFB9" blockId="16.[159,1156,142,653]" box="[160,276,142,168]" pageId="16" pageNumber="1835">
<emphasis id="68E1EAA4FFAC2E15FFE4219EFED6BFB9" bold="true" box="[160,276,142,168]" italics="true" pageId="16" pageNumber="1835">Diagnosis</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFAC2E15FFE421A2FC57BD9C" blockId="16.[159,1156,142,653]" pageId="16" pageNumber="1835">
<taxonomicName id="9D954D35FFAC2E15FFE421A2FEE6BFDA" box="[160,292,178,203]" class="Malacostraca" family="Munidopsidae" genus="Anoplonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1835" phylum="Arthropoda" rank="genus">
<emphasis id="68E1EAA4FFAC2E15FFE421A2FEE6BFDA" box="[160,292,178,203]" italics="true" pageId="16" pageNumber="1835">Anoplonotus</emphasis>
</taxonomicName>
group. Carapace dorsally uneven,covered with numerous granules and low small tubercles; epigastric bulges blunt, each bearing tuberculate ridge anteriorly; posterior cardiac region widely triangular; lateral margins subparallel, unarmed, each with rounded anterolateral angle. Rostrum narrow subtriangular, nearly horizontal, with mid-dorsal granular carina extending between epigastric bulges; width at base about 0.4 of distance between anterolateral angles of carapace. Thoracic sternite 4 broad, 2.6 times wider than, and not contiguous to, sternite 3; anterolateral margins gently concave. Pleonal tergites unarmed, tergites 24 each with elevated transverse ridge anteriorly, tergites 2 and 3 with posterior transverse ridge. No eye spines. Antennular article 1 with 2 strong distal spines, ventrodistal spine larger than dorsodistal spine. Antennal peduncle overreaching distal corneal margin by half length of article 4, article 1 with small subacute ventromesial projection. Mxp3 with 2 ventral spines, dorsal margin distally with distinct spine. P1 slender, 3.33.4 times PCL, unarmed, with numerous coarse granules; fingers 0.9 length of palm, not gaping when closed, dactylus with low tooth proximally on cutting edge. Epipods absent from P14.
</paragraph>
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<subSubSection id="128F653DFFAC2E17FFE423D9FD4DBFB6" lastPageId="18" lastPageNumber="1837" pageId="16" pageNumber="1835" type="description">
<paragraph id="5A2A36B6FFAC2E15FFE423D9FE74BDF2" blockId="16.[160,1157,713,1709]" box="[160,438,713,739]" pageId="16" pageNumber="1835">
<emphasis id="68E1EAA4FFAC2E15FFE423D9FE74BDF2" bold="true" box="[160,438,713,739]" italics="true" pageId="16" pageNumber="1835">
Description of
<typeStatus id="852E8814FFAC2E15FE0A23D9FE74BDF2" box="[334,438,713,739]" pageId="16" pageNumber="1835" type="holotype">holotype</typeStatus>
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</paragraph>
<paragraph id="5A2A36B6FFAC2E15FFE423FCFBCCBA1C" blockId="16.[160,1157,713,1709]" pageId="16" pageNumber="1835">
Carapace (
<figureCitation id="C2AE2A33FFAC2E15FE5223FCFEBFBC14" box="[278,381,748,773]" captionStart="Figure 3" captionStartId="5.[160,225,1345,1368]" captionTargetBox="[163,1152,142,1314]" captionTargetId="figure-50@5.[163,1152,142,1314]" captionTargetPageId="5" captionText="Figure 3. Carapace, pleon and cephalic appendages, dorsal view. (a) Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/00141; (b) R. shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140; (c) R. vittata Macpherson, 2009, ♀ (PCL 9.0 mm, CW 7.7 mm), IO/SS/ANO/00142;(d) Munidopsis bengala sp. nov., holotype ♀ (PCL 8.0 mm, CW 7.7 mm), IO/ SS/ANO/00145; (e) M. kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380770" httpUri="https://zenodo.org/record/7380770/files/figure.png" pageId="16" pageNumber="1835">Figures 3</figureCitation>
