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<mods:title id="7BEA43F022681F599BEA2D3CF1967650">The millipede genus Julus Linnaeus, 1758 in the Caucasus (Diplopoda: Julida: Julidae)</mods:title>
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<emphasis id="DBD3EAFF9315FF99FF0DF190FEAD18CF" bold="true" box="[151,339,1738,1764]" italics="true" pageId="3" pageNumber="92">Julus colchicus</emphasis>
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<figureCitation id="719C2A689315FF99FF0DF1B7FF0E192D" box="[151,240,1773,1798]" captionStart="FIGURE 1" captionStartId="4.[151,249,1820,1842]" captionTargetBox="[344,1236,854,1775]" captionTargetId="figure@4.[340,1248,841,1798]" captionTargetPageId="4" captionText="FIGURE 1. Antennae in Caucasian species of Julus, lateral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. dagestanus sp. n. H: J. khostensis sp. n. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459874/files/figure.png" pageId="3" pageNumber="92">Figs 1B</figureCitation>
,
<figureCitation id="719C2A689315FF99FF67F1B4FEDE192D" box="[253,288,1774,1798]" captionStart="FIGURE 2" captionStartId="5.[151,249,1601,1623]" captionTargetBox="[340,1248,985,1580]" captionTargetId="figure@5.[340,1248,985,1580]" captionTargetPageId="5" captionText="FIGURE 2. Gnathochilaria in Caucasian species of Julus, ventral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="3" pageNumber="92">2B</figureCitation>
,
<figureCitation id="719C2A689315FF99FEB6F1B4FE8F192D" box="[300,369,1774,1798]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF), J. kubanus Verhoeff, 1921 (GI), J. lignaui Verhoeff, 1910 (JM), J. lindholmi Lohmander, 1936 (N P), J. subalpinus Lohmander, 1936 (QR), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views BO; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (AC, E, G, H, JO, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="3" pageNumber="92">3BD</figureCitation>
,
<figureCitation id="719C2A689315FF99FEE6F1B4FE3C192D" box="[380,450,1774,1798]" captionStart="FIGURE 4" captionStartId="8.[151,250,1477,1499]" captionTargetBox="[340,1248,375,1442]" captionTargetId="figure@8.[340,1248,375,1455]" captionTargetPageId="8" captionText="FIGURE 4. Male legs 2 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF) and J. kubanus Verhoeff, 1921 (GI). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). Oral views C, E, H; caudal views A, B, D, F, G; mesal view I. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (AC, E, G, H), 0.2 mm (F), not to scale (D, I)." httpUri="https://zenodo.org/record/1459880/files/figure.png" pageId="3" pageNumber="92">4BD</figureCitation>
,
<figureCitation id="719C2A689315FF99FE57F1B7FE23192D" box="[461,477,1773,1798]" captionStart="FIGURE 5" captionStartId="9.[151,250,1527,1549]" captionTargetBox="[340,1247,1093,1506]" captionTargetId="figure@9.[340,1247,1093,1506]" captionTargetPageId="9" captionText="FIGURE 5. Julus colchicus Lohmander, 1936 from Krasnaya Polyana, Krasnodar Province. A: habitus of female (left) and male (right), lateral views. B and C: anterior part of male body, lateral and ventral views, respectively. D: midbody segments, lateral view. E: caudal part of body, lateral view. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459882/files/figure.png" pageId="3" pageNumber="92">5</figureCitation>
,
<figureCitation id="719C2A689315FF99FE73F1B7FDDD192D" box="[489,547,1773,1798]" captionStart="FIGURE 6" captionStartId="10.[151,250,1294,1316]" captionTargetBox="[340,1248,199,1261]" captionTargetId="figure@10.[340,1248,193,1273]" captionTargetPageId="10" captionText="FIGURE 6. Male legs 2 in Julus lignaui Verhoeff, 1910 (AD) and J. lindholmi Lohmander, 1936 (EG), caudal views. A: from Krasnaya Polyana, Krasnodar Province. B: from Teberda, Karachaevo-Cherkessia. C: from Mestia, Georgia. D: from Teberda, Karachaevo-Cherkessia. E: from Svanetia, Georgia. F: from near Teberda, Karachaevo-Cherkessia. G: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (AF), not to scale (G)." httpUri="https://zenodo.org/record/1459884/files/figure.png" pageId="3" pageNumber="92">6CI</figureCitation>
,
<figureCitation id="719C2A689315FF99FDB5F1B4FD9F192D" box="[559,609,1774,1798]" captionStart="FIGURE 14" captionStartId="20.[151,250,1014,1036]" captionTargetBox="[340,1248,373,993]" captionTargetId="figure@20.[340,1248,373,993]" captionTargetPageId="20" captionText="FIGURE 14. Mesomere and opisthomere in Caucasian species of Julus, oral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936 G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. Scale bars: 0.3 mm. Abbreviations: fl, flagellum; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere." httpUri="https://zenodo.org/record/1459900/files/figure.png" pageId="3" pageNumber="92">14B</figureCitation>
,
<figureCitation id="719C2A689315FF99FDF6F1B7FD63192D" box="[620,669,1773,1798]" captionStart="FIGURE 15" captionStartId="20.[151,249,1687,1709]" captionTargetBox="[340,1248,1178,1666]" captionTargetId="figure@20.[340,1248,1178,1666]" captionTargetPageId="20" captionText="FIGURE 15. Ventral edge of male segments 7 in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936 C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459902/files/figure.png" pageId="3" pageNumber="92">15B</figureCitation>
,
<figureCitation id="719C2A689315FF99FD32F1B7FD24192D" box="[680,730,1773,1798]" captionStart="FIGURE 16" captionStartId="21.[151,249,1286,1308]" captionTargetBox="[340,1247,583,1264]" captionTargetId="figure@21.[340,1247,583,1264]" captionTargetPageId="21" captionText="FIGURE 16. Striation on metazonites in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459904/files/figure.png" pageId="3" pageNumber="92">16B</figureCitation>
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<figureCitation id="719C2A689315FF99FD7FF1B4FCE8192D" box="[741,790,1774,1798]" captionStart="FIGURE 17" captionStartId="22.[151,249,1033,1055]" captionTargetBox="[340,1248,193,1012]" captionTargetId="figure@22.[340,1248,193,1012]" captionTargetPageId="22" captionText="FIGURE 17. Female vulvae and legs 2 in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. Abbreviations: vu, vulva; op, operculum; bu, bursa; am1 and am2, ampulla 1 and ampulla 2; l2, coxa and prefemur of leg 2. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459906/files/figure.png" pageId="3" pageNumber="92">17B</figureCitation>
, Map 1
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<emphasis id="DBD3EAFF9315FF99FF0DF06FFECD1960" box="[151,307,1845,1867]" italics="true" pageId="3" pageNumber="92">Julus colchicus</emphasis>
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(D).
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<emphasis id="DBD3EAFF9315FF99FF0DF00EFECC1941" box="[151,306,1876,1898]" italics="true" pageId="3" pageNumber="92">Julus colchicus</emphasis>
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: 1791
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(M);
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<bibRefCitation id="8D364B1C9315FF99FDA1F00EFD061941" box="[571,760,1876,1898]" pageId="3" pageNumber="92" refId="ref19152" refString="Kobakhidze, D. N. (1964) [Myriapoda]. Animal life of Georgia, 2 (Arthropoda), 186 - 195 [Tbilisi, in Georgian]" type="book chapter">Kobakhidze, 1964</bibRefCitation>
: 190
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(R); 1965: 393 (R);
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<bibRefCitation id="8D364B1C9315FF99FC60F00EFB581941" box="[1018,1190,1876,1898]" pageId="3" pageNumber="92" refId="ref18710" refString="Golovatch, S. I. (1977) [New or poorly-known species of Julida (Diplopoda) from the Sataplia Reservation, Georgia, Transcaucasia]. Byulleten' Moskovskogo obshchestva ispytatelei prirody, Otdel biologicheskii, 82 (4), 46 - 51. [in Russian, a summary in English]" type="journal article">Golovatch, 1977</bibRefCitation>
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(D, R);
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: 386
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(M);
<treatmentCitation id="680610FC9315FF99FE79F028FD4719A3" author="" box="[483,697,1906,1928]" page="143" pageId="3" pageNumber="92">
<bibRefCitation id="8D364B1C9315FF99FE79F028FD7619A3" box="[483,648,1906,1928]" pageId="3" pageNumber="92" refId="ref20114" refString="Talikadze, D. A. (1984) [On the millipede fauna (Diplopoda) of the Colchidan Province of the Caucasus]. Zoologicheskii zhurnal, 63 (1), 124 - 145 [in Russian, a summary in English]" type="journal article">Talikadze, 1984</bibRefCitation>
: 143
</treatmentCitation>
(M);
<treatmentCitation id="680610FC9315FF99FD6DF028FC4019A3" author="" box="[759,958,1906,1928]" page="181" pageId="3" pageNumber="92">
<bibRefCitation id="8D364B1C9315FF99FD6DF028FC7319A3" box="[759,909,1906,1928]" pageId="3" pageNumber="92" refId="ref18463" refString="Enghoff, H. (2006) The millipedes of Turkey (Diplopoda). Steenstrupia, 29 (2), 175 - 198." type="journal article">Enghoff, 2006</bibRefCitation>
: 181
</treatmentCitation>
(R);
<treatmentCitation id="680610FC9315FF99FC6CF028FB0319A3" author="" box="[1014,1277,1906,1928]" page="452" pageId="3" pageNumber="92">
<bibRefCitation id="8D364B1C9315FF99FC6CF028FB3319A3" box="[1014,1229,1906,1928]" pageId="3" pageNumber="92" refId="ref18252" refString="Chumachenko, Yu. A. (2012) [Species diversity and seasonal activity of millipedes (Diplopoda) in a Taxus and Buxus grove of the Caucasian Nature Reserve]. Shoba, S. A. (Ed.), Materials of the Lectures Presented at the 6 th Congress of the V. V. Dokuchaev Soil Science Society. All-Russia conference with international participation, Petrozavodsk, 13 - 18 August 2012: &quot; Soils of Russia: Present Condition, Prospects of Study and Use &quot;. A School for Young Researchers, l (2), 451 - 452. [508 pp., in Russian]" type="book chapter">Chumachenko, 2012</bibRefCitation>
: 452
</treatmentCitation>
(R); 2016: 409 (R);
<treatmentCitation id="680610FC9315FF99FF6CF0C8FE65198C" author="" box="[246,411,1937,1959]" page="350" pageId="3" pageNumber="92">
<bibRefCitation id="8D364B1C9315FF99FF6CF0C8FE92198C" box="[246,364,1937,1959]" pageId="3" pageNumber="92" refId="ref20367" refString="Zuev, R. V. (2014) Preliminary data on the millipedes (Diplopoda) of the Stavropol Territory, northern Caucasus, Russia. Arthropoda Selecta, 23 (4), 347 - 354." type="journal article">Zuev, 2014</bibRefCitation>
: 350
</treatmentCitation>
(R);
<treatmentCitation id="680610FC9315FF99FE4AF0CBFD0A198C" author="" box="[464,756,1937,1959]" page="25" pageId="3" pageNumber="92">
<bibRefCitation id="8D364B1C9315FF99FE4AF0CBFD2F198C" box="[464,721,1937,1959]" pageId="3" pageNumber="92" refId="ref19277" refString="Korobushkin, D. I., Semenyuk, I. I. &amp; Tuf, I. H. (2016) An annotated checklist of the Chilopoda and Diplopoda (Myriapoda) of the Abrau Peninsula, northwestern Caucasus, Russia. Biodiversity Data Journal, 4, e 7308. https: // doi. org / 10.3897 / BDJ. 4. e 7308" type="journal article">
Korobushkin
<emphasis id="DBD3EAFF9315FF99FDC1F0C8FD76198C" box="[603,648,1937,1959]" italics="true" pageId="3" pageNumber="92">et al</emphasis>
., 2016
</bibRefCitation>
: 25
</treatmentCitation>
(R);
<treatmentCitation id="680610FC9315FF99FCB3F0CBFB8D198C" author="" box="[809,1139,1937,1959]" page="40" pageId="3" pageNumber="92">
<bibRefCitation id="8D364B1C9315FF99FCB3F0CBFBAE198C" box="[809,1104,1937,1959]" pageId="3" pageNumber="92" refId="ref19228" refString="Kokhia, M. S. &amp; Golovatch, S. I. (2018) A checklist of the millipedes of Georgia, Caucasus (Diplopoda). ZooKeys, 741, 35 - 48. https: // doi. org / 10.3897 / zookeys. 741.20042" type="journal article">Kokhia and Golovatch, 2018</bibRefCitation>
: 40
</treatmentCitation>
(M).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1BD65669315FF9DFF0DF08EFD6D1DAB" lastPageId="7" lastPageNumber="96" pageId="3" pageNumber="92" type="materials_examined">
<paragraph id="E91836ED9315FF99FF0DF08EFA6219C7" blockId="3.[151,1436,2004,2029]" box="[151,1436,2004,2029]" pageId="3" pageNumber="92">
<materialsCitation id="59CF3CB09315FF99FF0DF08EFA6219C7" ID-GBIF-Occurrence="1919493213" box="[151,1436,2004,2029]" collectionCode="ZMUM" country="Russia" location="Severskaya Distr." pageId="3" pageNumber="92" specimenCount="35" specimenCount-female="30" specimenCount-juvenile="3" specimenCount-male="2" stateProvince="Krasnodar">
<emphasis id="DBD3EAFF9315FF99FF0DF08EFE7F19C6" bold="true" box="[151,385,2004,2029]" pageId="3" pageNumber="92">Material examined.</emphasis>
<specimenCount id="FFA1FD649315FF99FE13F08EFE1D19C6" box="[393,483,2004,2029]" pageId="3" pageNumber="92" type="male">2 males</specimenCount>
,
<specimenCount id="FFA1FD649315FF99FE75F08EFD9019C6" box="[495,622,2004,2029]" pageId="3" pageNumber="92" type="female">30 females</specimenCount>
,
<specimenCount id="FFA1FD649315FF99FDE0F08EFD0B19C6" box="[634,757,2004,2029]" pageId="3" pageNumber="92" type="juvenile">3 juveniles</specimenCount>
(
<collectionCode id="8FB6AE289315FF99FC9EF08EFC9F19C7" box="[772,865,2004,2028]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="3" pageNumber="92">ZMUM</collectionCode>
),
<emphasis id="DBD3EAFF9315FF99FCEFF08EFB5D19C6" bold="true" box="[885,1187,2004,2029]" pageId="3" pageNumber="92">
<collectingCountry id="91B0767D9315FF99FCEFF08EFC3819C6" box="[885,966,2004,2029]" name="Russia" pageId="3" pageNumber="92">Russia</collectingCountry>
,
<collectingRegion id="2B63F80F9315FF99FC4EF08EFBA919C6" box="[980,1111,2004,2029]" country="Russia" name="Krasnodar" pageId="3" pageNumber="92">Krasnodar</collectingRegion>
Prov.,
</emphasis>
<location id="EC7860369315FF99FB36F08EFA8A19C6" LSID="urn:lsid:plazi:treatment:610E87FB9315FF92FF0DF190FED71F6B:EC7860369315FF99FB36F08EFA8A19C6" box="[1196,1396,2004,2029]" country="Russia" name="Severskaya Distr." pageId="3" pageNumber="92" stateProvince="Krasnodar">Severskaya Distr.</location>
, 3
</materialsCitation>
</paragraph>
<paragraph id="E91836ED9312FF9EFF0DF7C2FEE11D3E" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="93">
10 km S of Ubinskaya,
<emphasis id="DBD3EAFF9312FF9EFE07F7C2FD3A1E84" box="[413,708,152,176]" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="2EA74D6E9312FF9EFE07F7C2FE001E84" box="[413,510,152,175]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFD93F7C3FDAC1E9B" box="[521,594,153,176]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFDC6F7C2FD3A1E84" box="[604,708,152,175]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Carpinus</taxonomicName>
</emphasis>
etc. forest,
<quantity id="2E5F9B089312FF9EFCDCF7CDFC3B1E84" box="[838,965,151,176]" metricMagnitude="2" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="4" pageNumber="93" unit="m" value="375.0" valueMax="450.0" valueMin="300.0">300450 m</quantity>
