222 lines
26 KiB
XML
222 lines
26 KiB
XML
<document id="C43557A37DFC05D006D16AC1B7F5B803" ID-DOI="10.5281/zenodo.183429" ID-GBIF-Dataset="1ca3efaf-ed1b-48b0-8b05-d6a5e6005e54" ID-ISSN="1175-5326" ID-Zenodo-Dep="183429" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459907369404" checkinUser="plazi" docAuthor="Rintelen, Thomas Von & Glaubrecht, Matthias" docDate="2008" docId="5F2DDF50FFB29339FF26F85391119A8E" docLanguage="en" docName="zt01852p049.pdf" docOrigin="Zootaxa 1852" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tylomelania baskasti Rintelen & Glaubrecht, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="41" masterDocId="A314A728FFB0933DFFB1FFDC921D9D4F" masterDocTitle="Three new species of the freshwater snail genus Tylomelania (Caenogastropoda: Pachychilidae) from the Malili lake system, Sulawesi, Indonesia" masterLastPageNumber="49" masterPageNumber="37" pageNumber="39" updateTime="1698229100631" updateUser="plazi">
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<mods:mods id="ED95541D64BA8CD4824DDFCAA65B33B4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A3AF69BCE793F207BFBA005C7241468F">
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<mods:title id="F7DCD917425D5AB208B29740B7D4705D">Three new species of the freshwater snail genus Tylomelania (Caenogastropoda: Pachychilidae) from the Malili lake system, Sulawesi, Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name id="40EAA5B390E98B03DA5C9D83CE76BB49" type="personal">
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<mods:role id="B802BFFBDECE69C64790844FB23E53D8">
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<mods:roleTerm id="87BF421A68D5604FCA00FD33CB0E1F03">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="10752DDB19A170011D1C3E094B771BF4">Rintelen, Thomas Von</mods:namePart>
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</mods:name>
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<mods:name id="AE1D09CD1690E10F9ABF71DF20F9CD25" type="personal">
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<mods:role id="BD64744A24F28061D6C904B0566ADAEA">
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<mods:roleTerm id="F6D34330CB027CA9E42BBBC258929F9A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="78120F575352152DC32018EDC43A53F0">Glaubrecht, Matthias</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="FCB01B29448676C8F5865B3944482050">text</mods:typeOfResource>
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<mods:relatedItem id="D9025BCE729E56111B9D8B838DA56DFA" type="host">
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<mods:titleInfo id="42F60CE78A200FCFC895681DB5FC7170">
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<mods:title id="55F39EBED8D95B1673B2AE96F99C0D93">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0821FF67232AE2A988033725840CDD29">
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<mods:date id="F8B0374187EA6D8F19D09163F60B66E8">2008</mods:date>
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<mods:detail id="97FF11BFD07298FAB6595645331C3E2A" type="volume">
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<mods:number id="ABB413870D5787D717862F8C77D818EC">1852</mods:number>
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</mods:detail>
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<mods:extent id="9A244F2B0BD156BC428BA91AD98FB1BB" unit="page">
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<mods:start id="39A6C22802F69502BDBF3CDF1716A2A7">37</mods:start>
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<mods:end id="19A7C3E0D32C7E3C1716AE5311AF4479">49</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="5CA0B8BA55883168EA04B6A65C6C03A2">journal article</mods:classification>
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<mods:identifier id="433DABA583F3AD79F0B437240F908ABB" type="DOI">10.5281/zenodo.183429</mods:identifier>
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<mods:identifier id="7A59A8A6B654ED1D71DEA37574AD9880" type="GBIF-Dataset">1ca3efaf-ed1b-48b0-8b05-d6a5e6005e54</mods:identifier>
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<mods:identifier id="16A3B32BCC1215E989A419B32FA35599" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="EA0EFB48DCAE51426145AA803FA884B0" type="Zenodo-Dep">183429</mods:identifier>
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<treatment id="5F2DDF50FFB29339FF26F85391119A8E" ID-DOI="http://doi.org/10.5281/zenodo.5621355" ID-GBIF-Taxon="119373540" ID-Zenodo-Dep="5621355" LSID="urn:lsid:plazi:treatment:5F2DDF50FFB29339FF26F85391119A8E" httpUri="http://treatment.plazi.org/id/5F2DDF50FFB29339FF26F85391119A8E" lastPageId="4" lastPageNumber="41" pageId="2" pageNumber="39">
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<subSubSection id="9F9E3DCDFFB2933FFF26F85393E69AE6" box="[151,507,1935,1961]" pageId="2" pageNumber="39" type="nomenclature">
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<paragraph id="D73B6E46FFB2933FFF26F85393E69AE6" blockId="2.[151,507,1935,1961]" box="[151,507,1935,1961]" pageId="2" pageNumber="39">
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<heading id="8C73D92AFFB2933FFF26F85393E69AE6" bold="true" box="[151,507,1935,1961]" fontSize="11" level="1" pageId="2" pageNumber="39" reason="1">
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<emphasis id="E5F0B254FFB2933FFF26F85393E69AE6" bold="true" box="[151,507,1935,1961]" pageId="2" pageNumber="39">
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<taxonomicName id="108415C5FFB2933FFF26F853938A9AE6" box="[151,407,1935,1961]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="2" pageNumber="39" phylum="Mollusca" rank="species" species="baskasti" status="sp. nov.">
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<emphasis id="E5F0B254FFB2933FFF26F853938A9AE6" bold="true" box="[151,407,1935,1961]" italics="true" pageId="2" pageNumber="39">Tylomelania baskasti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FEC30F2FFFB2933FFE2CF85393E69AE6" box="[413,507,1935,1961]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9F9E3DCDFFB3933EFF26FF4B935C9C65" pageId="3" pageNumber="40" type="materials_examined">
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<paragraph id="D73B6E46FFB3933EFF26FF4B935C9C65" blockId="3.[151,1436,151,298]" pageId="3" pageNumber="40">
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<emphasis id="E5F0B254FFB3933EFF26FF4B93539DFE" bold="true" box="[151,334,151,177]" pageId="3" pageNumber="40">
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<typeStatus id="083FD0E4FFB3933EFF26FF4B92CF9DFE" box="[151,210,151,177]" pageId="3" pageNumber="40">Type</typeStatus>
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material.