(e), 3(a, 3)) (exclusive of rostrum) quadrangular in general outline, 1.1 times as long as wide; lateral margins subparallel, each with small notch at lateral end of anterior cervical groove. Dorsal surface with regions well delineated by deep depressions, uneven in lateral view, unarmed, covered with granules and low small tubercles. Hepatic region not elevated. Gastric region strongly convex, with pair of blunt epigastric bulges, each bulge delimited anteriorly by tuberculate ridge. Cervical grooves deep. Cardiac region subdivided into anterior and posterior parts by deep transverse groove; posterior part widely triangular, more elevated than gastric region. Posterior branchial region subdivided into 2 parts by deep transverse groove. Frontal margin behind ocular peduncles oblique, gently convex between ocular and antennal peduncles; antennal spine absent; no submarginal protuberance between ocular and antennal peduncle. Anterolateral angle rounded. Posterior submarginal ridge elevated, sharp, unarmed. Rostrum (
<figureCitation id="C2AE2A33FFAC2E15FE4B259CFEACBBB4" box="[271,366,1164,1189]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="16" pageNumber="1835">Figure 8</figureCitation>
(a,)) about 0.3 times PCL, narrow, distinctly tapering distally in dorsal view, terminating in blunt tip, nearly horizontal; dorsal surface with median granular carina extending between epigastric bulges; surfaces lateral to carina minutely granulate; width at base about 0.4 of distance between anterolateral angles of carapace.
</paragraph>
<paragraph id="5A2A36B6FFAC2E15FF842407FE50BA42" blockId="16.[160,1157,713,1709]" pageId="16" pageNumber="1835">
Pterygostomian flap (
<figureCitation id="C2AE2A33FFAC2E15FEF42407FDCEBA21" box="[432,524,1303,1328]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="16" pageNumber="1835">Figure 8</figureCitation>
(b)) covered with short rows of granules; anterior margin bluntly subtriangular.
</paragraph>
<paragraph id="5A2A36B6FFAC2E15FF84244CFCD4B932" blockId="16.[160,1157,713,1709]" pageId="16" pageNumber="1835">
Sternum (
<figureCitation id="C2AE2A33FFAC2E15FE69244CFE4ABA64" box="[301,392,1372,1397]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="16" pageNumber="1835">Figure 8</figureCitation>
(c)) 1.2 times as wide as long, maximum width at sternite 7. Sternite 3 divided into 2 ovate lobes with distinctly sinuous anterior and posterior margins. Sternite 4 2.6 times wider than, and not contiguous to, sternite 3; surface smooth except for scattered granules, with subcircular median depression behind posterior margin of sternite 3; anterolateral margins gently concave, each with granulated ridge. Sternites 57 nearly smooth, each side with elevated anterior margin.
</paragraph>
<paragraph id="5A2A36B6FFAC2E15FF84273CFBB1B9BC" blockId="16.[160,1157,713,1709]" pageId="16" pageNumber="1835">
Pleon (
<figureCitation id="C2AE2A33FFAC2E15FE55273CFEB3B954" box="[273,369,1580,1605]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="16" pageNumber="1835">Figure 8</figureCitation>
(a,,)) unarmed, surfaces of tergites and pleura smooth. Tergites 24 each with elevated transverse ridge anteriorly, tergites 2 and 3 each with posterior transverse ridge. Tergites 5 and 6 without transverse ridges, distomedian margin of tergite 6 flattish. Telson (
<figureCitation id="C2AE2A33FFAC2E15FE222784FE00B9BC" box="[358,450,1684,1709]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="16" pageNumber="1835">Figure 8</figureCitation>
(e)) 1.3 times as wide as long, composed of 8 calcified plates.