a.s.l., litter and under bark,
<date id="9D19102D9312FF9EFB67F7CDFA691E9B" box="[1277,1431,151,176]" pageId="4" pageNumber="93" value="1986-07-03" valueMax="1986-07-04" valueMin="1986-07-03">34.VII.1986</date>
; 2 males, 22 females (ZMUM), Mostovskoy Distr., Shedok N of Psebai, karst funnel, wet
<emphasis id="DBD3EAFF9312FF9EFBE6F7E7FA901EFF" box="[1148,1390,188,212]" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="2EA74D6E9312FF9EFBE6F7E7FB3B1EFF" box="[1148,1221,189,212]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFB4AF7E6FAFB1EF8" box="[1232,1285,188,211]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFA8AF7E7FA901EFF" box="[1296,1390,189,212]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
</emphasis>
etc. forest, litter and under bark,
<quantity id="2E5F9B089312FF9EFE41F785FDDD1EDC" box="[475,547,223,248]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="4" pageNumber="93" unit="m" value="650.0">650 m</quantity>
a.s.l.,
<date id="9D19102D9312FF9EFDF4F785FD0B1ED3" box="[622,757,223,248]" pageId="4" pageNumber="93" value="1986-08-04">4.VIII.1986</date>
; 2 males, 1 female, 4 juveniles (ZMUM), Caucasian Nature Reserve, Pslukh
<emphasis id="DBD3EAFF9312FF9EFECCF65FFE8F1F37" box="[342,369,261,284]" italics="true" pageId="4" pageNumber="93">ca</emphasis>
<quantity id="2E5F9B089312FF9EFEE0F65EFE3C1F36" box="[378,450,260,285]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="93" unit="km" value="20.0">20 km</quantity>
E of Krasnaya Polyana,
<emphasis id="DBD3EAFF9312FF9EFD78F65FFBCA1F37" box="[738,1076,260,285]" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="2EA74D6E9312FF9EFD78F65FFCD41F37" box="[738,810,261,284]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFCA2F65FFC671F37" box="[824,921,261,284]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFC3CF65EFC191F36" box="[934,999,260,285]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Abies</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFC6EF65FFBCA1F37" box="[1012,1076,261,284]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
</emphasis>
etc. forest,
<quantity id="2E5F9B089312FF9EFB27F65EFAEA1F37" box="[1213,1300,260,284]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="93" unit="m" value="1000.0">1000 m</quantity>
a.s.l., litter,
<date id="9D19102D9312FF9EFF0DF67DFED21F6B" box="[151,300,295,320]" pageId="4" pageNumber="93" value="1986-08-12">12.VIII.1986</date>
; 2 males, 5 females, 2 juveniles (ZMUM), same locality, 1
<date id="9D19102D9312FF9EFC67F67DFB711F6B" box="[1021,1167,295,320]" pageId="4" pageNumber="93" value="1985-05-08" valueMax="1985-05-20" valueMin="1985-05-08">820.V.1985</date>
; 1 male, 6 females, 2 juveniles (ZMUM), Sochi,
<emphasis id="DBD3EAFF9312FF9EFE56F617FE181F4F" box="[460,486,333,356]" italics="true" pageId="4" pageNumber="93">ca</emphasis>
<quantity id="2E5F9B089312FF9EFE75F616FDD71F4E" box="[495,553,332,357]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="93" unit="km" value="8.0">8 km</quantity>
S of Khosta, Cave “Our Lady”,
<emphasis id="DBD3EAFF9312FF9EFC05F617FB821F48" box="[927,1148,332,356]" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="2EA74D6E9312FF9EFC05F617FC181F4F" box="[927,998,333,356]" class="Magnoliopsida" family="Buxaceae" genus="Buxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Buxales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Buxus</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFC69F617FBC21F4F" box="[1011,1084,333,356]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFBD2F616FB821F48" box="[1096,1148,332,355]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
</emphasis>
etc. forest near entrance, litter and under stones,
<date id="9D19102D9312FF9EFE29F635FDD71FA3" box="[435,553,367,392]" pageId="4" pageNumber="93" value="1985-05-16">16.V.1985</date>
; 1 male, 1 female (ZMUM), same locality,
<taxonomicName id="2EA74D6E9312FF9EFBA1F62BFB871FA3" box="[1083,1145,369,392]" class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFBA1F62BFB871FA3" box="[1083,1145,369,392]" italics="true" pageId="4" pageNumber="93">Taxus</emphasis>
</taxonomicName>
and
<taxonomicName id="2EA74D6E9312FF9EFB27F62BFAFC1FA3" box="[1213,1282,369,392]" class="Magnoliopsida" family="Buxaceae" genus="Buxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Buxales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFB27F62BFAFC1FA3" box="[1213,1282,369,392]" italics="true" pageId="4" pageNumber="93">Buxus</emphasis>
</taxonomicName>
forest, litter,
<date id="9D19102D9312FF9EFF0DF6CEFEEE1F87" box="[151,272,404,428]" pageId="4" pageNumber="93" value="1981-10-28">28.X.1981</date>
; 3 males, 5 females, 1 juvenile (ZMUM), same locality, Lazarevskoye, Saloniki, deciduous forest, litter,
<date id="9D19102D9312FF9EFF0DF6E2FEF11FFB" box="[151,271,440,464]" pageId="4" pageNumber="93" value="1981-10-28">28.X.1981</date>
, all leg. S. Golovatch; 5 males, 25 females, 4 juveniles (ZMUM), Belorechensky Distr., Dolgogusevsky, Belaya River valley, litter,
<date id="9D19102D9312FF9EFE5DF686FDC51FDF" box="[455,571,476,500]" pageId="4" pageNumber="93" value="2009-06-07">7.VI.2009</date>
, leg. L. Rybalov and V. Semenov; 2 males, 4 females, 7 juveniles (ZMUM),
<quantity id="2E5F9B089312FF9EFF0DF6A5FF161C33" box="[151,232,511,536]" metricMagnitude="3" metricUnit="m" metricValue="5.5" pageId="4" pageNumber="93" unit="km" value="5.5">5.5 km</quantity>
NE of Krasnaya Polyana, lower flow of Achipse River,
<quantity id="2E5F9B089312FF9EFC1BF6A5FC321C3C" box="[897,972,511,536]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="93" unit="m" value="600.0">600 m</quantity>
a.s.l., 43°4310”N, 40°1525”E, leg. K. Makarov and A. Matalin; 6 males, 6 females, 4 juveniles (AE), near Adler,
<date id="9D19102D9312FF9EFC6BF57EFBC21C17" box="[1009,1084,548,572]" pageId="4" pageNumber="93" value="2008-01">I.2008</date>
; 8 males, 8 females, 1 juvenile (AE), Caucasian Nature Reserve, near Khosta,
<taxonomicName id="2EA74D6E9312FF9EFD33F513FD181C4B" box="[681,742,585,608]" class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFD33F513FD181C4B" box="[681,742,585,608]" italics="true" pageId="4" pageNumber="93">Taxus</emphasis>
</taxonomicName>
and
<taxonomicName id="2EA74D6E9312FF9EFCBBF513FC981C4B" box="[801,870,585,608]" class="Magnoliopsida" family="Buxaceae" genus="Buxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Buxales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFCBBF513FC981C4B" box="[801,870,585,608]" italics="true" pageId="4" pageNumber="93">Buxus</emphasis>
</taxonomicName>
grove,
<date id="9D19102D9312FF9EFC21F51DFBE51C4B" box="[955,1051,583,608]" pageId="4" pageNumber="93" value="2006-06">VI.2006</date>
; 17 males, 11 females, 2 juveniles (AE), same locality,
<date id="9D19102D9312FF9EFE18F536FE2D1CAE" box="[386,467,620,645]" pageId="4" pageNumber="93" value="2006-05">V.2006</date>
, all leg. Yu. Chumachenko; 2 females, 1 juvenile (SMNG), Khosta,
<taxonomicName id="2EA74D6E9312FF9EFB44F537FAE21CAF" box="[1246,1308,621,644]" class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFB44F537FAE21CAF" box="[1246,1308,621,644]" italics="true" pageId="4" pageNumber="93">Taxus</emphasis>
</taxonomicName>
and
<taxonomicName id="2EA74D6E9312FF9EFACDF537FA621CAF" box="[1367,1436,621,644]" class="Magnoliopsida" family="Buxaceae" genus="Buxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Buxales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFACDF537FA621CAF" box="[1367,1436,621,644]" italics="true" pageId="4" pageNumber="93">Buxus</emphasis>
</taxonomicName>
forest, 43.5317°N, 39.8772°E,
<quantity id="2E5F9B089312FF9EFE6BF5CAFDD41C8C" box="[497,554,656,680]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="93" unit="m" value="80.0">80 m</quantity>
a.s.l.,
<date id="9D19102D9312FF9EFDE8F5CAFCF91C83" box="[626,775,656,680]" pageId="4" pageNumber="93" value="2012-08-07">07.VIII.2012</date>
, leg. F. Walther; 1 male, 1 female (SMNG), Utrish,
<emphasis id="DBD3EAFF9312FF9EFAD3F5CAFCB41CE7" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="2EA74D6E9312FF9EFAD3F5CAFEF61CE7" authority=", Juniperus" authorityName="Juniperus" class="Magnoliopsida" family="Betulaceae" genus="Ostrya" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Ostrya, Juniperus</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFE82F5EEFE301CE7" box="[280,462,692,716]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="species" species="campestre">Acer campestre</taxonomicName>
,
<taxonomicName id="2EA74D6E9312FF9EFE44F5EFFD5F1CE7" authority=", Cornus" authorityName="Cornus" box="[478,673,693,716]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">Quercus, Cornus</taxonomicName>
mas, Pistacia
</emphasis>
etc. coppice, 44.7111°N, 37.4825°E,
<date id="9D19102D9312FF9EFA93F5EEFA801CE7" box="[1289,1406,692,716]" pageId="4" pageNumber="93" value="2004-05-11">11.V.2004</date>
; 1 female, 1 juvenile (SMNG), Mezmai, valley with
<taxonomicName id="2EA74D6E9312FF9EFD5CF583FCF41CDB" box="[710,778,729,752]" class="Magnoliopsida" family="Buxaceae" genus="Buxus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Buxales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFD5CF583FCF41CDB" box="[710,778,729,752]" italics="true" pageId="4" pageNumber="93">Buxus</emphasis>
</taxonomicName>
, under bark, 44.1902°N, 39.9677°E,
<date id="9D19102D9312FF9EFB37F58DFADC1CDB" box="[1197,1314,727,752]" pageId="4" pageNumber="93" value="2004-05-16">16.V.2004</date>
, all leg. K. Voigtländer.
</paragraph>
<caption id="BDD866659312FF9EFF0DF046FBA91944" httpUri="https://zenodo.org/record/1459874/files/figure.png" pageId="4" pageNumber="93" startId="4.[151,249,1820,1842]" targetBox="[344,1236,854,1775]" targetPageId="4">
<paragraph id="E91836ED9312FF9EFF0DF046FBA91944" blockId="4.[151,1437,1820,1903]" pageId="4" pageNumber="93">
<emphasis id="DBD3EAFF9312FF9EFF0DF046FEE61919" bold="true" box="[151,280,1820,1842]" pageId="4" pageNumber="93">FIGURE 1.</emphasis>
Antennae in Caucasian species of
<taxonomicName id="2EA74D6E9312FF9EFD01F046FD2E1919" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[667,720,1820,1842]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFD01F046FD2E1919" box="[667,720,1820,1842]" italics="true" pageId="4" pageNumber="93">Julus</emphasis>
</taxonomicName>
, lateral views.
<emphasis id="DBD3EAFF9312FF9EFCE6F046FC681919" bold="true" box="[892,918,1820,1842]" pageId="4" pageNumber="93">A:</emphasis>
<taxonomicName id="2EA74D6E9312FF9EFC39F047FB141919" authority="Evsyukov, 2016" authorityName="Evsyukov" authorityYear="2016" box="[931,1258,1820,1842]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="DBD3EAFF9312FF9EFC39F047FBCC1919" box="[931,1074,1820,1842]" italics="true" pageId="4" pageNumber="93">J. alexandrae</emphasis>
Evsyukov, 2016
</taxonomicName>
.
<emphasis id="DBD3EAFF9312FF9EFB63F046FAED1919" bold="true" box="[1273,1299,1820,1842]" pageId="4" pageNumber="93">B:</emphasis>
<taxonomicName id="2EA74D6E9312FF9EFA85F047FEAE197B" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="colchicus">
<emphasis id="DBD3EAFF9312FF9EFA85F047FA621919" box="[1311,1436,1820,1842]" italics="true" pageId="4" pageNumber="93">J. colchicus</emphasis>
Lohmander, 1936
</taxonomicName>
.
<emphasis id="DBD3EAFF9312FF9EFEC1F060FE88197B" bold="true" box="[347,374,1850,1872]" pageId="4" pageNumber="93">C:</emphasis>
<taxonomicName id="2EA74D6E9312FF9EFEE4F061FD1B197B" authority="Golovatch, 1981" authorityName="Golovatch" authorityYear="1981" box="[382,741,1850,1872]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="jedryczkowskii">
<emphasis id="DBD3EAFF9312FF9EFEE4F061FDD0197B" box="[382,558,1850,1872]" italics="true" pageId="4" pageNumber="93">J. jedryczkowskii</emphasis>
Golovatch, 1981
</taxonomicName>
.
<emphasis id="DBD3EAFF9312FF9EFD6BF060FCF5197B" bold="true" box="[753,779,1850,1872]" pageId="4" pageNumber="93">D:</emphasis>
<taxonomicName id="2EA74D6E9312FF9EFC8EF061FBD7197B" authority="Verhoeff, 1921" authorityName="Verhoeff" authorityYear="1921" box="[788,1065,1850,1872]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="kubanus">
<emphasis id="DBD3EAFF9312FF9EFC8EF061FC7C197B" box="[788,898,1850,1872]" italics="true" pageId="4" pageNumber="93">J. kubanus</emphasis>
Verhoeff, 1921
</taxonomicName>
.
<emphasis id="DBD3EAFF9312FF9EFBAEF060FBB3197B" bold="true" box="[1076,1101,1850,1872]" pageId="4" pageNumber="93">E:</emphasis>
<taxonomicName id="2EA74D6E9312FF9EFBCCF061FAA0197B" authority="Verhoeff, 1910" authorityName="Verhoeff" authorityYear="1910" box="[1110,1374,1850,1872]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9312FF9EFBCCF061FB47197B" box="[1110,1209,1850,1872]" italics="true" pageId="4" pageNumber="93">J. lignaui</emphasis>
Verhoeff, 1910
</taxonomicName>
.
<emphasis id="DBD3EAFF9312FF9EFAF0F060FA7C197B" bold="true" box="[1386,1410,1850,1872]" pageId="4" pageNumber="93">F:</emphasis>
<taxonomicName id="2EA74D6E9312FF9EFA10F061FE4A1944" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="lindholmi">
<emphasis id="DBD3EAFF9312FF9EFA10F061FF061944" italics="true" pageId="4" pageNumber="93">J. lindholmi</emphasis>
Lohmander, 1936
</taxonomicName>
.
<emphasis id="DBD3EAFF9312FF9EFE24F003FE241944" bold="true" box="[446,474,1881,1903]" pageId="4" pageNumber="93">G:</emphasis>
<taxonomicName id="2EA74D6E9312FF9EFE7AF000FD941944" authority="Evsyukov &amp; Golovatch &amp; Reip, 2018" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[480,618,1881,1903]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="dagestanus" status="sp. nov.">
<emphasis id="DBD3EAFF9312FF9EFE7AF000FD941944" box="[480,618,1881,1903]" italics="true" pageId="4" pageNumber="93">J. dagestanus</emphasis>
</taxonomicName>
<emphasis id="DBD3EAFF9312FF9EFDEAF003FD351944" bold="true" box="[624,715,1881,1903]" pageId="4" pageNumber="93">
<taxonomicNameLabel id="C0E057849312FF9EFDEAF003FD571944" box="[624,681,1881,1903]" pageId="4" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
H:
</emphasis>
<taxonomicName id="2EA74D6E9312FF9EFD48F000FCAC1944" authority="Evsyukov &amp; Golovatch &amp; Reip, 2018" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[722,850,1881,1903]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="khostensis" status="sp. nov.">
<emphasis id="DBD3EAFF9312FF9EFD48F000FCAC1944" box="[722,850,1881,1903]" italics="true" pageId="4" pageNumber="93">J. khostensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0E057849312FF9EFCC2F003FC6F1944" box="[856,913,1881,1903]" pageId="4" pageNumber="93" rank="species">
<emphasis id="DBD3EAFF9312FF9EFCC2F003FC6F1944" bold="true" box="[856,913,1881,1903]" pageId="4" pageNumber="93">sp. n.</emphasis>
</taxonomicNameLabel>
Scale bar: 0.3 mm.
</paragraph>
</caption>
<paragraph id="E91836ED9312FF9EFF5DF0C6FA62199F" blockId="4.[151,1436,1947,2008]" box="[199,1436,1947,1972]" pageId="4" pageNumber="93">
<materialsCitation id="59CF3CB09312FF9EFF5DF0C6FA62199F" ID-GBIF-Occurrence="1919493225" box="[199,1436,1947,1972]" collectionCode="ZMUM" collectorName="Novoprokhladnoe, S" country="Russia" elevation="700" location="Maikop" municipality="Sakhrai" pageId="4" pageNumber="93" specimenCount="2" specimenCount-male="2" stateProvince="Adygea">
<specimenCount id="FFA1FD649312FF9EFF5DF0C6FEDF199F" box="[199,289,1947,1972]" pageId="4" pageNumber="93" type="male">2 males</specimenCount>
(
<collectionCode id="8FB6AE289312FF9EFEAEF0C6FE6F199F" box="[308,401,1948,1972]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="4" pageNumber="93">ZMUM</collectionCode>
),
<emphasis id="DBD3EAFF9312FF9EFE32F0C1FD1F199F" bold="true" box="[424,737,1947,1972]" pageId="4" pageNumber="93">
<collectingCountry id="91B0767D9312FF9EFE32F0C1FE07199F" box="[424,505,1947,1972]" name="Russia" pageId="4" pageNumber="93">Russia</collectingCountry>
,
<collectingRegion id="2B63F80F9312FF9EFD90F0C1FD9A199F" box="[522,612,1947,1972]" country="Russia" name="Adygeya" pageId="4" pageNumber="93">Adygea</collectingRegion>
Republic,
</emphasis>
<collectingMunicipality id="097CAC979312FF9EFD76F0C1FCBF199F" box="[748,833,1947,1972]" pageId="4" pageNumber="93">Sakhrai</collectingMunicipality>
near
<collectorName id="4452533B9312FF9EFC12F0C1FB8B199F" box="[904,1141,1947,1972]" pageId="4" pageNumber="93">Novoprokhladnoe, S</collectorName>
of
<location id="EC7860369312FF9EFB38F0C1FB02199F" LSID="urn:lsid:plazi:treatment:610E87FB9315FF92FF0DF190FED71F6B:EC7860369312FF9EFB38F0C1FB02199F" box="[1186,1276,1947,1972]" country="Russia" municipality="Sakhrai" name="Maikop" pageId="4" pageNumber="93" stateProvince="Adygea">Maikop</location>
,
<elevation id="628AD1DE9312FF9EFA91F0C6FA69199F" box="[1291,1431,1947,1972]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="93" unit="m" value="700.0">
<quantity id="2E5F9B089312FF9EFA91F0C6FAA81998" box="[1291,1366,1948,1972]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="93" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
,
</materialsCitation>
</paragraph>
<paragraph id="E91836ED9312FF9EFF0DF09BFA6219F3" blockId="4.[151,1436,1947,2008]" box="[151,1436,1983,2008]" pageId="4" pageNumber="93">
<taxonomicName id="2EA74D6E9312FF9EFF0DF09BFF2319F3" box="[151,221,1985,2008]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFF0DF09BFF2319F3" box="[151,221,1985,2008]" italics="true" pageId="4" pageNumber="93">Fagus</emphasis>
</taxonomicName>
and
<taxonomicName id="2EA74D6E9312FF9EFE8DF0E5FEA919F3" box="[279,343,1983,2008]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9312FF9EFE8DF0E5FEA919F3" box="[279,343,1983,2008]" italics="true" pageId="4" pageNumber="93">Alnus</emphasis>
</taxonomicName>
forest, litter,
<date id="9D19102D9312FF9EFE69F0E5FD8F19F3" box="[499,625,1983,2008]" pageId="4" pageNumber="93" value="1986-07-07">7.VII.1986</date>
, leg. S. Golovatch; 1 female (SMNG), Sachrai, Novopchladnoe, slope,
</paragraph>
<paragraph id="E91836ED9313FF9FFF0DF7CDFE5E1D8F" blockId="5.[151,1436,151,933]" pageId="5" pageNumber="94">
mixed forest, 44.1363°N, 40.2880°E,
<date id="9D19102D9313FF9FFDD9F7CDFD261E9B" box="[579,728,151,176]" pageId="5" pageNumber="94" value="2005-08-31">31.VIII.2005</date>
, leg. R. Bischof; 7 juveniles (SMNG), S of Novoprokhladnoe, Bzhebs River valley, 44.1365°N, 40.3179°E, under logs,
<date id="9D19102D9313FF9FFCCBF7E6FC181EFE" box="[849,998,188,213]" pageId="5" pageNumber="94" value="2005-08-19">19.VIII.2005</date>
; 1 juvenile (SMNG), same locality,
<date id="9D19102D9313FF9FFF0DF785FED21ED3" box="[151,300,223,248]" pageId="5" pageNumber="94" value="2005-08-30">30.VIII.2005</date>
;2 males, 2 females (SMNG), SE of Novoprokhladnoe, Kuna Valley, wet
<taxonomicName id="2EA74D6E9313FF9FFB17F7BBFAB41ED3" authorityName="Lipsky" authorityYear="1898" box="[1165,1354,223,248]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="DBD3EAFF9313FF9FFB17F7BBFAB41ED3" box="[1165,1354,223,248]" italics="true" pageId="5" pageNumber="94">Fagus orientalis</emphasis>
</taxonomicName>
forest, litter, 44.1131°N, 40.3214°E,
<date id="9D19102D9313FF9FFE73F65EFD811F36" box="[489,639,260,285]" pageId="5" pageNumber="94" value="2005-08-30">30.VIII.2005</date>
; 1 female (SMNG), Ust-Sakhrai, Kuna River valley,
<taxonomicName id="2EA74D6E9313FF9FFB78F65FFA651F36" authorityName="Lipsky" authorityYear="1898" box="[1250,1435,260,285]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="DBD3EAFF9313FF9FFB78F65FFA651F36" box="[1250,1435,260,285]" italics="true" pageId="5" pageNumber="94">Fagus orientalis</emphasis>
</taxonomicName>
etc. forest, under logs, 44.2086°N, 40.2816°E,
<date id="9D19102D9313FF9FFD39F672FCE71F6B" box="[675,793,296,320]" pageId="5" pageNumber="94" value="2004-05-19">19.V.2004</date>
; 2 female, 4 juveniles (SMNG), SE of Novoprokhladnoe, 44.0866°N, 40.4183°E,
<date id="9D19102D9313FF9FFE34F616FDDA1F4F" box="[430,548,332,356]" pageId="5" pageNumber="94" value="2004-05-23">23.V.2004</date>
; 1 male (SMNG), same locality, 44.1000°N, 40.3666°E,
<date id="9D19102D9313FF9FFB5CF616FAC31F4E" box="[1222,1341,332,357]" pageId="5" pageNumber="94" value="2004-05-26">26.V.2004</date>
; 1 male (SMNG), same locality,
<taxonomicName id="2EA74D6E9313FF9FFE59F62BFD441FA3" authority=", Heracleum" authorityName="Heracleum" box="[451,698,367,392]" class="Magnoliopsida" family="Asteraceae" genus="Petasites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9313FF9FFE59F62BFD441FA3" box="[451,698,367,392]" italics="true" pageId="5" pageNumber="94">Petasites, Heracleum</emphasis>
</taxonomicName>
etc. meadow, 44.0999°N, 40.3998°E,
<date id="9D19102D9313FF9FFB0EF635FACA1FA3" box="[1172,1332,367,392]" pageId="5" pageNumber="94" value="2004-05-22" valueMax="2004-05-26" valueMin="2004-05-22">2226.V.2004</date>
; 1 male (SMNG), E of Novoprokhladnoe, Sachrai Valley,
<taxonomicName id="2EA74D6E9313FF9FFD6DF6CFFC461F86" authorityName="Lipsky" authorityYear="1898" box="[759,952,404,429]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="DBD3EAFF9313FF9FFD6DF6CFFC461F86" box="[759,952,404,429]" italics="true" pageId="5" pageNumber="94">Fagus orientalis</emphasis>
</taxonomicName>
etc. forest, 44.1365°N, 40.3179°E,
<date id="9D19102D9313FF9FFAE8F6CEFEF21FFB" pageId="5" pageNumber="94" value="2004-05-19" valueMax="2004-05-21" valueMin="2004-05-19">19 21.V.2004</date>
, all leg. K. Voigtländer; 4 males, 6 females (AE), Caucasian Nature Reserve, near Guzeripl,
<taxonomicName id="2EA74D6E9313FF9FFABFF6E3FA951FFB" box="[1317,1387,441,464]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9313FF9FFABFF6E3FA951FFB" box="[1317,1387,441,464]" italics="true" pageId="5" pageNumber="94">Fagus</emphasis>
</taxonomicName>
and
<taxonomicName id="2EA74D6E9313FF9FFF0DF686FF281FDE" box="[151,214,476,501]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9313FF9FFF0DF686FF281FDE" box="[151,214,476,501]" italics="true" pageId="5" pageNumber="94">Abies</emphasis>
</taxonomicName>
forest,
<quantity id="2E5F9B089313FF9FFEAAF686FE761FDF" box="[304,392,476,500]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="94" unit="m" value="1000.0">1000 m</quantity>
a.s.l.,
<date id="9D19102D9313FF9FFE4DF686FDD51FDF" box="[471,555,476,500]" pageId="5" pageNumber="94" value="2009-05">V.2009</date>
; 1 male, 1 female (AE), same locality,
<quantity id="2E5F9B089313FF9FFC62F686FBBC1FDF" box="[1016,1090,476,500]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="94" unit="m" value="700.0">700 m</quantity>
a.s.l.,
<date id="9D19102D9313FF9FFB08F686FAF61FDF" box="[1170,1288,476,500]" pageId="5" pageNumber="94" value="2009-05-21">21.V.2009</date>
; 4 males, 11 females (AE), same locality,
<quantity id="2E5F9B089313FF9FFE7CF55AFDC31C3C" box="[486,573,512,536]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="94" unit="m" value="1000.0">1000 m</quantity>
a.s.l.,
<date id="9D19102D9313FF9FFD11F55AFC931C33" box="[651,877,512,536]" pageId="5" pageNumber="94" value="2009-06-23" valueMax="2009-07-24" valueMin="2009-06-23">23.VI.24.VII.2009</date>
; 2 males, 7 females (AE), same locality,
<date id="9D19102D9313FF9FFAD1F55AFEDD1C17" pageId="5" pageNumber="94" value="2009-07-08" valueMax="2009-07-24" valueMin="2009-07-08">8.VII. 24.VII.2009</date>
; 14 males, 17 females (AE), same locality,
<date id="9D19102D9313FF9FFC86F57EFC851C17" box="[796,891,548,572]" pageId="5" pageNumber="94" value="2009-06">VI.2009</date>
, all leg. Yu. Chumachenko; 2 males, 5 females (ZMUM), same locality, forest, summer 1976, leg. B. Striganova; 3 males, 2 females (AE), Caucasian Nature Reserve, NE slope of Mt Oshten,
<quantity id="2E5F9B089313FF9FFDBDF536FD811CAF" box="[551,639,620,645]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="5" pageNumber="94" unit="m" value="2150.0">2150 m</quantity>
a.s.l.,
<date id="9D19102D9313FF9FFD55F536FCA21CAF" box="[719,860,620,644]" pageId="5" pageNumber="94" value="2004-07-17">17.VII.2004</date>
; 5 males, 6 females (AE), near Maikop,
<taxonomicName id="2EA74D6E9313FF9FFAA4F537FA621CAF" box="[1342,1436,621,644]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9313FF9FFAA4F537FA621CAF" box="[1342,1436,621,644]" italics="true" pageId="5" pageNumber="94">Quercus</emphasis>
</taxonomicName>
forest, pitfall trapping,
<date id="9D19102D9313FF9FFE30F5CAFD8C1C83" box="[426,626,656,680]" pageId="5" pageNumber="94" value="2009-04-14" valueMax="2009-05-01" valueMin="2009-04-14">14.IV.01.V.2009</date>
; 7 males, 4 females (AE), near Maikop, 44°20.724N, 40°11.368E,
<date id="9D19102D9313FF9FFF0DF5EEFEA51CE6" box="[151,347,692,717]" pageId="5" pageNumber="94" value="2011-03-16" valueMax="2011-04-01" valueMin="2011-03-16">16.III.1.IV.2011</date>
; 3 males, 5 females (AE), same locality,
<date id="9D19102D9313FF9FFCB8F5EEFC221CE6" box="[802,988,692,717]" pageId="5" pageNumber="94" value="2011-04-15" valueMax="2011-05-02" valueMin="2011-04-15">15.IV.2.V.2011</date>
; 2 males, 5 females (AE), same locality,
<date id="9D19102D9313FF9FFF0DF58DFEAD1CDB" box="[151,339,727,752]" pageId="5" pageNumber="94" value="2011-05-15" valueMax="2011-06-04" valueMin="2011-05-15">15.V.4.VI.2011</date>
; 1 male, 2 females (AE), same locality,
<date id="9D19102D9313FF9FFCBEF58DFC131CDB" box="[804,1005,727,752]" pageId="5" pageNumber="94" value="2011-06-04" valueMax="2011-06-16" valueMin="2011-06-04">4.VI.16.VI.2011</date>
; 2 males, 8 females, 1 juvenile (AE), same locality,
<date id="9D19102D9313FF9FFEDAF5A6FDDE1D3E" box="[320,544,764,789]" pageId="5" pageNumber="94" value="2011-06-16" valueMax="2011-07-15" valueMin="2011-06-16">16.VI.15.VII.2011</date>
; 12 females, 1 juvenile (AE), same locality,
<date id="9D19102D9313FF9FFBA8F5A6FAE21D3E" box="[1074,1308,764,789]" pageId="5" pageNumber="94" value="2011-07-15" valueMax="2011-07-29" valueMin="2011-07-15">15.VII.29.VII.2011</date>
; 7 females (AE), same locality,
<date id="9D19102D9313FF9FFE1DF445FD851D13" box="[391,635,799,824]" pageId="5" pageNumber="94" value="2011-07-29" valueMax="2011-08-16" valueMin="2011-07-29">29.VII.16.VIII.2011</date>
; 1 male, 6 females (AE), same locality,
<date id="9D19102D9313FF9FFBCEF445FAEA1D13" box="[1108,1300,799,824]" pageId="5" pageNumber="94" value="2011-09-16" valueMax="2011-10-02" valueMin="2011-09-16">16.IX.2.X.2011</date>
; 3 males, 4 females, 2 juveniles (AE), same locality,
<date id="9D19102D9313FF9FFDF6F41EFCDC1D77" box="[620,802,836,860]" pageId="5" pageNumber="94" value="2011-10-01" valueMax="2011-10-17" valueMin="2011-10-01">1.X.17.X.2011</date>
, all leg. Yu. Chumachenko; 1 male (AE),
<quantity id="2E5F9B089313FF9FFA9BF41EFAC51D76" box="[1281,1339,836,861]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="94" unit="km" value="6.0">6 km</quantity>
NNE of Guzeripl, mouth of Kisha River, mixed forest,
<date id="9D19102D9313FF9FFD31F432FCD31DAB" box="[683,813,872,896]" pageId="5" pageNumber="94" value="2013-06-11">11.VI.2013</date>
, leg. R. Zuev; 1 male, 1 female (AE), Guzeripl, mixed forest,
<date id="9D19102D9313FF9FFF7EF4D6FEA61D8F" box="[228,344,908,932]" pageId="5" pageNumber="94" value="2012-06-03">3.VI.2012</date>
, leg.?
</paragraph>
<caption id="BDD866659313FF9FFF0DF11BFF381898" httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="5" pageNumber="94" startId="5.[151,249,1601,1623]" targetBox="[340,1248,985,1580]" targetPageId="5">
<paragraph id="E91836ED9313FF9FFF0DF11BFF381898" blockId="5.[151,1437,1601,1715]" pageId="5" pageNumber="94">
<emphasis id="DBD3EAFF9313FF9FFF0DF11BFEEB187C" bold="true" box="[151,277,1601,1623]" pageId="5" pageNumber="94">FIGURE 2.</emphasis>
Gnathochilaria in Caucasian species of
<taxonomicName id="2EA74D6E9313FF9FFD22F11BFD13187C" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[696,749,1601,1623]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="DBD3EAFF9313FF9FFD22F11BFD13187C" box="[696,749,1601,1623]" italics="true" pageId="5" pageNumber="94">Julus</emphasis>
</taxonomicName>
, ventral views.
<emphasis id="DBD3EAFF9313FF9FFC0FF11BFC51187C" bold="true" box="[917,943,1601,1623]" pageId="5" pageNumber="94">A:</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFC22F118FB0B187C" authority="Evsyukov, 2016" authorityName="Evsyukov" authorityYear="2016" box="[952,1269,1601,1623]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="DBD3EAFF9313FF9FFC22F118FBBA187C" box="[952,1092,1601,1623]" italics="true" pageId="5" pageNumber="94">J. alexandrae</emphasis>
Evsyukov, 2016
</taxonomicName>
.
<emphasis id="DBD3EAFF9313FF9FFA9AF11BFAE4187C" bold="true" box="[1280,1306,1601,1623]" pageId="5" pageNumber="94">B:</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFAB8F118FEAE185D" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="colchicus">
<emphasis id="DBD3EAFF9313FF9FFAB8F118FA62187C" box="[1314,1436,1601,1623]" italics="true" pageId="5" pageNumber="94">J. colchicus</emphasis>
Lohmander, 1936
</taxonomicName>
.
<emphasis id="DBD3EAFF9313FF9FFEC1F13AFE88185D" bold="true" box="[347,374,1632,1654]" pageId="5" pageNumber="94">C:</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFEE4F13BFD1B185D" authority="Golovatch, 1981" authorityName="Golovatch" authorityYear="1981" box="[382,741,1632,1654]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="jedryczkowskii">
<emphasis id="DBD3EAFF9313FF9FFEE4F13BFDD0185D" box="[382,558,1632,1654]" italics="true" pageId="5" pageNumber="94">J. jedryczkowskii</emphasis>
Golovatch, 1981
</taxonomicName>
.
<emphasis id="DBD3EAFF9313FF9FFD6BF13AFCF5185D" bold="true" box="[753,779,1632,1654]" pageId="5" pageNumber="94">D:</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFC8EF13BFBD6185D" authority="Verhoeff, 1921" authorityName="Verhoeff" authorityYear="1921" box="[788,1064,1632,1654]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="kubanus">
<emphasis id="DBD3EAFF9313FF9FFC8EF13BFC7C185D" box="[788,898,1632,1654]" italics="true" pageId="5" pageNumber="94">J. kubanus</emphasis>
Verhoeff, 1921
</taxonomicName>
.
<emphasis id="DBD3EAFF9313FF9FFBAEF13AFBB3185D" bold="true" box="[1076,1101,1632,1654]" pageId="5" pageNumber="94">E:</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFBCCF13BFAA0185D" authority="Verhoeff, 1910" authorityName="Verhoeff" authorityYear="1910" box="[1110,1374,1632,1654]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9313FF9FFBCCF13BFB47185D" box="[1110,1209,1632,1654]" italics="true" pageId="5" pageNumber="94">J. lignaui</emphasis>
Verhoeff, 1910
</taxonomicName>
.
<emphasis id="DBD3EAFF9313FF9FFAF0F13AFA7C185D" bold="true" box="[1386,1410,1632,1654]" pageId="5" pageNumber="94">F:</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFA10F13BFE4818BF" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="lindholmi">
<emphasis id="DBD3EAFF9313FF9FFA10F13BFF0618BF" italics="true" pageId="5" pageNumber="94">J. lindholmi</emphasis>
Lohmander, 1936
</taxonomicName>
.