</paragraph>
<caption id="0EEA663EFFAD2E14FFE422E3FC17BB6E" ID-DOI="http://doi.org/10.5281/zenodo.7380786" ID-Zenodo-Dep="7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="17" pageNumber="1836" startId="17.[160,225,1011,1034]" targetBox="[166,1149,145,975]" targetPageId="17">
<paragraph id="5A2A36B6FFAD2E14FFE422E3FC17BB6E" blockId="17.[160,1156,1011,1151]" pageId="17" pageNumber="1836">
<emphasis id="68E1EAA4FFAD2E14FFE422E3FF3EBB1B" bold="true" box="[160,252,1011,1034]" pageId="17" pageNumber="1836">Figure 8.</emphasis>
<taxonomicName id="9D954D35FFAD2E14FE4222E3FE72BB1B" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[262,432,1011,1034]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="1836" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAD2E14FE4222E3FE72BB1B" box="[262,432,1011,1034]" italics="true" pageId="17" pageNumber="1836">Munidopsis kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAD2E14FEFC22E3FDC8BB1B" box="[440,522,1011,1034]" pageId="17" pageNumber="1836" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm.
</paragraph>
</caption>
<paragraph id="5A2A36B6FFAD2E14FF8425A9FEDFBA09" blockId="17.[159,1156,1209,1720]" pageId="17" pageNumber="1836">
Ocular peduncle (
<figureCitation id="C2AE2A33FFAD2E14FEC225A9FE23BBC3" box="[390,481,1209,1234]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="17" pageNumber="1836">Figure 8</figureCitation>
(a)) short, stout, movable, unarmed. Cornea subglobular, not dilated, weakly pigmented, slightly longer than remaining peduncle, reaching 0.4 length of rostrum.
</paragraph>
<paragraph id="5A2A36B6FFAD2E14FF842431FD7BBA4C" blockId="17.[159,1156,1209,1720]" pageId="17" pageNumber="1836">
Antennular article 1 (
<figureCitation id="C2AE2A33FFAD2E14FEF62431FDD3BA2B" box="[434,529,1313,1338]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="17" pageNumber="1836">Figure 8</figureCitation>
(f)) distally armed with 2 strong, well-separated spines, ventrodistal spine larger than dorsodistal spine.
</paragraph>
<paragraph id="5A2A36B6FFAD2E14FF842477FB8ABAB3" blockId="17.[159,1156,1209,1720]" pageId="17" pageNumber="1836">
Antennal peduncle (
<figureCitation id="C2AE2A33FFAD2E14FEEC2477FDC5BA91" box="[424,519,1383,1408]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="17" pageNumber="1836">Figure 8</figureCitation>
(f)) overreaching distal corneal margin by half length of article 4; article 1 with small subacute distomesial projection; articles 24 unarmed.
</paragraph>
<paragraph id="5A2A36B6FFAD2E14FF8424BCFB46B91B" blockId="17.[159,1156,1209,1720]" pageId="17" pageNumber="1836">
Mxp3 (
<figureCitation id="C2AE2A33FFAD2E14FE4A24BCFEA8BAD4" box="[270,362,1452,1477]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="17" pageNumber="1836">Figure 8</figureCitation>
(g)) ischium 0.8 merus length; dorsal margin sharply ridged, with small distolateral spine, crista dentata consisting of 19 corneous denticles. Merus with 2 spines on ventral margin,distal spine smaller than proximal spine;dorsal margin with strong distal spine.