<emphasis id="DBD3EAFF9313FF9FFE25F124FE2518BF" bold="true" box="[447,475,1662,1684]" pageId="5" pageNumber="94">G:</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFE79F1DAFCD918BF" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" box="[483,807,1662,1685]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="subalpinus">
<emphasis id="DBD3EAFF9313FF9FFE79F1DAFD9718BF" box="[483,617,1662,1685]" italics="true" pageId="5" pageNumber="94">J. subalpinus</emphasis>
Lohmander, 1936
</taxonomicName>
.
<emphasis id="DBD3EAFF9313FF9FFCAAF124FCB218BF" bold="true" box="[816,844,1662,1684]" pageId="5" pageNumber="94">H:</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFCCEF1DAFC2018BF" authority="Evsyukov &amp; Golovatch &amp; Reip, 2018" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[852,990,1662,1685]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="dagestanus" status="sp. nov.">
<emphasis id="DBD3EAFF9313FF9FFCCEF1DAFC2018BF" box="[852,990,1662,1685]" italics="true" pageId="5" pageNumber="94">J. dagestanus</emphasis>
</taxonomicName>
<emphasis id="DBD3EAFF9313FF9FFC7FF124FBC918BF" bold="true" box="[997,1079,1662,1684]" pageId="5" pageNumber="94">
<taxonomicNameLabel id="C0E057849313FF9FFC7FF124FBE018BF" box="[997,1054,1662,1684]" pageId="5" pageNumber="94" rank="species">sp. n.</taxonomicNameLabel>
I:
</emphasis>
<taxonomicName id="2EA74D6E9313FF9FFBA4F1DAFB4118BF" authority="Evsyukov &amp; Golovatch &amp; Reip, 2018" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[1086,1215,1662,1685]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="khostensis" status="sp. nov.">
<emphasis id="DBD3EAFF9313FF9FFBA4F1DAFB4118BF" box="[1086,1215,1662,1685]" italics="true" pageId="5" pageNumber="94">J. khostensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0E057849313FF9FFB5CF124FB0118BF" box="[1222,1279,1662,1684]" pageId="5" pageNumber="94" rank="species">
<emphasis id="DBD3EAFF9313FF9FFB5CF124FB0118BF" bold="true" box="[1222,1279,1662,1684]" pageId="5" pageNumber="94">sp. n.</emphasis>
</taxonomicNameLabel>
Scale bars: 0.3 mm.
</paragraph>
</caption>
<paragraph id="E91836ED9313FF9FFF5DF1BAFB3E196A" blockId="5.[151,1436,1760,2037]" pageId="5" pageNumber="94">
<materialsCitation id="59CF3CB09313FF9FFF5DF1BAFA811936" ID-GBIF-Occurrence="1919493218" collectingDate="1986-08-02" collectionCode="ZMUM" collectorName="S. Golovatch" country="Russia" county="Pass Klukhor" elevation="2805" location="Karachaevo-Cherkessia" municipality="Canyon Gonachkhir" pageId="5" pageNumber="94" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<specimenCount id="FFA1FD649313FF9FFF5DF1BAFED018D2" box="[199,302,1760,1785]" pageId="5" pageNumber="94" type="male">3 males</specimenCount>
,
<specimenCount id="FFA1FD649313FF9FFEDCF1BAFE3E18D2" box="[326,448,1760,1785]" pageId="5" pageNumber="94" type="female">3 females</specimenCount>
(
<collectionCode id="8FB6AE289313FF9FFE47F1BAFDC418D3" box="[477,570,1760,1784]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="5" pageNumber="94">ZMUM</collectionCode>
),
<emphasis id="DBD3EAFF9313FF9FFDC1F1BAFC1518D2" bold="true" box="[603,1003,1760,1785]" pageId="5" pageNumber="94">
<collectingCountry id="91B0767D9313FF9FFDC1F1BAFD5218D2" box="[603,684,1760,1785]" name="Russia" pageId="5" pageNumber="94">Russia</collectingCountry>
,
<location id="EC7860369313FF9FFD5CF1BAFC1818D2" LSID="urn:lsid:plazi:treatment:610E87FB9315FF92FF0DF190FED71F6B:EC7860369313FF9FFD5CF1BAFC1818D2" box="[710,998,1760,1785]" country="Russia" county="Pass Klukhor" municipality="Canyon Gonachkhir" name="Karachaevo-Cherkessia" pageId="5" pageNumber="94">Karachaevo-Cherkessia</location>
,
</emphasis>
<location id="EC7860369313FF9FFC65F1BAFAD518D3" LSID="urn:lsid:plazi:treatment:610E87FB9315FF92FF0DF190FED71F6B:EC7860369313FF9FFC65F1BAFAD518D3" box="[1023,1323,1760,1785]" country="Russia" county="Pass Klukhor" municipality="Canyon Gonachkhir" name="Teberda Nature Reserve" pageId="5" pageNumber="94">Teberda Nature Reserve</location>
,
<collectingMunicipality id="097CAC979313FF9FFADFF1BAFEE31936" pageId="5" pageNumber="94">Canyon Gonachkhir</collectingMunicipality>
,
<collectingCounty id="00794E619313FF9FFEB1F05FFE381936" box="[299,454,1796,1821]" pageId="5" pageNumber="94">Pass Klukhor</collectingCounty>
,
<elevation id="628AD1DE9313FF9FFE4EF05EFD4C1936" box="[468,690,1796,1821]" metricMagnitude="3" metricUnit="m" metricValue="2.805" metricValueMax="2.85" metricValueMin="2.76" pageId="5" pageNumber="94" unit="m" value="2805.0" valueMax="2850.0" valueMin="2760.0">
<quantity id="2E5F9B089313FF9FFE4EF05EFD8C1937" box="[468,626,1796,1821]" metricMagnitude="3" metricUnit="m" metricValue="2.805" metricValueMax="2.85" metricValueMin="2.76" pageId="5" pageNumber="94" unit="m" value="2805.0" valueMax="2850.0" valueMin="2760.0">27602850 m</quantity>
a.s.l.
</elevation>
, alpine meadow, under stones,
<collectingDate id="8D5DE9C59313FF9FFB85F05EFB5B1936" box="[1055,1189,1796,1821]" pageId="5" pageNumber="94" value="1986-08-02">
<date id="9D19102D9313FF9FFB85F05EFB5B1936" box="[1055,1189,1796,1821]" pageId="5" pageNumber="94" value="1986-08-02">2.VIII.1986</date>
</collectingDate>
, leg.
<collectorName id="4452533B9313FF9FFB7DF05FFA811936" box="[1255,1407,1796,1821]" pageId="5" pageNumber="94">S. Golovatch</collectorName>
</materialsCitation>
;
<materialsCitation id="59CF3CB09313FF9FFA14F05FFB42196A" ID-GBIF-Occurrence="1919493206" collectingDate="2009-06-30" collectingDateMax="2009-07-09" collectingDateMin="2009-06-30" collectorName="Yu. Chumachenko" country="Russia" location="Caucasian Nature Reserve" municipality="Zakan" pageId="5" pageNumber="94" specimenCount="1" specimenCount-male="1">
<specimenCount id="FFA1FD649313FF9FFA14F05FFF33196A" pageId="5" pageNumber="94" type="male">1 male</specimenCount>
(AE),
<location id="EC7860369313FF9FFE80F072FDBA196B" LSID="urn:lsid:plazi:treatment:610E87FB9315FF92FF0DF190FED71F6B:EC7860369313FF9FFE80F072FDBA196B" box="[282,580,1832,1857]" country="Russia" municipality="Zakan" name="Caucasian Nature Reserve" pageId="5" pageNumber="94">Caucasian Nature Reserve</location>
, near
<collectingMunicipality id="097CAC979313FF9FFD1DF072FD2E196A" box="[647,720,1832,1857]" pageId="5" pageNumber="94">Zakan</collectingMunicipality>
,
<collectingDate id="8D5DE9C59313FF9FFD41F072FC50196B" box="[731,942,1832,1856]" pageId="5" pageNumber="94" value="2009-06-30" valueMax="2009-07-09" valueMin="2009-06-30">
<date id="9D19102D9313FF9FFD41F072FC50196B" box="[731,942,1832,1856]" pageId="5" pageNumber="94" value="2009-06-30" valueMax="2009-07-09" valueMin="2009-06-30">30.VI.9.VII.2009</date>
</collectingDate>
, leg.
<collectorName id="4452533B9313FF9FFC73F073FB42196A" box="[1001,1212,1832,1857]" pageId="5" pageNumber="94">Yu. Chumachenko</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="E91836ED9313FF9FFF5DF017FAE91987" blockId="5.[151,1436,1760,2037]" pageId="5" pageNumber="94">
1 male (ZMUM),
<emphasis id="DBD3EAFF9313FF9FFE0AF016FD52194E" bold="true" box="[400,684,1868,1893]" pageId="5" pageNumber="94">Russia, Stavropol Prov.,</emphasis>
Georgievsk,
<emphasis id="DBD3EAFF9313FF9FFCDBF017FC1C194F" box="[833,994,1869,1892]" italics="true" pageId="5" pageNumber="94">
<taxonomicName id="2EA74D6E9313FF9FFCDBF017FC5C194F" box="[833,930,1869,1892]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="2EA74D6E9313FF9FFC37F017FC1C194F" box="[941,994,1869,1892]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
</emphasis>
and
<taxonomicName id="2EA74D6E9313FF9FFB82F017FBA0194F" box="[1048,1118,1869,1892]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9313FF9FFB82F017FBA0194F" box="[1048,1118,1869,1892]" italics="true" pageId="5" pageNumber="94">Fagus</emphasis>
</taxonomicName>
forest, litter and under bark,
<date id="9D19102D9313FF9FFF0DF02AFEF519A2" box="[151,267,1904,1929]" pageId="5" pageNumber="94" value="1985-06-07">7.VI.1985</date>
, leg. S. Golovatch; 1 male, 3 females, 1 juvenile (AE), Kochubeevsky Distr., near Nevinnomyssk, floodplain of Bolshoy Zelenchuk River, 44.5970°N, 41,9252°E,
<quantity id="2E5F9B089313FF9FFCF0F0CFFC4F1987" box="[874,945,1941,1965]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="94" unit="m" value="340.0">340 m</quantity>
a.s.l.,
<date id="9D19102D9313FF9FFC63F0CEFB821986" box="[1017,1148,1940,1965]" pageId="5" pageNumber="94" value="2017-11-06">06.XI.2017</date>
, leg. R. Zuev.
</paragraph>
<paragraph id="E91836ED9313FF9CFF5DF0E2FD5E1F86" blockId="5.[151,1436,1760,2037]" lastBlockId="6.[151,1436,151,429]" lastPageId="6" lastPageNumber="95" pageId="5" pageNumber="94">
1 male, 3 females, 1 juvenile (ZMUM),
<emphasis id="DBD3EAFF9313FF9FFD0BF0E2FCF519FA" bold="true" box="[657,779,1976,2001]" pageId="5" pageNumber="94">Abkhazia,</emphasis>
Lake Ritsa,
<quantity id="2E5F9B089313FF9FFC07F0E2FBD419FB" box="[925,1066,1976,2001]" metricMagnitude="3" metricUnit="m" metricValue="1.025" metricValueMax="1.1" metricValueMin="0.95" pageId="5" pageNumber="94" unit="m" value="1025.0" valueMax="1100.0" valueMin="950.0">9501100 m</quantity>
a.s.l.,
<emphasis id="DBD3EAFF9313FF9FFBEFF0E3FA6219FB" box="[1141,1436,1976,2001]" italics="true" pageId="5" pageNumber="94">
<taxonomicName id="2EA74D6E9313FF9FFBEFF0E3FB4019FB" box="[1141,1214,1977,2000]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="2EA74D6E9313FF9FFB51F0E2FAF219FA" box="[1227,1292,1976,2001]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">Abies</taxonomicName>
,
<taxonomicName id="2EA74D6E9313FF9FFA82F0E3FAA419FB" box="[1304,1370,1977,2000]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
,
<taxonomicName id="2EA74D6E9313FF9FFAFDF0E3FA6219FB" box="[1383,1436,1977,2000]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="5" pageNumber="94" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
</emphasis>
etc. forest, litter, under bark and stones,
<date id="9D19102D9313FF9FFDC7F086FCE219DE" box="[605,796,2012,2037]" pageId="5" pageNumber="94" value="1986-08-13" valueMax="1986-08-14" valueMin="1986-08-13">1314.VIII.1986</date>
; 19 males, 14 females, 15 juveniles (ZMUM), Myussera Nature Reserve,
<quantity id="2E5F9B089310FF9CFECCF7C2FE361E84" box="[342,456,152,176]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="13.0" metricValueMin="2.0" pageId="6" pageNumber="95" unit="m" value="75.0" valueMax="130.0" valueMin="20.0">20130 m</quantity>
a.s.l., mixed deciduous forest (
<emphasis id="DBD3EAFF9310FF9CFCAAF7C2FC171E9B" box="[816,1001,151,176]" italics="true" pageId="6" pageNumber="95">
<taxonomicName id="2EA74D6E9310FF9CFCAAF7C2FC621E84" box="[816,924,152,175]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">Castanea</taxonomicName>
,
<taxonomicName id="2EA74D6E9310FF9CFC33F7CDFC171E9B" box="[937,1001,151,176]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">Alnus</taxonomicName>
</emphasis>
etc.), litter, under bark and stones,
<date id="9D19102D9310FF9CFA1AF7C2FEE81EFF" pageId="6" pageNumber="95" value="1983-04-08" valueMax="1983-04-10" valueMin="1983-04-08">8 10.IV.1983</date>
; 1 male (ZMUM), same locality and dates; 1 male, 9 females (ZMUM), Sukhumi Distr., Bzyb River Valley, Pskhu,
<quantity id="2E5F9B089310FF9CFED4F7BAFE2B1EDC" box="[334,469,224,248]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="6" pageNumber="95" unit="m" value="800.0" valueMax="900.0" valueMin="700.0">700900 m</quantity>
a.s.l.,
<emphasis id="DBD3EAFF9310FF9CFDB4F7BBFC961EDC" box="[558,872,224,248]" italics="true" pageId="6" pageNumber="95">
<taxonomicName id="2EA74D6E9310FF9CFDB4F7BBFD881ED3" box="[558,630,225,248]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="2EA74D6E9310FF9CFD10F7BAFD151EDC" box="[650,747,224,247]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="2EA74D6E9310FF9CFD64F7BAFC961EDC" box="[766,872,224,247]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">Castanea</taxonomicName>
</emphasis>
etc. forest, litter, under bark and stones,
<date id="9D19102D9310FF9CFAE8F785FED21F36" pageId="6" pageNumber="95" value="1986-08-15" valueMax="1986-08-16" valueMin="1986-08-15">15 16.VIII.1986</date>
; 5 males, 2 females, 1 juvenile (ZMUM), environs of Sukhumi, near Cave Kelassuri, litter,
<date id="9D19102D9310FF9CFF0DF672FEEA1F6B" box="[151,276,296,320]" pageId="6" pageNumber="95" value="1983-04-11">11.IV.1983</date>
, all leg. S. Golovatch; 1 male (ZMUM), near Sukhumi, under stones,
<date id="9D19102D9310FF9CFBB1F672FB571F6B" box="[1067,1193,296,320]" pageId="6" pageNumber="95" value="1971-04-22">22.IV.1971</date>
, leg. N. Zalesskaja; 2 males, 3 females (ZMUM), Abkhazia, Sukhumi Distr., Yashtukha, tree nursery,
<date id="9D19102D9310FF9CFB86F616FB631F4E" box="[1052,1181,332,357]" pageId="6" pageNumber="95" value="1995-03-29">29.III.1995</date>
, leg. A. Markossian; 1 male (ZMUM), Abkhazia, Sukhum, 42°58.4N, 41°04.04E,
<emphasis id="DBD3EAFF9310FF9CFCCCF62BFC8E1FA3" box="[854,880,369,392]" italics="true" pageId="6" pageNumber="95">ca</emphasis>
<quantity id="2E5F9B089310FF9CFCE0F635FC481FAC" box="[890,950,367,392]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="95" unit="m" value="50.0">50 m</quantity>
a.s.l., litter,
<taxonomicName id="2EA74D6E9310FF9CFBD0F635FB381FA3" box="[1098,1222,367,392]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9310FF9CFBD0F635FB381FA3" box="[1098,1222,367,392]" italics="true" pageId="6" pageNumber="95">Eucalyptus</emphasis>
</taxonomicName>
, tree shaking etc.,
<date id="9D19102D9310FF9CFF0DF6CEFEEA1F87" box="[151,276,404,428]" pageId="6" pageNumber="95" value="2003-12-07">7.XII.2003</date>
, leg. Yu. Marusik and G. Antipova.