</paragraph>
<paragraph id="5A2A36B6FFAD2E14FF842704FCB0B9A9" blockId="17.[159,1156,1209,1720]" pageId="17" pageNumber="1836">
P1 subequal (
<figureCitation id="C2AE2A33FFAD2E14FE1C2704FE77B93C" box="[344,437,1556,1581]" captionStart="Figure 8" captionStartId="17.[160,225,1011,1034]" captionTargetBox="[166,1149,145,975]" captionTargetId="figure-180@17.[163,1152,142,979]" captionTargetPageId="17" captionText="Figure 8. Munidopsis kadal sp. nov., holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. (a) carapace, pleon and cephalic appendages (dorsal view); (b) carapace (right lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (dorsal view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) right P1, dorsal view. Scale bars: ae, h = 1.0 mm; f, g = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380786" httpUri="https://zenodo.org/record/7380786/files/figure.png" pageId="17" pageNumber="1836">Figure 8</figureCitation>
(h)), 3.33.4 times PCL, unarmed, covered with numerous coarse granules. Merus somewhat concave on mesial surface; lateral surface with short subdistal ridge. Carpus and chela slightly compressed dorsoventrally. Palm 1.8 times longer than carpus. Fingers 0.9 length of palm, not gaping when closed; cutting edges each with row of small blunt teeth, also with low tooth proximally on dactylus.
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF84219EFD4DBFB6" blockId="18.[192,655,142,167]" box="[192,655,142,167]" pageId="18" pageNumber="1837">P24 missing. Epipods absent from P14.</paragraph>
</subSubSection>
<subSubSection id="128F653DFFAE2E17FFE421C7FE75BAE6" pageId="18" pageNumber="1837" type="discussion">
<paragraph id="5A2A36B6FFAE2E17FFE421C7FEC4BFE0" blockId="18.[160,1156,215,899]" box="[160,262,215,241]" pageId="18" pageNumber="1837">
<emphasis id="68E1EAA4FFAE2E17FFE421C7FEC4BFE0" bold="true" box="[160,262,215,241]" italics="true" pageId="18" pageNumber="1837">Remarks</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FFE421EAFBDDBE8F" blockId="18.[160,1156,215,899]" pageId="18" pageNumber="1837">
As discussed above in the Remarks section for
<taxonomicName id="9D954D35FFAE2E17FDF821EAFCF5BE02" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[700,823,250,275]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FDF821EAFCF5BE02" box="[700,823,250,275]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FC0421EAFC59BE02" box="[832,923,250,275]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="9D954D35FFAE2E17FCE321EAFBC5BE02" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[935,1031,250,275]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FCE321EAFBC5BE02" box="[935,1031,250,275]" italics="true" pageId="18" pageNumber="1837">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FB5521EAFBABBE02" box="[1041,1129,250,275]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
is morphologically similar to
<taxonomicName id="9D954D35FFAE2E17FE95200DFD8FBE27" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[465,589,285,310]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FE95200DFD8FBE27" box="[465,589,285,310]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FD13200DFD71BE27" box="[599,691,285,310]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="68E1EAA4FFAE2E17FD84200DFC6FBE27" box="[704,941,285,310]" italics="true" pageId="18" pageNumber="1837">
<taxonomicName id="9D954D35FFAE2E17FD84200DFCE2BE27" authorityName="Macpherson" authorityYear="2007" box="[704,800,285,310]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bruta">M. bruta</taxonomicName>
,
<taxonomicName id="9D954D35FFAE2E17FC6B200DFC6FBE27" authorityName="Vazquez-Bader, Gracia and Lemaitre" authorityYear="2014" box="[815,941,285,310]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="shulerae">M. shulerae</taxonomicName>
</emphasis>
and
<taxonomicName id="9D954D35FFAE2E17FCA8200DFBBFBE27" authorityName="Macpherson and Segonzac" authorityYear="2005" box="[1004,1149,285,310]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="truculenta">
<emphasis id="68E1EAA4FFAE2E17FCA8200DFBBFBE27" box="[1004,1149,285,310]" italics="true" pageId="18" pageNumber="1837">M. truculenta</emphasis>
</taxonomicName>
. However,
<taxonomicName id="9D954D35FFAE2E17FE48202FFEA5BE49" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[268,359,319,344]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FE48202FFEA5BE49" box="[268,359,319,344]" italics="true" pageId="18" pageNumber="1837">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FE28202FFE7DBE49" box="[364,447,319,344]" pageId="18" pageNumber="1837" rank="species">sp.nov.</taxonomicNameLabel>
is distinguished from the other four species by the proportionally shorter antennal peduncle which overreaches the distal corneal margin by half the length of article 4, rather than by at least the full length of article 4 as in the other species.