</paragraph>
<caption id="BDD866659310FF9CFF0DF281FDD11969" httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="6" pageNumber="95" startId="6.[151,249,1499,1521]" targetBox="[340,1248,481,1476]" targetPageId="6">
<paragraph id="E91836ED9310FF9CFF0DF281FDD11969" blockId="6.[151,1437,1499,1858]" pageId="6" pageNumber="95">
<emphasis id="DBD3EAFF9310FF9CFF0DF281FEEA1BDA" bold="true" box="[151,276,1499,1521]" pageId="6" pageNumber="95">FIGURE 3.</emphasis>
Male legs 1 in
<taxonomicName id="2EA74D6E9310FF9CFE25F281FCE11BDA" authority="Evsyukov, 2016" authorityName="Evsyukov" authorityYear="2016" box="[447,799,1499,1521]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="DBD3EAFF9310FF9CFE25F281FD8E1BDA" box="[447,624,1499,1521]" italics="true" pageId="6" pageNumber="95">Julus alexandrae</emphasis>
Evsyukov, 2016
</taxonomicName>
(
<emphasis id="DBD3EAFF9310FF9CFCAAF281FCBC1BDA" bold="true" box="[816,834,1499,1521]" pageId="6" pageNumber="95">A</emphasis>
),
<taxonomicName id="2EA74D6E9310FF9CFCC0F286FB6D1BDA" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" box="[858,1171,1499,1521]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="colchicus">
<emphasis id="DBD3EAFF9310FF9CFCC0F286FC2B1BDA" box="[858,981,1499,1521]" italics="true" pageId="6" pageNumber="95">J. colchicus</emphasis>
Lohmander, 1936
</taxonomicName>
(
<emphasis id="DBD3EAFF9310FF9CFB3EF281FB2A1BDA" bold="true" box="[1188,1236,1499,1521]" pageId="6" pageNumber="95">BD</emphasis>
),
<taxonomicName id="2EA74D6E9310FF9CFB76F286FEBE183B" authority="Golovatch, 1981" authorityName="Golovatch" authorityYear="1981" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="jedryczkowskii">
<emphasis id="DBD3EAFF9310FF9CFB76F286FA621BDA" box="[1260,1436,1499,1521]" italics="true" pageId="6" pageNumber="95">J. jedryczkowskii</emphasis>
Golovatch, 1981
</taxonomicName>
(
<emphasis id="DBD3EAFF9310FF9CFED4F2A0FE85183B" bold="true" box="[334,379,1530,1552]" pageId="6" pageNumber="95">EF</emphasis>
),
<taxonomicName id="2EA74D6E9310FF9CFE0BF2A1FD65183B" authority="Verhoeff, 1921" authorityName="Verhoeff" authorityYear="1921" box="[401,667,1530,1552]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="kubanus">
<emphasis id="DBD3EAFF9310FF9CFE0BF2A1FE03183B" box="[401,509,1530,1552]" italics="true" pageId="6" pageNumber="95">J. kubanus</emphasis>
Verhoeff, 1921
</taxonomicName>
(
<emphasis id="DBD3EAFF9310FF9CFD30F2A0FD2D183B" bold="true" box="[682,723,1530,1552]" pageId="6" pageNumber="95">GI</emphasis>
),
<taxonomicName id="2EA74D6E9310FF9CFD73F2A1FC19183B" authority="Verhoeff, 1910" authorityName="Verhoeff" authorityYear="1910" box="[745,999,1530,1552]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="lignaui">
<emphasis id="DBD3EAFF9310FF9CFD73F2A1FCB7183B" box="[745,841,1530,1552]" italics="true" pageId="6" pageNumber="95">J. lignaui</emphasis>
Verhoeff, 1910
</taxonomicName>
(
<emphasis id="DBD3EAFF9310FF9CFC6FF2A0FBD9183B" bold="true" box="[1013,1063,1530,1552]" pageId="6" pageNumber="95">JM</emphasis>
),
<taxonomicName id="2EA74D6E9310FF9CFBA6F2A1FA90183B" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" box="[1084,1390,1530,1552]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="lindholmi">
<emphasis id="DBD3EAFF9310FF9CFBA6F2A1FB4A183B" box="[1084,1204,1530,1552]" italics="true" pageId="6" pageNumber="95">J. lindholmi</emphasis>
Lohmander, 1936
</taxonomicName>
(
<emphasis id="DBD3EAFF9310FF9CFAE7F2A0FF581805" bold="true" pageId="6" pageNumber="95">N P</emphasis>
),
<taxonomicName id="2EA74D6E9310FF9CFF27F140FDFE1805" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" box="[189,512,1560,1583]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="subalpinus">
<emphasis id="DBD3EAFF9310FF9CFF27F140FEBA1805" box="[189,324,1560,1583]" italics="true" pageId="6" pageNumber="95">J. subalpinus</emphasis>
Lohmander, 1936
</taxonomicName>
(
<emphasis id="DBD3EAFF9310FF9CFD8AF142FDBC1805" bold="true" box="[528,578,1560,1582]" pageId="6" pageNumber="95">QR</emphasis>
),
<taxonomicName id="2EA74D6E9310FF9CFDC3F140FD1B1805" authority="Evsyukov &amp; Golovatch &amp; Reip, 2018" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[601,741,1560,1583]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="dagestanus" status="sp. nov.">
<emphasis id="DBD3EAFF9310FF9CFDC3F140FD1B1805" box="[601,741,1560,1583]" italics="true" pageId="6" pageNumber="95">J. dagestanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0E057849310FF9CFD76F142FCD91805" box="[748,807,1560,1582]" pageId="6" pageNumber="95" rank="species">
<emphasis id="DBD3EAFF9310FF9CFD76F142FCD91805" bold="true" box="[748,807,1560,1582]" pageId="6" pageNumber="95">sp. n.</emphasis>
</taxonomicNameLabel>
(
<emphasis id="DBD3EAFF9310FF9CFCADF142FCBB1805" bold="true" box="[823,837,1560,1582]" pageId="6" pageNumber="95">S</emphasis>
) and
<taxonomicName id="2EA74D6E9310FF9CFC1BF140FBFD1805" authority="Evsyukov &amp; Golovatch &amp; Reip, 2018" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[897,1027,1560,1583]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="khostensis" status="sp. nov.">
<emphasis id="DBD3EAFF9310FF9CFC1BF140FBFD1805" box="[897,1027,1560,1583]" italics="true" pageId="6" pageNumber="95">J. khostensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0E057849310FF9CFB90F142FBBB1805" box="[1034,1093,1560,1582]" pageId="6" pageNumber="95" rank="species">
<emphasis id="DBD3EAFF9310FF9CFB90F142FBBB1805" bold="true" box="[1034,1093,1560,1582]" pageId="6" pageNumber="95">sp. n.</emphasis>
</taxonomicNameLabel>
(
<emphasis id="DBD3EAFF9310FF9CFBCFF142FB981805" bold="true" box="[1109,1126,1560,1582]" pageId="6" pageNumber="95">T</emphasis>
).
<emphasis id="DBD3EAFF9310FF9CFBE6F142FB691805" bold="true" box="[1148,1175,1560,1582]" pageId="6" pageNumber="95">A:</emphasis>
from the Rostov-on-Don Region.
<emphasis id="DBD3EAFF9310FF9CFF69F16DFEF51866" bold="true" box="[243,267,1591,1613]" pageId="6" pageNumber="95">B:</emphasis>
from Krasnaya Polyana, Krasnodar Province.
<emphasis id="DBD3EAFF9310FF9CFC93F16DFCDA1866" bold="true" box="[777,804,1591,1613]" pageId="6" pageNumber="95">C:</emphasis>
from Sakhrai, Adygea Republic.
<emphasis id="DBD3EAFF9310FF9CFB02F16DFB4C1866" bold="true" box="[1176,1202,1591,1613]" pageId="6" pageNumber="95">D:</emphasis>
syntype from Gagry, Abkhazia, after Lohmander (1936).
<emphasis id="DBD3EAFF9310FF9CFD93F10CFDDC1847" bold="true" box="[521,546,1622,1644]" pageId="6" pageNumber="95">E:</emphasis>
from North Ossetia.
<emphasis id="DBD3EAFF9310FF9CFD67F10CFCEB1847" bold="true" box="[765,789,1622,1644]" pageId="6" pageNumber="95">F:</emphasis>
paratype from North Ossetia, after Golovatch (1981).
<emphasis id="DBD3EAFF9310FF9CFADDF10CFA9D1847" bold="true" box="[1351,1379,1622,1644]" pageId="6" pageNumber="95">G:</emphasis>
from Maikop, Adygea Republic.
<emphasis id="DBD3EAFF9310FF9CFE2DF12EFE2D18A1" bold="true" box="[439,467,1652,1674]" pageId="6" pageNumber="95">H:</emphasis>
from Pskhu, Abkhazia.
<emphasis id="DBD3EAFF9310FF9CFD4EF12EFD1818A1" bold="true" box="[724,742,1652,1674]" pageId="6" pageNumber="95">I:</emphasis>
from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936).
<emphasis id="DBD3EAFF9310FF9CFA1DF12EFA6218A1" bold="true" box="[1415,1436,1652,1674]" pageId="6" pageNumber="95">J:</emphasis>
from Krasnaya Polyana, Krasnodar Province.
<emphasis id="DBD3EAFF9310FF9CFDE9F1C9FD711882" bold="true" box="[627,655,1683,1705]" pageId="6" pageNumber="95">K:</emphasis>
from Teberda, Karachaevo-Cherkessia.
<emphasis id="DBD3EAFF9310FF9CFBAAF1C9FBB61882" bold="true" box="[1072,1096,1683,1705]" pageId="6" pageNumber="95">L:</emphasis>
from Mestia, Svanetia, Georgia.
<emphasis id="DBD3EAFF9310FF9CFF0DF1E8FF4918E3" bold="true" box="[151,183,1714,1736]" pageId="6" pageNumber="95">M:</emphasis>
from Teberda, Karachaevo-Cherkessia.
<emphasis id="DBD3EAFF9310FF9CFDFAF1E8FD8418E3" bold="true" box="[608,634,1714,1736]" pageId="6" pageNumber="95">N:</emphasis>
from Svanetia, Georgia.
<emphasis id="DBD3EAFF9310FF9CFC10F1E8FC5B18E3" bold="true" box="[906,933,1714,1736]" pageId="6" pageNumber="95">O:</emphasis>
from near Teberda, Karachaevo-Cherkessia.
<emphasis id="DBD3EAFF9310FF9CFA1EF1E8FA6218E3" bold="true" box="[1412,1436,1714,1736]" pageId="6" pageNumber="95">P:</emphasis>
holotype from Olginskoe, Abkhazia, after Lohmander (1936).
<emphasis id="DBD3EAFF9310FF9CFCA6F18AFCA918CD" bold="true" box="[828,855,1744,1766]" pageId="6" pageNumber="95">Q:</emphasis>
from North Ossetia.
<emphasis id="DBD3EAFF9310FF9CFBDDF18AFB9F18CD" bold="true" box="[1095,1121,1744,1766]" pageId="6" pageNumber="95">R:</emphasis>
syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936).
<emphasis id="DBD3EAFF9310FF9CFDF5F1B5FD7B192E" bold="true" box="[623,645,1775,1797]" pageId="6" pageNumber="95">S:</emphasis>
holotype from Dagestan.
<emphasis id="DBD3EAFF9310FF9CFC14F1B5FC5A192E" bold="true" box="[910,932,1775,1797]" pageId="6" pageNumber="95">T:</emphasis>
holotype from Khosta, Krasnodar Province. Oral views
<emphasis id="DBD3EAFF9310FF9CFF42F054FEF7190F" bold="true" box="[216,265,1806,1828]" pageId="6" pageNumber="95">BO</emphasis>
; caudal views
<emphasis id="DBD3EAFF9310FF9CFE05F054FE34190F" bold="true" box="[415,458,1806,1828]" pageId="6" pageNumber="95">A, P</emphasis>
.
<emphasis id="DBD3EAFF9310FF9CFE4CF054FD91190F" bold="true" box="[470,623,1806,1828]" pageId="6" pageNumber="95">Abbreviations</emphasis>
:
<emphasis id="DBD3EAFF9310FF9CFDE6F054FD64190F" bold="true" box="[636,666,1806,1828]" pageId="6" pageNumber="95">cx,</emphasis>
coxa;
<emphasis id="DBD3EAFF9310FF9CFD47F054FD09190F" bold="true" box="[733,759,1806,1828]" pageId="6" pageNumber="95">te,</emphasis>
telopodite remnant. Scale bars: 0.3 mm (
<emphasis id="DBD3EAFF9310FF9CFB0DF054FF561969" bold="true" pageId="6" pageNumber="95">AC, E, G, H, JO, Q, S, T</emphasis>
), 0.2 mm (
<emphasis id="DBD3EAFF9310FF9CFE83F076FED61969" bold="true" box="[281,296,1836,1858]" pageId="6" pageNumber="95">F</emphasis>
), not to scale (
<emphasis id="DBD3EAFF9310FF9CFE25F076FDDF1969" bold="true" box="[447,545,1836,1858]" pageId="6" pageNumber="95">D, I, P, R</emphasis>
).