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF8420B7FE12BD7F" blockId="18.[160,1156,215,899]" pageId="18" pageNumber="1837">
<taxonomicName id="9D954D35FFAE2E17FF8420B7FE44BED1" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[192,390,423,448]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FF8420B7FE44BED1" box="[192,390,423,448]" italics="true" pageId="18" pageNumber="1837">Munidopsis kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FED520B7FE2FBED1" box="[401,493,423,448]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
is morphologically closest to
<taxonomicName id="9D954D35FFAE2E17FC1220B7FC1ABED1" authorityName="Vazquez-Bader, Gracia and Lemaitre" authorityYear="2014" box="[854,984,423,448]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="shulerae">
<emphasis id="68E1EAA4FFAE2E17FC1220B7FC1ABED1" box="[854,984,423,448]" italics="true" pageId="18" pageNumber="1837">M. shulerae</emphasis>
</taxonomicName>
in having the combination of the following characters: the carapace lateral margins are subparallel; the epigastric bulges each have a tuberculate ridge anteriorly; and the rostrum has a middorsal carina. However, the new species differs from
<taxonomicName id="9D954D35FFAE2E17FC44231FFCBCBD39" authorityName="Vazquez-Bader, Gracia and Lemaitre" authorityYear="2014" box="[768,894,527,552]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="shulerae">
<emphasis id="68E1EAA4FFAE2E17FC44231FFCBCBD39" box="[768,894,527,552]" italics="true" pageId="18" pageNumber="1837">M. shulerae</emphasis>
</taxonomicName>
in lacking dense short setae along the cutting edges of the P1 finger, in addition to having a proportionally shorter antennal peduncle.
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF842367FE72BC2F" blockId="18.[160,1156,215,899]" pageId="18" pageNumber="1837">
The new species further differs from
<taxonomicName id="9D954D35FFAE2E17FD1A2367FD79BD81" authorityName="Macpherson" authorityYear="2007" box="[606,699,631,656]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bruta">
<emphasis id="68E1EAA4FFAE2E17FD1A2367FD79BD81" box="[606,699,631,656]" italics="true" pageId="18" pageNumber="1837">M. bruta</emphasis>
</taxonomicName>
in the dorsal margin of the Mxp3 merus being unarmed exclusive of a strong distal spine, instead of being crenulated or having one or two small spines; and from
<taxonomicName id="9D954D35FFAE2E17FD7C23ADFD09BDC7" authorityName="Macpherson and Segonzac" authorityYear="2005" box="[568,715,701,726]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="truculenta">
<emphasis id="68E1EAA4FFAE2E17FD7C23ADFD09BDC7" box="[568,715,701,726]" italics="true" pageId="18" pageNumber="1837">M. truculenta</emphasis>
</taxonomicName>
in the lateral carapace margins being subparallel, rather than divergent posteriorly, and the dorsal surface of the carapace with coarse granules and small tubercles, rather than being distinctly tuberculate, particularly on the branchial region.
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF842258FEDBBC92" blockId="18.[160,1156,215,899]" pageId="18" pageNumber="1837">
<taxonomicName id="9D954D35FFAE2E17FF842258FEBCBC70" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[192,382,840,865]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FF842258FEBCBC70" box="[192,382,840,865]" italics="true" pageId="18" pageNumber="1837">Munidopsis kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FEC12258FE1EBC70" box="[389,476,840,865]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="9D954D35FFAE2E17FD572258FD4FBC70" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[531,653,840,865]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FD572258FD4FBC70" box="[531,653,840,865]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FDD02258FD28BC70" box="[660,746,840,865]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
are distinguishable by the following characters.