</paragraph>
</caption>
<paragraph id="E91836ED9310FF9DFF5DF035FD6D1DAB" blockId="6.[151,1436,1903,2036]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="96" pageId="6" pageNumber="95">
5 males, 2 females, 1 juvenile (ZMUM),
<emphasis id="DBD3EAFF9310FF9CFD01F035FCFC19A3" bold="true" box="[667,770,1903,1928]" pageId="6" pageNumber="95">Georgia,</emphasis>
Borzhomi Nature Reserve, Banishkevi Valley,
<quantity id="2E5F9B089310FF9CFA86F02AFA6219AC" box="[1308,1436,1904,1928]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="6" pageNumber="95" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800900 m</quantity>
a.s.l.,
<emphasis id="DBD3EAFF9310FF9CFF43F0CEFE931980" box="[217,365,1940,1963]" italics="true" pageId="6" pageNumber="95">
<taxonomicName id="2EA74D6E9310FF9CFF43F0CEFEE51980" box="[217,283,1940,1963]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">Picea</taxonomicName>
,
<taxonomicName id="2EA74D6E9310FF9CFEBDF0CEFE931980" box="[295,365,1940,1963]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
</emphasis>
and
<taxonomicName id="2EA74D6E9310FF9CFE39F0CEFDF21980" box="[419,524,1940,1963]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="95" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9310FF9CFE39F0CEFDF21980" box="[419,524,1940,1963]" italics="true" pageId="6" pageNumber="95">Carpinus</emphasis>
</taxonomicName>
forest, litter, logs, under stones,
<date id="9D19102D9310FF9CFCE0F0CEFBBC1987" box="[890,1090,1939,1964]" pageId="6" pageNumber="95" value="1983-05-12" valueMax="1983-05-16" valueMin="1983-05-12">12 and 16.V.1983</date>
; 1 male, 8 females, 3 juveniles (ZMUM), Ajaria, Khulo Distr.,
<quantity id="2E5F9B089310FF9CFD9DF0E2FDBD19FB" box="[519,579,1975,2000]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="95" unit="km" value="3.0">3 km</quantity>
W of Danisparauli, deciduous forest, litter,
<date id="9D19102D9310FF9CFBDFF0E2FB4119FB" box="[1093,1215,1976,2000]" pageId="6" pageNumber="95" value="1981-10-10">10.X.1981</date>
; 1 male, 3 females (ZMUM), Ajaria, Batumi Botanical Garden, litter,
<date id="9D19102D9310FF9CFD4CF086FCB119DF" box="[726,847,2012,2036]" pageId="6" pageNumber="95" value="1978-10-13">13.X.1978</date>
; all leg S. Golovatch; 2 males, 1 juvenile (ZMUM), Ajaria, Kintrish Nature Reserve, Zeraboseli,
<quantity id="2E5F9B089311FF9DFD02F7CDFCE71E84" box="[664,793,151,176]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="7" pageNumber="96" unit="m" value="525.0" valueMax="600.0" valueMin="450.0">450600 m</quantity>
a.s.l.,
<date id="9D19102D9311FF9DFCFDF7C2FC081E9B" box="[871,1014,152,176]" pageId="7" pageNumber="96" value="1981-06-01" valueMax="1981-06-03" valueMin="1981-06-01">13.VI.1981</date>
, leg. S. Golovatch and J. Martens; 1 male (SMNG), Khashuri, Akhaldaba, Nedzvi Managemened Reserve, Nedzvistskali River, 41.9082°N, 43.5151°E, humid
<taxonomicName id="2EA74D6E9311FF9DFF73F7BBFED71ED3" box="[233,297,225,248]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9311FF9DFF73F7BBFED71ED3" box="[233,297,225,248]" italics="true" pageId="7" pageNumber="96">Picea</emphasis>
</taxonomicName>
forest,
<date id="9D19102D9311FF9DFE1FF7BAFDF91ED3" box="[389,519,224,248]" pageId="7" pageNumber="96" value="2011-06-01">01.VI.2011</date>
; 1 male (SMNG), Akhaltsikhe to Borjomi, S Chobiskhevi, Chobiskhevi River, 41.7651°N, 43.3265°E,
<taxonomicName id="2EA74D6E9311FF9DFE32F65FFE101F37" box="[424,494,261,284]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="genus">
<emphasis id="DBD3EAFF9311FF9DFE32F65FFE101F37" box="[424,494,261,284]" italics="true" pageId="7" pageNumber="96">Fagus</emphasis>
</taxonomicName>
forest,
<date id="9D19102D9311FF9DFDDCF65EFD361F37" box="[582,712,260,284]" pageId="7" pageNumber="96" value="2011-06-01">01.VI.2011</date>
; 1 male (SMNG), Sairme Gorge, 41.9569°N 42.7692°E, mixed forest, leaf litter (
<taxonomicName id="2EA74D6E9311FF9DFEFCF673FDDC1F6B" authorityName="Lipsky" authorityYear="1898" box="[358,546,295,320]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="DBD3EAFF9311FF9DFEFCF673FDDC1F6B" box="[358,546,295,320]" italics="true" pageId="7" pageNumber="96">Fagus orientalis</emphasis>
</taxonomicName>
and
<taxonomicName id="2EA74D6E9311FF9DFDFAF672FCE61F6B" box="[608,792,296,320]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="species" species="sativa">
<emphasis id="DBD3EAFF9311FF9DFDFAF672FCE61F6B" box="[608,792,296,320]" italics="true" pageId="7" pageNumber="96">Castanea sativa</emphasis>
</taxonomicName>
),
<date id="9D19102D9311FF9DFCA9F672FC401F6B" box="[819,958,296,320]" pageId="7" pageNumber="96" value="2013-07-14">14.VII.2013</date>
, all leg. L. Mumladze; 1 male, 1 female (SMNG), Samzche-Dschawachetien, Abastumani Observatory on Mt Kanobili, 41.7544°N, 42.8202°E,
<quantity id="2E5F9B089311FF9DFADEF616FA621F4F" box="[1348,1436,332,357]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="7" pageNumber="96" unit="m" value="1580.0">1580 m</quantity>
a.s.l.,
<date id="9D19102D9311FF9DFF42F62AFEAC1FA3" box="[216,338,368,392]" pageId="7" pageNumber="96" value="2009-10-28">28.X.2009</date>
; 1 male, 3 females, 1 juvenile (SMNG), Shida Kartli, Ormotsi, Tana valley
<quantity id="2E5F9B089311FF9DFB06F635FB091FA3" box="[1180,1271,367,392]" metricMagnitude="4" metricUnit="m" metricValue="1.66" pageId="7" pageNumber="96" unit="km" value="16.6">16.6 km</quantity>
upstream Didi Ateni, 41.8611°N, 43.9444°E,
<quantity id="2E5F9B089311FF9DFD9AF6CEFDA61F87" box="[512,600,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="7" pageNumber="96" unit="m" value="1110.0">1110 m</quantity>
a.s.l.,
<date id="9D19102D9311FF9DFD36F6CEFCD31F86" box="[684,813,404,429]" pageId="7" pageNumber="96" value="2011-09-16">16.IX.2011</date>
; 1 male (SMNG), Samtskhe-Javakheti,
<quantity id="2E5F9B089311FF9DFA9CF6CEFAAC1F87" box="[1286,1362,404,429]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="96" unit="m" value="500.0">500 m</quantity>
W of Tsaghveri, Gujarula River valley, 41.8078°N, 43.4672°E,
<quantity id="2E5F9B089311FF9DFCA1F6E2FC6A1FE4" box="[827,916,440,464]" metricMagnitude="3" metricUnit="m" metricValue="1.01" pageId="7" pageNumber="96" unit="m" value="1010.0">1010 m</quantity>
a.s.l.,
<date id="9D19102D9311FF9DFC7CF6E2FB961FFB" box="[998,1128,440,464]" pageId="7" pageNumber="96" value="2011-09-17">17.IX.2011</date>
; 1 male (SMNG), Kvemo Kartli, SW of Kojori, 41.6472°N, 44.6936°E,
<quantity id="2E5F9B089311FF9DFD04F686FD0D1FDF" box="[670,755,476,500]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="96" unit="m" value="1200.0">1200 m</quantity>
a.s.l.,
<date id="9D19102D9311FF9DFCA4F686FC481FDE" box="[830,950,476,501]" pageId="7" pageNumber="96" value="2011-10-06">06.X.2011</date>
; 1 male (SMNG), Guria, SW of Kvabgha, 41.9131°N, 42.4000°E,
<quantity id="2E5F9B089311FF9DFE32F6A5FE0E1C3C" box="[424,496,511,536]" metricMagnitude="2" metricUnit="m" metricValue="5.9" pageId="7" pageNumber="96" unit="m" value="590.0">590 m</quantity>
a.s.l.,
<date id="9D19102D9311FF9DFDA6F6A5FD401C33" box="[572,702,511,536]" pageId="7" pageNumber="96" value="2011-09-26">26.IX.2011</date>
; 1 male (SMNG), Ajaria, Qorolistavi towards Mt Mta Mtirala, 41.6494°N, 41.7731°E,
<quantity id="2E5F9B089311FF9DFE3FF57EFE131C17" box="[421,493,548,572]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="96" unit="m" value="730.0">730 m</quantity>
a.s.l.,
<date id="9D19102D9311FF9DFDACF57EFD4E1C17" box="[566,688,548,572]" pageId="7" pageNumber="96" value="2012-10-01">01.X.2012</date>
; 1 female, 1 juvenile (SMNG), Mtskheta-Mtianeti, Darial gorge at
<quantity id="2E5F9B089311FF9DFF0DF512FEF31C4B" box="[151,269,583,608]" metricMagnitude="4" metricUnit="m" metricValue="6.95" metricValueMax="13.1" metricValueMin="0.8" pageId="7" pageNumber="96" unit="km" value="69.5" valueMax="131.0" valueMin="8.0">8131 km</quantity>
, 42.7039°N, 44.6264°E,
<quantity id="2E5F9B089311FF9DFDB4F512FD781C74" box="[558,646,584,608]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="7" pageNumber="96" unit="m" value="1480.0">1480 m</quantity>
.a.s.l.,
<date id="9D19102D9311FF9DFD46F512FC8E1C4B" box="[732,880,584,608]" pageId="7" pageNumber="96" value="2014-08-10">10.VIII.2014</date>
, all leg. F. Walther; 1 male, 1 female (ZMUM), Tskhaltubo, near Cave Belaya (= Tetri), deciduous forest on limestone rock, litter and under stones,
<date id="9D19102D9311FF9DFA92F536FA7C1CAE" box="[1288,1410,620,645]" pageId="7" pageNumber="96" value="1981-10-26">26.X.1981</date>
; 1 male (ZMUM), Kutaisi Distr., Sataplia Nature Reserve near Tskhaltubo, forest, litter and under stones,
<date id="9D19102D9311FF9DFA85F5D5FA661C83" box="[1311,1432,655,680]" pageId="7" pageNumber="96" value="1981-10-25">25.X.1981</date>
, all leg. S. Golovatch; 1 male, 1 female (ZMUM), same locality,
<date id="9D19102D9311FF9DFC1BF5EEFC231CE7" box="[897,989,692,716]" pageId="7" pageNumber="96" value="1973-04">IV.1973</date>
, leg. D. Krivolutsky; 1 male, 1 female (ZMUM), Okriba, Mukhura
<emphasis id="DBD3EAFF9311FF9DFE43F583FE0D1CDB" box="[473,499,729,752]" italics="true" pageId="7" pageNumber="96">ca</emphasis>
<quantity id="2E5F9B089311FF9DFE60F58DFDBF1CDB" box="[506,577,727,752]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="96" unit="km" value="15.0">15 km</quantity>
E of Tkibuli,
<quantity id="2E5F9B089311FF9DFD7AF582FCA11CC4" box="[736,863,728,752]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="7" pageNumber="96" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">700800 m</quantity>
a.s.l.,
<emphasis id="DBD3EAFF9311FF9DFC33F582FB231CC4" box="[937,1245,728,752]" italics="true" pageId="7" pageNumber="96">
<taxonomicName id="2EA74D6E9311FF9DFC33F582FBEB1CC4" box="[937,1045,728,751]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="genus">Castanea</taxonomicName>
,
<taxonomicName id="2EA74D6E9311FF9DFBBBF583FB941CDB" box="[1057,1130,729,752]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="2EA74D6E9311FF9DFBEFF582FB231CC4" box="[1141,1245,728,751]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="genus">Carpinus</taxonomicName>
</emphasis>
etc. forest, litter, under bark and stones,
<date id="9D19102D9311FF9DFE31F5A6FDD01D3F" box="[427,558,764,788]" pageId="7" pageNumber="96" value="1987-05-07" valueMax="1987-05-09" valueMin="1987-05-07">79.V.1987</date>
; 4 males, 8 females, 4 juveniles (ZMUM), Surami Mt. Ridge, Pass Djvari between Gomi and Sachkhere,
<quantity id="2E5F9B089311FF9DFD92F445FDAB1D1C" box="[520,597,799,824]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="96" unit="m" value="850.0">850 m</quantity>
a.s.l.,
<emphasis id="DBD3EAFF9311FF9DFD32F445FC071D13" box="[680,1017,799,824]" italics="true" pageId="7" pageNumber="96">
<taxonomicName id="2EA74D6E9311FF9DFD32F445FD151D13" box="[680,747,799,824]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="genus">Alnus</taxonomicName>
,
<taxonomicName id="2EA74D6E9311FF9DFD61F47BFCBA1D13" box="[763,836,801,824]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="2EA74D6E9311FF9DFCCEF445FC071D13" box="[852,1017,799,824]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="7" pageNumber="96" phylum="Tracheophyta" rank="genus">Rhododendron</taxonomicName>
</emphasis>
etc. forest, litter,
<date id="9D19102D9311FF9DFB49F47AFAC41D13" box="[1235,1338,800,824]" pageId="7" pageNumber="96" value="1987-05-07">7.V.1987</date>
; 1 male (ZMUM), Kakhetia, Babaneuri Nature Reserve,
<emphasis id="DBD3EAFF9311FF9DFD51F41FFD1B1D77" box="[715,741,837,860]" italics="true" pageId="7" pageNumber="96">ca</emphasis>
<quantity id="2E5F9B089311FF9DFD6BF41EFCC21D76" box="[753,828,836,861]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="7" pageNumber="96" unit="km" value="16.0">16 km</quantity>
ENE of Akhmeta, near Babaneuri,
<quantity id="2E5F9B089311FF9DFB7EF41EFAD11D77" box="[1252,1327,836,861]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="96" unit="m" value="500.0">500 m</quantity>
a.s.l.,
<date id="9D19102D9311FF9DFA1AF41EFF001DAB" pageId="7" pageNumber="96" value="1987-05-04" valueMax="1987-05-05" valueMin="1987-05-04">4 5.V.1987</date>
, all leg. S. Golovatch and K. Eskov.
</paragraph>
</subSubSection>
<subSubSection id="A1BD65669311FF9DFF5DF4D6FA73195B" pageId="7" pageNumber="96" type="diagnosis">
<paragraph id="E91836ED9311FF9DFF5DF4D6FC4E1A57" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="96">
<emphasis id="DBD3EAFF9311FF9DFF5DF4D6FEBF1D8E" bold="true" box="[199,321,908,933]" pageId="7" pageNumber="96">Diagnosis.</emphasis>
Differs from other species of the genus by the following combination of characters. Main process (
<emphasis id="DBD3EAFF9311FF9DFF3AF4F5FF391DE3" bold="true" box="[160,199,943,968]" pageId="7" pageNumber="96">mp</emphasis>
) of male leg-pair 2 long, with a ventral spike-like outgrowth (
<figureCitation id="719C2A689311FF9DFC11F4F5FBE91DE3" box="[907,1047,943,968]" captionStart="FIGURE 2" captionStartId="5.[151,249,1601,1623]" captionTargetBox="[340,1248,985,1580]" captionTargetId="figure@5.[340,1248,985,1580]" captionTargetPageId="5" captionText="FIGURE 2. Gnathochilaria in Caucasian species of Julus, ventral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="7" pageNumber="96">Figs 2B, 2C</figureCitation>
,
<figureCitation id="719C2A689311FF9DFBBEF4EAFBB91DE3" box="[1060,1095,944,968]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF), J. kubanus Verhoeff, 1921 (GI), J. lignaui Verhoeff, 1910 (JM), J. lindholmi Lohmander, 1936 (N P), J. subalpinus Lohmander, 1936 (QR), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views BO; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (AC, E, G, H, JO, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="7" pageNumber="96">3B</figureCitation>
,
<figureCitation id="719C2A689311FF9DFBCEF4EAFB881DE3" box="[1108,1142,944,968]" captionStart="FIGURE 4" captionStartId="8.[151,250,1477,1499]" captionTargetBox="[340,1248,375,1442]" captionTargetId="figure@8.[340,1248,375,1455]" captionTargetPageId="8" captionText="FIGURE 4. Male legs 2 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF) and J. kubanus Verhoeff, 1921 (GI). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). Oral views C, E, H; caudal views A, B, D, F, G; mesal view I. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (AC, E, G, H), 0.2 mm (F), not to scale (D, I)." httpUri="https://zenodo.org/record/1459880/files/figure.png" pageId="7" pageNumber="96">4B</figureCitation>
,
<figureCitation id="719C2A689311FF9DFB1EF4F5FB591DE3" box="[1156,1191,943,968]" captionStart="FIGURE 5" captionStartId="9.[151,250,1527,1549]" captionTargetBox="[340,1247,1093,1506]" captionTargetId="figure@9.[340,1247,1093,1506]" captionTargetPageId="9" captionText="FIGURE 5. Julus colchicus Lohmander, 1936 from Krasnaya Polyana, Krasnodar Province. A: habitus of female (left) and male (right), lateral views. B and C: anterior part of male body, lateral and ventral views, respectively. D: midbody segments, lateral view. E: caudal part of body, lateral view. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459882/files/figure.png" pageId="7" pageNumber="96">5B</figureCitation>
). Ends of flagella (
<emphasis id="DBD3EAFF9311FF9DFA1BF4F5FA6C1DE3" bold="true" box="[1409,1426,943,968]" pageId="7" pageNumber="96">fl</emphasis>
) upturned (
<figureCitation id="719C2A689311FF9DFE93F48EFE6B1DC7" box="[265,405,980,1005]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF), J. kubanus Verhoeff, 1921 (GI), J. lignaui Verhoeff, 1910 (JM), J. lindholmi Lohmander, 1936 (N P), J. subalpinus Lohmander, 1936 (QR), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views BO; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (AC, E, G, H, JO, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="7" pageNumber="96">Figs 3C, 3D</figureCitation>
,
<figureCitation id="719C2A689311FF9DFE3BF48EFE381DC7" box="[417,454,980,1004]" captionStart="FIGURE 4" captionStartId="8.[151,250,1477,1499]" captionTargetBox="[340,1248,375,1442]" captionTargetId="figure@8.[340,1248,375,1455]" captionTargetPageId="8" captionText="FIGURE 4. Male legs 2 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF) and J. kubanus Verhoeff, 1921 (GI). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). Oral views C, E, H; caudal views A, B, D, F, G; mesal view I. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (AC, E, G, H), 0.2 mm (F), not to scale (D, I)." httpUri="https://zenodo.org/record/1459880/files/figure.png" pageId="7" pageNumber="96">4D</figureCitation>
,
<figureCitation id="719C2A689311FF9DFE4BF48EFE0A1DC6" box="[465,500,980,1005]" captionStart="FIGURE 5" captionStartId="9.[151,250,1527,1549]" captionTargetBox="[340,1247,1093,1506]" captionTargetId="figure@9.[340,1247,1093,1506]" captionTargetPageId="9" captionText="FIGURE 5. Julus colchicus Lohmander, 1936 from Krasnaya Polyana, Krasnodar Province. A: habitus of female (left) and male (right), lateral views. B and C: anterior part of male body, lateral and ventral views, respectively. D: midbody segments, lateral view. E: caudal part of body, lateral view. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459882/files/figure.png" pageId="7" pageNumber="96">5E</figureCitation>
). Distal part of
<emphasis id="DBD3EAFF9311FF9DFD39F48EFD4A1DC6" bold="true" box="[675,692,980,1005]" pageId="7" pageNumber="96">fl</emphasis>
thin, with lamellae. Gonopodal promere (
<emphasis id="DBD3EAFF9311FF9DFB16F48EFB561DC6" bold="true" box="[1164,1192,980,1005]" pageId="7" pageNumber="96">pr</emphasis>
) wide, round at apex, with a triangular spike (
<emphasis id="DBD3EAFF9311FF9DFE3FF4ADFE501A3B" bold="true" box="[421,430,1015,1040]" pageId="7" pageNumber="96">t</emphasis>
) in middle part (
<figureCitation id="719C2A689311FF9DFDF4F4ADFD021A3B" box="[622,764,1015,1040]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF), J. kubanus Verhoeff, 1921 (GI), J. lignaui Verhoeff, 1910 (JM), J. lindholmi Lohmander, 1936 (N P), J. subalpinus Lohmander, 1936 (QR), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views BO; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (AC, E, G, H, JO, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="7" pageNumber="96">Figs 3C, 3D</figureCitation>
,
<figureCitation id="719C2A689311FF9DFC92F4A2FCA31A3B" box="[776,861,1016,1040]" captionStart="FIGURE 4" captionStartId="8.[151,250,1477,1499]" captionTargetBox="[340,1248,375,1442]" captionTargetId="figure@8.[340,1248,375,1455]" captionTargetPageId="8" captionText="FIGURE 4. Male legs 2 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF) and J. kubanus Verhoeff, 1921 (GI). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). Oral views C, E, H; caudal views A, B, D, F, G; mesal view I. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (AC, E, G, H), 0.2 mm (F), not to scale (D, I)." httpUri="https://zenodo.org/record/1459880/files/figure.png" pageId="7" pageNumber="96">4C, 4D</figureCitation>
,
<figureCitation id="719C2A689311FF9DFCF3F4ADFC131A3B" box="[873,1005,1015,1040]" captionStart="FIGURE 5" captionStartId="9.[151,250,1527,1549]" captionTargetBox="[340,1247,1093,1506]" captionTargetId="figure@9.[340,1247,1093,1506]" captionTargetPageId="9" captionText="FIGURE 5. Julus colchicus Lohmander, 1936 from Krasnaya Polyana, Krasnodar Province. A: habitus of female (left) and male (right), lateral views. B and C: anterior part of male body, lateral and ventral views, respectively. D: midbody segments, lateral view. E: caudal part of body, lateral view. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459882/files/figure.png" pageId="7" pageNumber="96">5C, 5D, 5E</figureCitation>
). Lateral outgrowth (
<emphasis id="DBD3EAFF9311FF9DFB46F4ADFB0C1A3B" bold="true" box="[1244,1266,1015,1040]" pageId="7" pageNumber="96">lo</emphasis>
) of
<emphasis id="DBD3EAFF9311FF9DFAB8F4ADFAC11A3B" bold="true" box="[1314,1343,1015,1040]" pageId="7" pageNumber="96">op</emphasis>
narrow, pointed (
<figureCitation id="719C2A689311FF9DFF60F346FEA81A1F" box="[250,342,1052,1077]" captionStart="FIGURE 4" captionStartId="8.[151,250,1477,1499]" captionTargetBox="[340,1248,375,1442]" captionTargetId="figure@8.[340,1248,375,1455]" captionTargetPageId="8" captionText="FIGURE 4. Male legs 2 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF) and J. kubanus Verhoeff, 1921 (GI). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). Oral views C, E, H; caudal views A, B, D, F, G; mesal view I. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (AC, E, G, H), 0.2 mm (F), not to scale (D, I)." httpUri="https://zenodo.org/record/1459880/files/figure.png" pageId="7" pageNumber="96">Figs 4E</figureCitation>
,
<figureCitation id="719C2A689311FF9DFEF8F346FE7C1A1E" box="[354,386,1052,1077]" captionStart="FIGURE 5" captionStartId="9.[151,250,1527,1549]" captionTargetBox="[340,1247,1093,1506]" captionTargetId="figure@9.[340,1247,1093,1506]" captionTargetPageId="9" captionText="FIGURE 5. Julus colchicus Lohmander, 1936 from Krasnaya Polyana, Krasnodar Province. A: habitus of female (left) and male (right), lateral views. B and C: anterior part of male body, lateral and ventral views, respectively. D: midbody segments, lateral view. E: caudal part of body, lateral view. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459882/files/figure.png" pageId="7" pageNumber="96">5E</figureCitation>
) or rounded (
<figureCitation id="719C2A689311FF9DFD87F346FD851A1F" box="[541,635,1052,1077]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF), J. kubanus Verhoeff, 1921 (GI), J. lignaui Verhoeff, 1910 (JM), J. lindholmi Lohmander, 1936 (N P), J. subalpinus Lohmander, 1936 (QR), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views BO; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (AC, E, G, H, JO, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="7" pageNumber="96">Figs 3D</figureCitation>
,
<figureCitation id="719C2A689311FF9DFD12F346FD551A1E" box="[648,683,1052,1077]" captionStart="FIGURE 5" captionStartId="9.[151,250,1527,1549]" captionTargetBox="[340,1247,1093,1506]" captionTargetId="figure@9.[340,1247,1093,1506]" captionTargetPageId="9" captionText="FIGURE 5. Julus colchicus Lohmander, 1936 from Krasnaya Polyana, Krasnodar Province. A: habitus of female (left) and male (right), lateral views. B and C: anterior part of male body, lateral and ventral views, respectively. D: midbody segments, lateral view. E: caudal part of body, lateral view. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459882/files/figure.png" pageId="7" pageNumber="96">5D</figureCitation>
) at apex. Metazonae with shallow and weakly expressed striations (
<figureCitation id="719C2A689311FF9DFF05F365FF031A73" box="[159,253,1087,1112]" captionStart="FIGURE 2" captionStartId="5.[151,249,1601,1623]" captionTargetBox="[340,1248,985,1580]" captionTargetId="figure@5.[340,1248,985,1580]" captionTargetPageId="5" captionText="FIGURE 2. Gnathochilaria in Caucasian species of Julus, ventral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="7" pageNumber="96">Figs 2D</figureCitation>
, 22B). Female vulvae (
<emphasis id="DBD3EAFF9311FF9DFD92F365FDDB1A73" bold="true" box="[520,549,1087,1112]" pageId="7" pageNumber="96">vu</emphasis>
) relatively broad; ampulla 1 (
<emphasis id="DBD3EAFF9311FF9DFCE1F365FC511A73" bold="true" box="[891,943,1087,1112]" pageId="7" pageNumber="96">am1</emphasis>
) located slightly basal to ampulla 2 (
<emphasis id="DBD3EAFF9311FF9DFACDF365FA741A73" bold="true" box="[1367,1418,1087,1112]" pageId="7" pageNumber="96">am2</emphasis>
); coxa 2 (
<emphasis id="DBD3EAFF9311FF9DFF6BF33EFEF81A56" bold="true" box="[241,262,1124,1149]" pageId="7" pageNumber="96">l2</emphasis>
) with a weakly developed apicomesal outgrowth (Fig. 23B).
</paragraph>
<paragraph id="E91836ED9311FF9DFF5DF3DDFCF81B26" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="96">
<emphasis id="DBD3EAFF9311FF9DFF5DF3DDFE611A8B" bold="true" box="[199,415,1159,1184]" pageId="7" pageNumber="96">Descriptive notes.</emphasis>
Length of adults
<quantity id="2E5F9B089311FF9DFDE3F3DDFD041A8B" box="[633,762,1159,1184]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.6" metricValueMin="1.9" pageId="7" pageNumber="96" unit="mm" value="27.5" valueMax="36.0" valueMin="19.0">1936 mm</quantity>
, width
<quantity id="2E5F9B089311FF9DFCCCF3DDFC1B1A8B" box="[854,997,1159,1184]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.5" metricValueMin="1.3" pageId="7" pageNumber="96" unit="mm" value="1.9" valueMax="2.5" valueMin="1.3">1.32.5 mm</quantity>
. Number of segments in adults from 44+1+T to 50+1+T. Body grey-brown with a thin, black, dorsal, axial line, venter lighter; legs light brown to yellow; antennae and anal valves brown (
<figureCitation id="719C2A689311FF9DFD0BF395FD231AC3" box="[657,733,1231,1256]" captionStart="FIGURE 5" captionStartId="9.[151,250,1527,1549]" captionTargetBox="[340,1247,1093,1506]" captionTargetId="figure@9.[340,1247,1093,1506]" captionTargetPageId="9" captionText="FIGURE 5. Julus colchicus Lohmander, 1936 from Krasnaya Polyana, Krasnodar Province. A: habitus of female (left) and male (right), lateral views. B and C: anterior part of male body, lateral and ventral views, respectively. D: midbody segments, lateral view. E: caudal part of body, lateral view. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459882/files/figure.png" pageId="7" pageNumber="96">Fig. 5</figureCitation>
). Eye patches composed of 4045 ocelli. Metazonae with superficial, weakly expressed striations (
<figureCitation id="719C2A689311FF9DFDC4F3AEFD451B26" box="[606,699,1268,1293]" captionStart="FIGURE 5" captionStartId="9.[151,250,1527,1549]" captionTargetBox="[340,1247,1093,1506]" captionTargetId="figure@9.[340,1247,1093,1506]" captionTargetPageId="9" captionText="FIGURE 5. Julus colchicus Lohmander, 1936 from Krasnaya Polyana, Krasnodar Province. A: habitus of female (left) and male (right), lateral views. B and C: anterior part of male body, lateral and ventral views, respectively. D: midbody segments, lateral view. E: caudal part of body, lateral view. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459882/files/figure.png" pageId="7" pageNumber="96">Figs 5D</figureCitation>
,
<figureCitation id="719C2A689311FF9DFD5DF3AEFD071B26" box="[711,761,1268,1293]" captionStart="FIGURE 16" captionStartId="21.[151,249,1286,1308]" captionTargetBox="[340,1247,583,1264]" captionTargetId="figure@21.[340,1247,583,1264]" captionTargetPageId="21" captionText="FIGURE 16. Striation on metazonites in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Pictures taken not to scale." httpUri="https://zenodo.org/record/1459904/files/figure.png" pageId="7" pageNumber="96">16B</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9311FF9DFF5DF24DFCD71B53" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="96">
All other characters as in
<taxonomicName id="2EA74D6E9311FF9DFE72F243FD7F1B1B" authorityName="Evsyukov" authorityYear="2016" box="[488,641,1303,1328]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="alexandrae">
<emphasis id="DBD3EAFF9311FF9DFE72F243FD7F1B1B" box="[488,641,1303,1328]" italics="true" pageId="7" pageNumber="96">J. alexandrae</emphasis>
</taxonomicName>
, except as follows. Antennae
<emphasis id="DBD3EAFF9311FF9DFC4EF243FBE41B1B" box="[980,1050,1305,1328]" italics="true" pageId="7" pageNumber="96">in situ</emphasis>
reaching back to body segment 4. Antennomeres 5 and 6 each with a corolla of bacilliform sensilla (
<figureCitation id="719C2A689311FF9DFC19F266FC241B7F" box="[899,986,1340,1365]" captionStart="FIGURE 1" captionStartId="4.[151,249,1820,1842]" captionTargetBox="[344,1236,854,1775]" captionTargetId="figure@4.[340,1248,841,1798]" captionTargetPageId="4" captionText="FIGURE 1. Antennae in Caucasian species of Julus, lateral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. dagestanus sp. n. H: J. khostensis sp. n. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459874/files/figure.png" pageId="7" pageNumber="96">Fig. 1B</figureCitation>
). Each stipes of gnathochilarium with a group of a few setae parabasally, below mentum (
<figureCitation id="719C2A689311FF9DFD59F205FCE51B53" box="[707,795,1375,1400]" captionStart="FIGURE 2" captionStartId="5.[151,249,1601,1623]" captionTargetBox="[340,1248,985,1580]" captionTargetId="figure@5.[340,1248,985,1580]" captionTargetPageId="5" captionText="FIGURE 2. Gnathochilaria in Caucasian species of Julus, ventral views. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. dagestanus sp. n. I: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459876/files/figure.png" pageId="7" pageNumber="96">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9311FF9DFF5DF2DEFC791807" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="96">
Male leg-pair 1 (
<figureCitation id="719C2A689311FF9DFE10F2DEFDF71BB7" box="[394,521,1412,1437]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF), J. kubanus Verhoeff, 1921 (GI), J. lignaui Verhoeff, 1910 (JM), J. lindholmi Lohmander, 1936 (N P), J. subalpinus Lohmander, 1936 (QR), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views BO; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (AC, E, G, H, JO, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="7" pageNumber="96">Figs 3BD</figureCitation>
) with a well-developed suture between coxa (
<emphasis id="DBD3EAFF9311FF9DFB84F2DEFBC61BB6" bold="true" box="[1054,1080,1412,1437]" pageId="7" pageNumber="96">cx</emphasis>
) and a 1-segmented telopodite remnant (
<emphasis id="DBD3EAFF9311FF9DFE98F2FDFEE61BEB" bold="true" box="[258,280,1447,1472]" pageId="7" pageNumber="96">te</emphasis>
). Apex of lateral process of
<emphasis id="DBD3EAFF9311FF9DFDCEF2FDFD911BEB" bold="true" box="[596,623,1447,1472]" pageId="7" pageNumber="96">cx</emphasis>
varying from acuminate (
<figureCitation id="719C2A689311FF9DFC15F2FDFBE61BEB" box="[911,1048,1447,1472]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF), J. kubanus Verhoeff, 1921 (GI), J. lignaui Verhoeff, 1910 (JM), J. lindholmi Lohmander, 1936 (N P), J. subalpinus Lohmander, 1936 (QR), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views BO; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (AC, E, G, H, JO, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="7" pageNumber="96">Figs 3B, 3D</figureCitation>
) to round (
<figureCitation id="719C2A689311FF9DFB09F2FDFB151BEB" box="[1171,1259,1447,1472]" captionStart="FIGURE 3" captionStartId="6.[151,249,1499,1521]" captionTargetBox="[340,1248,481,1476]" captionTargetId="figure@6.[340,1248,481,1478]" captionTargetPageId="6" captionText="FIGURE 3. Male legs 1 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF), J. kubanus Verhoeff, 1921 (GI), J. lignaui Verhoeff, 1910 (JM), J. lindholmi Lohmander, 1936 (N P), J. subalpinus Lohmander, 1936 (QR), J. dagestanus sp. n. (S) and J. khostensis sp. n. (T). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). J: from Krasnaya Polyana, Krasnodar Province. K: from Teberda, Karachaevo-Cherkessia. L: from Mestia, Svanetia, Georgia. M: from Teberda, Karachaevo-Cherkessia. N: from Svanetia, Georgia. O: from near Teberda, Karachaevo-Cherkessia. P: holotype from Olginskoe, Abkhazia, after Lohmander (1936). Q: from North Ossetia. R: syntype from Mount Djuga, Krasnodar Province, after Lohmander (1936). S: holotype from Dagestan. T: holotype from Khosta, Krasnodar Province. Oral views BO; caudal views A, P. Abbreviations: cx, coxa; te, telopodite remnant. Scale bars: 0.3 mm (AC, E, G, H, JO, Q, S, T), 0.2 mm (F), not to scale (D, I, P, R)." httpUri="https://zenodo.org/record/1459878/files/figure.png" pageId="7" pageNumber="96">Fig. 3C</figureCitation>
);
<emphasis id="DBD3EAFF9311FF9DFB65F2FDFAEB1BEB" bold="true" box="[1279,1301,1447,1472]" pageId="7" pageNumber="96">te</emphasis>
round, with several thin setae. Male leg-pair 2 with three coxal processes (
<figureCitation id="719C2A689311FF9DFCFCF296FC171BCF" box="[870,1001,1484,1509]" captionStart="FIGURE 4" captionStartId="8.[151,250,1477,1499]" captionTargetBox="[340,1248,375,1442]" captionTargetId="figure@8.[340,1248,375,1455]" captionTargetPageId="8" captionText="FIGURE 4. Male legs 2 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF) and J. kubanus Verhoeff, 1921 (GI). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). Oral views C, E, H; caudal views A, B, D, F, G; mesal view I. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (AC, E, G, H), 0.2 mm (F), not to scale (D, I)." httpUri="https://zenodo.org/record/1459880/files/figure.png" pageId="7" pageNumber="96">Figs 4BD</figureCitation>
): a short anterolateral process (
<emphasis id="DBD3EAFF9311FF9DFACAF296FA8B1BCE" bold="true" box="[1360,1397,1484,1509]" pageId="7" pageNumber="96">alp</emphasis>
), a long, distally thickened main process (
<emphasis id="DBD3EAFF9311FF9DFDCDF2B5FD801823" bold="true" box="[599,638,1519,1544]" pageId="7" pageNumber="96">mp</emphasis>
) with a distocaudal spike-like outgrowth, and a short and acuminate posterior process (
<emphasis id="DBD3EAFF9311FF9DFEFFF14EFE7A1806" bold="true" box="[357,388,1556,1581]" pageId="7" pageNumber="96">pp</emphasis>
). Penes (
<emphasis id="DBD3EAFF9311FF9DFE74F14EFE031806" bold="true" box="[494,509,1556,1581]" pageId="7" pageNumber="96">p</emphasis>
) short, slightly bifurcate (
<figureCitation id="719C2A689311FF9DFCBBF14EFC841807" box="[801,890,1556,1581]" captionStart="FIGURE 4" captionStartId="8.[151,250,1477,1499]" captionTargetBox="[340,1248,375,1442]" captionTargetId="figure@8.[340,1248,375,1455]" captionTargetPageId="8" captionText="FIGURE 4. Male legs 2 in Julus alexandrae Evsyukov, 2016 (A), J. colchicus Lohmander, 1936 (BD), J. jedryczkowskii Golovatch, 1981 (EF) and J. kubanus Verhoeff, 1921 (GI). A: from the Rostov-on-Don Region. B: from Krasnaya Polyana, Krasnodar Province. C: from Sakhrai, Adygea Republic. D: syntype from Gagry, Abkhazia, after Lohmander (1936). E: from North Ossetia. F: paratype from North Ossetia, after Golovatch (1981). G: from Maikop, Adygea Republic. H: from Pskhu, Abkhazia. I: from Aksaut, Karachaevo-Cherkessia, after Lohmander (1936). Oral views C, E, H; caudal views A, B, D, F, G; mesal view I. Abbreviations: alp, anterolateral process of coxa 2; mp, main process of coxa 2; p, penes; pp, posterior process of coxa 2. Scale bars: 0.3 mm (AC, E, G, H), 0.2 mm (F), not to scale (D, I)." httpUri="https://zenodo.org/record/1459880/files/figure.png" pageId="7" pageNumber="96">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9311FF9DFF5DF162FEBC18BC" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="96">
Promere (
<emphasis id="DBD3EAFF9311FF9DFEADF16DFEAD187B" bold="true" box="[311,339,1591,1616]" pageId="7" pageNumber="96">pr</emphasis>
) broad, clearly rounded at apex, with a triangular mesal spike or tooth (
<emphasis id="DBD3EAFF9311FF9DFB16F16DFB6B187B" bold="true" box="[1164,1173,1591,1616]" pageId="7" pageNumber="96">t</emphasis>
) near distal third (
<figureCitation id="719C2A689311FF9DFAF0F16DFF2C185F" captionStart="FIGURE 8" captionStartId="12.[151,250,1726,1748]" captionTargetBox="[340,1248,625,1695]" captionTargetId="figure@12.[340,1248,625,1705]" captionTargetPageId="12" captionText="FIGURE 8. Male gonopods in Julus alexandrae Evsyukov, 2018 (AB from the Rostov-on-Don Region) and J. colchicus Lohmander, 1936 (CD from Krasnaya Polyana, Krasnodar Province, EF from Sakhrai, Adygea Republic, GH syntype from Gagry, Abkhazia, after Lohmander (1936), I from Sataplia State Reserve, Georgia, after Golovatch (1977)). A, E, G: promere, caudal view. B, D: left gonopods, mesal view. C: promere with flagella, caudal view. F: right gonopods, mesal view. H, I: left gonopods, mesal view. Scale bars: 0.3 mm. G, H, I not to scale. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere; t, mesal tooth on promere." httpUri="https://zenodo.org/record/1459888/files/figure.png" pageId="7" pageNumber="96">Figs 8CI</figureCitation>
), flagella (
<emphasis id="DBD3EAFF9311FF9DFECEF106FE9B185E" bold="true" box="[340,357,1628,1653]" pageId="7" pageNumber="96">fl</emphasis>
) long and mostly relatively slender, with inconspicuous lamellae over most of their extent, curved at apex.