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF8422A0FD1EBCFA" blockId="18.[192,1156,944,1384]" pageId="18" pageNumber="1837">
(1) The dorsal surface of the carapace is more strongly uneven and more tuberculate in
<taxonomicName id="9D954D35FFAE2E17FFAD22C2FE85BCFA" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[233,327,978,1003]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FFAD22C2FE85BCFA" box="[233,327,978,1003]" italics="true" pageId="18" pageNumber="1837">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FE0A22C2FE64BCFA" box="[334,422,978,1003]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
than in
<taxonomicName id="9D954D35FFAE2E17FD4022C2FDBFBCFA" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[516,637,978,1003]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FD4022C2FDBFBCFA" box="[516,637,978,1003]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FDC122C2FD1EBCFA" box="[645,732,978,1003]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF8422E5FB46BB20" blockId="18.[192,1156,944,1384]" pageId="18" pageNumber="1837">
(2) The lateral margins of the carapace are subparallel in
<taxonomicName id="9D954D35FFAE2E17FC7D22E5FC57BB1F" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[825,917,1013,1038]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FC7D22E5FC57BB1F" box="[825,917,1013,1038]" italics="true" pageId="18" pageNumber="1837">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FCDF22E5FC36BB1F" box="[923,1012,1013,1038]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
, but they are gently divergent posteriorly along the anterior three-fourths in
<taxonomicName id="9D954D35FFAE2E17FCEA2508FBE5BB20" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[942,1063,1048,1073]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FCEA2508FBE5BB20" box="[942,1063,1048,1073]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FB692508FB46BB20" box="[1069,1156,1048,1073]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF84252AFB82BB88" blockId="18.[192,1156,944,1384]" pageId="18" pageNumber="1837">
(3) The pair of epigastric bulges on the carapace are each delimited anteriorly by a tuberculate ridge in
<taxonomicName id="9D954D35FFAE2E17FEBF254DFD9DBB67" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[507,607,1117,1142]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FEBF254DFD9DBB67" box="[507,607,1117,1142]" italics="true" pageId="18" pageNumber="1837">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FD2F254DFD09BB67" box="[619,715,1117,1142]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
, whereas the anterior margin of the epigastric bulges is rounded and such ridge is absent in
<taxonomicName id="9D954D35FFAE2E17FC232590FC23BB88" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[871,993,1152,1177]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FC232590FC23BB88" box="[871,993,1152,1177]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FCAC2590FB82BB88" box="[1000,1088,1152,1177]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF8425B2FD98BA32" blockId="18.[192,1156,944,1384]" pageId="18" pageNumber="1837">
(4) The rostrum is wider at the base and tapers more strongly in
<taxonomicName id="9D954D35FFAE2E17FCD525B2FC2CBBAA" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[913,1006,1186,1211]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FCD525B2FC2CBBAA" box="[913,1006,1186,1211]" italics="true" pageId="18" pageNumber="1837">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FCB025B2FB88BBAA" box="[1012,1098,1186,1211]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
than in
<taxonomicName id="9D954D35FFAE2E17FE4225D5FE43BBCF" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[262,385,1221,1246]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FE4225D5FE43BBCF" box="[262,385,1221,1246]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FECE25D5FE21BBCF" box="[394,483,1221,1246]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
The basal width of the rostrum is about 0.4 and 0.3 of the distance between the anterolateral angles of the carapace in
<taxonomicName id="9D954D35FFAE2E17FCD325F7FC37BA11" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[919,1013,1255,1280]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FCD325F7FC37BA11" box="[919,1013,1255,1280]" italics="true" pageId="18" pageNumber="1837">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FCB825F7FB90BA11" box="[1020,1106,1255,1280]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="9D954D35FFAE2E17FFAD241AFEA1BA32" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[233,355,1290,1315]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FFAD241AFEA1BA32" box="[233,355,1290,1315]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FE2E241AFE06BA32" box="[362,452,1290,1315]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
, respectively.
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FF84243DFDF3BA79" blockId="18.[192,1156,944,1384]" pageId="18" pageNumber="1837">
(5) The Mxp3 merus has a strong dorsodistal spine in
<taxonomicName id="9D954D35FFAE2E17FC74243DFC4DBA57" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[816,911,1325,1350]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FC74243DFC4DBA57" box="[816,911,1325,1350]" italics="true" pageId="18" pageNumber="1837">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FCDD243DFC36BA57" box="[921,1012,1325,1350]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
; the spine is absent in
<taxonomicName id="9D954D35FFAE2E17FE1C245FFE10BA79" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[344,466,1359,1384]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="1837" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFAE2E17FE1C245FFE10BA79" box="[344,466,1359,1384]" italics="true" pageId="18" pageNumber="1837">M. bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFAE2E17FE9E245FFDF3BA79" box="[474,561,1359,1384]" pageId="18" pageNumber="1837" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FFE42488FE12BAA3" blockId="18.[160,1156,1432,1527]" box="[160,464,1432,1458]" pageId="18" pageNumber="1837">
<emphasis id="68E1EAA4FFAE2E17FFE42488FE12BAA3" bold="true" box="[160,464,1432,1458]" italics="true" pageId="18" pageNumber="1837">Geographical distribution</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFAE2E17FFE424ABFE75BAE6" blockId="18.[160,1156,1432,1527]" pageId="18" pageNumber="1837">
<materialsCitation id="EAFD3CEBFFAE2E17FFE424ABFE71BAE6" ID-GBIF-Occurrence="3978782307" country="India" elevation="1018" location="Arabian Sea" pageId="18" pageNumber="1837" specimenCount="1" stateProvince="Kerala">
So far known only from the
<typeStatus id="852E8814FFAE2E17FEAB24ABFDE3BAC5" box="[495,545,1467,1492]" pageId="18" pageNumber="1837">type</typeStatus>
locality, south-eastern
<location id="5F4A606DFFAE2E17FC7424ABFC7EBAC5" LSID="urn:lsid:plazi:treatment:D23C87A0FFB32E17FEB2258CFE75BAE6:5F4A606DFFAE2E17FC7424ABFC7EBAC5" box="[816,956,1467,1492]" country="India" name="Arabian Sea" pageId="18" pageNumber="1837" stateProvince="Kerala">Arabian Sea</location>
, off
<collectingRegion id="9851F854FFAE2E17FCB624ABFBFBBAC5" box="[1010,1081,1467,1492]" country="India" name="Kerala" pageId="18" pageNumber="1837">Kerala</collectingRegion>
,
<collectingCountry id="22827626FFAE2E17FB0224ABFB42BAC5" box="[1094,1152,1467,1492]" name="India" pageId="18" pageNumber="1837">India</collectingCountry>
,
<elevation id="D1B8D185FFAE2E17FFE424CEFEFDBAE6" box="[160,319,1502,1527]" metricMagnitude="3" metricUnit="m" metricValue="1.018" pageId="18" pageNumber="1837" unit="m" value="1018.0">
<quantity id="9D6D9B53FFAE2E17FFE424CEFF36BAE6" box="[160,244,1502,1527]" metricMagnitude="3" metricUnit="m" metricValue="1.018" pageId="18" pageNumber="1837" unit="m" value="1018.0">1018 m</quantity>
depth
</elevation>
(
<figureCitation id="C2AE2A33FFAE2E17FE0A24CEFE69BAE6" box="[334,427,1502,1527]" captionStart="Figure 1" captionStartId="2.[160,225,819,842]" captionTargetBox="[164,1152,143,786]" captionTargetId="figure-273@2.[163,1153,142,787]" captionTargetPageId="2" captionText="Figure 1. Map of India indicating collection locations for the material examined in the present study. 1, Raymunida shraddhanandi sp. nov. and R. vittata Macpherson, 2009; 2, Munidopsis bengala sp. nov.; 3, Munidopsis kadal sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7380766" httpUri="https://zenodo.org/record/7380766/files/figure.png" pageId="18" pageNumber="1837">Figure 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>