</paragraph>
<paragraph id="E91836ED9311FF9DFF5DF1FFFB39192F" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="96">
Mesomere (
<emphasis id="DBD3EAFF9311FF9DFECEF1FEFE881896" bold="true" box="[340,374,1700,1725]" pageId="7" pageNumber="96">ms</emphasis>
) rounded at apex (
<figureCitation id="719C2A689311FF9DFDFAF1FEFCAB1897" box="[608,853,1700,1725]" captionStart="FIGURE 8" captionStartId="12.[151,250,1726,1748]" captionTargetBox="[340,1248,625,1695]" captionTargetId="figure@12.[340,1248,625,1705]" captionTargetPageId="12" captionText="FIGURE 8. Male gonopods in Julus alexandrae Evsyukov, 2018 (AB from the Rostov-on-Don Region) and J. colchicus Lohmander, 1936 (CD from Krasnaya Polyana, Krasnodar Province, EF from Sakhrai, Adygea Republic, GH syntype from Gagry, Abkhazia, after Lohmander (1936), I from Sataplia State Reserve, Georgia, after Golovatch (1977)). A, E, G: promere, caudal view. B, D: left gonopods, mesal view. C: promere with flagella, caudal view. F: right gonopods, mesal view. H, I: left gonopods, mesal view. Scale bars: 0.3 mm. G, H, I not to scale. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere; t, mesal tooth on promere." httpUri="https://zenodo.org/record/1459888/files/figure.png" pageId="7" pageNumber="96">
Figs 8D, 8F, 8H,
<date id="9D19102D9311FF9DFCA6F1FEFCAB1897" box="[828,853,1700,1724]" pageId="7" pageNumber="96">8I</date>
</figureCitation>
). Opisthomere (
<emphasis id="DBD3EAFF9311FF9DFB8DF1FEFBCA1896" bold="true" box="[1047,1076,1700,1725]" pageId="7" pageNumber="96">op</emphasis>
) somewhat variable in shape: flattened (
<figureCitation id="719C2A689311FF9DFE96F19DFE6518CB" box="[268,411,1735,1760]" captionStart="FIGURE 8" captionStartId="12.[151,250,1726,1748]" captionTargetBox="[340,1248,625,1695]" captionTargetId="figure@12.[340,1248,625,1705]" captionTargetPageId="12" captionText="FIGURE 8. Male gonopods in Julus alexandrae Evsyukov, 2018 (AB from the Rostov-on-Don Region) and J. colchicus Lohmander, 1936 (CD from Krasnaya Polyana, Krasnodar Province, EF from Sakhrai, Adygea Republic, GH syntype from Gagry, Abkhazia, after Lohmander (1936), I from Sataplia State Reserve, Georgia, after Golovatch (1977)). A, E, G: promere, caudal view. B, D: left gonopods, mesal view. C: promere with flagella, caudal view. F: right gonopods, mesal view. H, I: left gonopods, mesal view. Scale bars: 0.3 mm. G, H, I not to scale. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere; t, mesal tooth on promere." httpUri="https://zenodo.org/record/1459888/files/figure.png" pageId="7" pageNumber="96">Figs 8D, 8F</figureCitation>
) or widely rounded (
<figureCitation id="719C2A689311FF9DFD0CF19DFCE118CB" box="[662,799,1735,1760]" captionStart="FIGURE 8" captionStartId="12.[151,250,1726,1748]" captionTargetBox="[340,1248,625,1695]" captionTargetId="figure@12.[340,1248,625,1705]" captionTargetPageId="12" captionText="FIGURE 8. Male gonopods in Julus alexandrae Evsyukov, 2018 (AB from the Rostov-on-Don Region) and J. colchicus Lohmander, 1936 (CD from Krasnaya Polyana, Krasnodar Province, EF from Sakhrai, Adygea Republic, GH syntype from Gagry, Abkhazia, after Lohmander (1936), I from Sataplia State Reserve, Georgia, after Golovatch (1977)). A, E, G: promere, caudal view. B, D: left gonopods, mesal view. C: promere with flagella, caudal view. F: right gonopods, mesal view. H, I: left gonopods, mesal view. Scale bars: 0.3 mm. G, H, I not to scale. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere; t, mesal tooth on promere." httpUri="https://zenodo.org/record/1459888/files/figure.png" pageId="7" pageNumber="96">
Figs 8H,
<date id="9D19102D9311FF9DFC9DF192FCE118CB" box="[775,799,1736,1760]" pageId="7" pageNumber="96">8I</date>
</figureCitation>
) (see also
<bibRefCitation id="8D364B1C9311FF9DFC39F19DFB9418CB" box="[931,1130,1735,1760]" pageId="7" pageNumber="96" refId="ref19572" refString="Lohmander, H. (1936) Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar, 5, 5 B (1), 1 - 196." type="journal article">Lohmander 1936</bibRefCitation>
;
<bibRefCitation id="8D364B1C9311FF9DFBE0F19DFAC818CB" box="[1146,1334,1735,1760]" pageId="7" pageNumber="96" refId="ref18710" refString="Golovatch, S. I. (1977) [New or poorly-known species of Julida (Diplopoda) from the Sataplia Reservation, Georgia, Transcaucasia]. Byulleten' Moskovskogo obshchestva ispytatelei prirody, Otdel biologicheskii, 82 (4), 46 - 51. [in Russian, a summary in English]" type="journal article">Golovatch 1977</bibRefCitation>
). Lateral outgrowth (
<emphasis id="DBD3EAFF9311FF9DFE80F1B6FECE192E" bold="true" box="[282,304,1772,1797]" pageId="7" pageNumber="96">lo</emphasis>
) of
<emphasis id="DBD3EAFF9311FF9DFEC5F1B6FE82192E" bold="true" box="[351,380,1772,1797]" pageId="7" pageNumber="96">op</emphasis>
narrow, pointed (
<figureCitation id="719C2A689311FF9DFDD8F1B6FD62192F" box="[578,668,1772,1797]" captionStart="FIGURE 8" captionStartId="12.[151,250,1726,1748]" captionTargetBox="[340,1248,625,1695]" captionTargetId="figure@12.[340,1248,625,1705]" captionTargetPageId="12" captionText="FIGURE 8. Male gonopods in Julus alexandrae Evsyukov, 2018 (AB from the Rostov-on-Don Region) and J. colchicus Lohmander, 1936 (CD from Krasnaya Polyana, Krasnodar Province, EF from Sakhrai, Adygea Republic, GH syntype from Gagry, Abkhazia, after Lohmander (1936), I from Sataplia State Reserve, Georgia, after Golovatch (1977)). A, E, G: promere, caudal view. B, D: left gonopods, mesal view. C: promere with flagella, caudal view. F: right gonopods, mesal view. H, I: left gonopods, mesal view. Scale bars: 0.3 mm. G, H, I not to scale. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere; t, mesal tooth on promere." httpUri="https://zenodo.org/record/1459888/files/figure.png" pageId="7" pageNumber="96">Fig. 8H</figureCitation>
), rounded (
<figureCitation id="719C2A689311FF9DFC81F1B6FC8C192F" box="[795,882,1772,1797]" captionStart="FIGURE 8" captionStartId="12.[151,250,1726,1748]" captionTargetBox="[340,1248,625,1695]" captionTargetId="figure@12.[340,1248,625,1705]" captionTargetPageId="12" captionText="FIGURE 8. Male gonopods in Julus alexandrae Evsyukov, 2018 (AB from the Rostov-on-Don Region) and J. colchicus Lohmander, 1936 (CD from Krasnaya Polyana, Krasnodar Province, EF from Sakhrai, Adygea Republic, GH syntype from Gagry, Abkhazia, after Lohmander (1936), I from Sataplia State Reserve, Georgia, after Golovatch (1977)). A, E, G: promere, caudal view. B, D: left gonopods, mesal view. C: promere with flagella, caudal view. F: right gonopods, mesal view. H, I: left gonopods, mesal view. Scale bars: 0.3 mm. G, H, I not to scale. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere; t, mesal tooth on promere." httpUri="https://zenodo.org/record/1459888/files/figure.png" pageId="7" pageNumber="96">Fig. 8D</figureCitation>
) or bifurcate (
<figureCitation id="719C2A689311FF9DFB89F1B6FBA1192F" box="[1043,1119,1772,1797]" captionStart="FIGURE 8" captionStartId="12.[151,250,1726,1748]" captionTargetBox="[340,1248,625,1695]" captionTargetId="figure@12.[340,1248,625,1705]" captionTargetPageId="12" captionText="FIGURE 8. Male gonopods in Julus alexandrae Evsyukov, 2018 (AB from the Rostov-on-Don Region) and J. colchicus Lohmander, 1936 (CD from Krasnaya Polyana, Krasnodar Province, EF from Sakhrai, Adygea Republic, GH syntype from Gagry, Abkhazia, after Lohmander (1936), I from Sataplia State Reserve, Georgia, after Golovatch (1977)). A, E, G: promere, caudal view. B, D: left gonopods, mesal view. C: promere with flagella, caudal view. F: right gonopods, mesal view. H, I: left gonopods, mesal view. Scale bars: 0.3 mm. G, H, I not to scale. Abbreviations: fl, flagellum; la, lamella on promere; lo, lateral outgrowth of opisthomere; ms, mesomere; op, opisthomere; pr, promere; t, mesal tooth on promere." httpUri="https://zenodo.org/record/1459888/files/figure.png" pageId="7" pageNumber="96">
Fig.
<date id="9D19102D9311FF9DFBDDF1B6FBA1192F" box="[1095,1119,1772,1796]" pageId="7" pageNumber="96">8I</date>
</figureCitation>
) at apex.
</paragraph>
<paragraph id="E91836ED9311FF9DFF5DF055FAA01903" blockId="7.[151,1437,151,2013]" box="[199,1374,1807,1832]" pageId="7" pageNumber="96">
Ventral edge of male segment 7 with small curved lamellae bordering the gonopodal aperture (
<figureCitation id="719C2A689311FF9DFB71F055FAAE1903" box="[1259,1360,1807,1832]" captionStart="FIGURE 15" captionStartId="20.[151,249,1687,1709]" captionTargetBox="[340,1248,1178,1666]" captionTargetId="figure@20.[340,1248,1178,1666]" captionTargetPageId="20" captionText="FIGURE 15. Ventral edge of male segments 7 in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936 C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. E: J. lignaui Verhoeff, 1910. F: J. lindholmi Lohmander, 1936. G: J. subalpinus Lohmander, 1936. H: J. khostensis sp. n. Scale bars: 0.3 mm." httpUri="https://zenodo.org/record/1459902/files/figure.png" pageId="7" pageNumber="96">Fig. 15B</figureCitation>
).
</paragraph>
<paragraph id="E91836ED9311FF9DFF5DF06EFA73195B" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="96">
Female vulvae (
<emphasis id="DBD3EAFF9311FF9DFE19F06EFE5E1966" bold="true" box="[387,416,1844,1869]" pageId="7" pageNumber="96">vu</emphasis>
) relatively wide; bursa (
<emphasis id="DBD3EAFF9311FF9DFD58F06EFD1F1966" bold="true" box="[706,737,1844,1869]" pageId="7" pageNumber="96">bu</emphasis>
) with a small, rounded, apical outgrowth; ampulla 1 (
<emphasis id="DBD3EAFF9311FF9DFAC5F06EFA6C1966" bold="true" box="[1375,1426,1844,1869]" pageId="7" pageNumber="96">am1</emphasis>
) located slightly basal to ampulla 2 (
<emphasis id="DBD3EAFF9311FF9DFDBCF00DFDA7195B" bold="true" box="[550,601,1879,1904]" pageId="7" pageNumber="96">am2</emphasis>
); coxa 2
<emphasis id="DBD3EAFF9311FF9DFD59F00DFD0F195B" bold="true" box="[707,753,1879,1904]" pageId="7" pageNumber="96">(12)</emphasis>
with a weakly expressed apicomesal outgrowth (
<figureCitation id="719C2A689311FF9DFA83F00DFA81195B" box="[1305,1407,1879,1904]" captionStart="FIGURE 17" captionStartId="22.[151,249,1033,1055]" captionTargetBox="[340,1248,193,1012]" captionTargetId="figure@22.[340,1248,193,1012]" captionTargetPageId="22" captionText="FIGURE 17. Female vulvae and legs 2 in Caucasian species of Julus. A: J. alexandrae Evsyukov, 2016. B: J. colchicus Lohmander, 1936. C: J. jedryczkowskii Golovatch, 1981. D: J. kubanus Verhoeff, 1921. Abbreviations: vu, vulva; op, operculum; bu, bursa; am1 and am2, ampulla 1 and ampulla 2; l2, coxa and prefemur of leg 2. Scale bar: 0.3 mm." httpUri="https://zenodo.org/record/1459906/files/figure.png" pageId="7" pageNumber="96">Fig. 17B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A1BD65669311FF92FF5DF026FED71F6B" lastPageId="8" lastPageNumber="97" pageId="7" pageNumber="96" type="discussion">
<paragraph id="E91836ED9311FF92FF5DF026FDEC1ED3" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1436,151,320]" lastPageId="8" lastPageNumber="97" pageId="7" pageNumber="96">
<emphasis id="DBD3EAFF9311FF9DFF5DF026FEC519BE" bold="true" box="[199,315,1916,1941]" pageId="7" pageNumber="96">Remarks.</emphasis>
This is the most common and widespread species of
<taxonomicName id="2EA74D6E9311FF9DFC20F026FC0B19BE" authorityName="Evsyukov &amp; Golovatch &amp; Reip" authorityYear="2018" box="[954,1013,1916,1941]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="DBD3EAFF9311FF9DFC20F026FC0B19BE" box="[954,1013,1916,1941]" italics="true" pageId="7" pageNumber="96">Julus</emphasis>
</taxonomicName>
in the Caucasus, recorded from the
<collectingRegion id="2B63F80F9311FF9DFF0DF0FAFEF21992" box="[151,268,1952,1977]" country="Russia" name="Krasnodar" pageId="7" pageNumber="96">Krasnodar</collectingRegion>
and
<collectingRegion id="2B63F80F9311FF9DFED8F0FAFE501992" box="[322,430,1952,1977]" country="Russia" name="Stavropol'" pageId="7" pageNumber="96">Stavropol</collectingRegion>
provinces, the
<collectingRegion id="2B63F80F9311FF9DFDC0F0FAFD4F1992" box="[602,689,1952,1977]" country="Russia" name="Adygeya" pageId="7" pageNumber="96">Adygea</collectingRegion>
and Karachaevo-Cherkessia republics,
<collectingRegion id="2B63F80F9311FF9DFBF7F0FAFB221992" box="[1133,1244,1952,1977]" country="Georgia" name="Abkhazia" pageId="7" pageNumber="96">Abkhazia</collectingRegion>
, most of
<collectingCountry id="91B0767D9311FF9DFAD8F0FAFA621992" box="[1346,1436,1952,1977]" name="Georgia" pageId="7" pageNumber="96">Georgia</collectingCountry>
and the adjacent parts of
<collectingCountry id="91B0767D9311FF9DFE35F09EFE0119F6" box="[431,511,1988,2013]" name="Turkey" pageId="7" pageNumber="96">Turkey</collectingCountry>
(
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;
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;
<bibRefCitation id="8D364B1C9311FF9DFCFFF09EFBD019F6" box="[869,1070,1988,2013]" pageId="7" pageNumber="96" refId="ref19152" refString="Kobakhidze, D. N. (1964) [Myriapoda]. Animal life of Georgia, 2 (Arthropoda), 186 - 195 [Tbilisi, in Georgian]" type="book chapter">Kobakhidze 1964</bibRefCitation>
,
<bibRefCitation id="8D364B1C9311FF9DFBA3F09EFB8A19F6" box="[1081,1140,1988,2013]" pageId="7" pageNumber="96" refId="ref19184" refString="Kobakhidze, D. N. (1965) [A list of millipedes (Diplopoda) of SSR Georgia]. Fragmenta Faunistica, 11 (21), 390 - 398. [in Russian, summaries in Polish and German]" type="journal article">1965</bibRefCitation>
;
<bibRefCitation id="8D364B1C9311FF9DFB1AF09EFAC919F7" box="[1152,1335,1988,2013]" pageId="7" pageNumber="96" refId="ref18710" refString="Golovatch, S. I. (1977) [New or poorly-known species of Julida (Diplopoda) from the Sataplia Reservation, Georgia, Transcaucasia]. Byulleten' Moskovskogo obshchestva ispytatelei prirody, Otdel biologicheskii, 82 (4), 46 - 51. [in Russian, a summary in English]" type="journal article">Golovatch 1977</bibRefCitation>
;
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;
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;
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;
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Korobushkin
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). It inhabits mixed and deciduous forests, as well as subalpine and alpine meadows, occurring at a range of elevations from the Black Sea coast to the alpine belt.
</paragraph>
<paragraph id="E91836ED931EFF92FF5DF65EFED71F6B" blockId="8.[151,1436,151,320]" pageId="8" pageNumber="97">The species shows only minor morphological variations in the structure of male coxa 1 and gonopodal opisthomere.</paragraph>
</subSubSection>
</treatment>
</document>