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</emphasis>
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<collectingCountry id="AF932ED6FFB3933EFEEDFF4493CC9DFD" box="[348,465,152,178]" name="Indonesia" pageId="3" pageNumber="40">Indonesia</collectingCountry>
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, Sulawesi, Larona River:
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<typeStatus id="083FD0E4FFB3933EFCBFFF4491669DFD" box="[782,891,152,178]" pageId="3" pageNumber="40" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="4FBF72C3FFB3933EFC3EFF4491C49DFD" box="[911,985,152,178]" captionStart="FIGURE 2. T" captionStartId="3.[151,254,1485,1509]" captionTargetBox="[188,1397,355,1450]" captionTargetId="figure@3.[188,1398,354,1461]" captionTargetPageId="3" captionText="FIGURE 2. T. baskasti sp. nov.; A – D. shells, A. holotype, MZB Gst. 12.108 (loc. 71 - 02), B. paratypes, ZMB Moll. 190535 (loc. 71 - 02), C. paratypes, ZMB Moll. 190534 (loc. 15 - 02), D. paratypes, ZMB Moll. 190533 (loc. 14 - 02). Scale bar = 1 cm. E – F. opercula, E. paratype, ZMB Moll. 190534 (loc. 15 - 02), F. paratype, ZMB Moll. 190533 (loc. 14 - 02). Scale bar = 0.5 cm." httpUri="https://zenodo.org/record/183431/files/figure.png" pageId="3" pageNumber="40">Fig. 2</figureCitation>
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A;
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<quantity id="107CC3A3FFB3933EFBB0FF4496709DFD" box="[1025,1133,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="5.2299999999999995" pageId="3" pageNumber="40" unit="mm" value="52.3">52.3 mm</quantity>
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x
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<quantity id="107CC3A3FFB3933EFB23FF44971C9DFD" box="[1170,1281,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="3" pageNumber="40" unit="mm" value="17.7">17.7 mm</quantity>
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, loc. 71-02),
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<collectionCode id="B195F683FFB3933EFF26FF1C92CA9D95" box="[151,215,192,218]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="40">MZB</collectionCode>
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Gst. 12.109;
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<typeStatus id="083FD0E4FFB3933EFEC8FF1C93F49D95" box="[377,489,192,218]" pageId="3" pageNumber="40" type="paratype">paratypes</typeStatus>
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(
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<figureCitation id="4FBF72C3FFB3933EFE4BFF1C905C9D95" box="[506,577,192,218]" captionStart="FIGURE 2. T" captionStartId="3.[151,254,1485,1509]" captionTargetBox="[188,1397,355,1450]" captionTargetId="figure@3.[188,1398,354,1461]" captionTargetPageId="3" captionText="FIGURE 2. T. baskasti sp. nov.; A – D. shells, A. holotype, MZB Gst. 12.108 (loc. 71 - 02), B. paratypes, ZMB Moll. 190535 (loc. 71 - 02), C. paratypes, ZMB Moll. 190534 (loc. 15 - 02), D. paratypes, ZMB Moll. 190533 (loc. 14 - 02). Scale bar = 1 cm. E – F. opercula, E. paratype, ZMB Moll. 190534 (loc. 15 - 02), F. paratype, ZMB Moll. 190533 (loc. 14 - 02). Scale bar = 0.5 cm." httpUri="https://zenodo.org/record/183431/files/figure.png" pageId="3" pageNumber="40">Fig. 2</figureCitation>
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B-D): loc. 14–02,
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<collectionCode id="B195F683FFB3933EFCAAFF1C91469D95" box="[795,859,192,218]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="40">MZB</collectionCode>
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Gst. 12.110, n=5;
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<collectionCode id="B195F683FFB3933EFB88FF1C96649D95" box="[1081,1145,192,218]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="40">ZMB</collectionCode>
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Moll.
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<date id="A33A4886FFB3933EFB7BFF1C97389D95" box="[1226,1317,192,218]" pageId="3" pageNumber="40" value="33-05-19">190533</date>
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, n=7; loc. 15-02,
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<collectionCode id="B195F683FFB3933EFF5AFF3493369C4D" box="[235,299,232,258]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="40">MZB</collectionCode>
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Gst. 12.111, n=12;
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<collectionCode id="B195F683FFB3933EFDA9FF3490459C4D" box="[536,600,232,258]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="40">ZMB</collectionCode>
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Moll.
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<date id="A33A4886FFB3933EFD18FF3491199C4D" box="[681,772,232,258]" pageId="3" pageNumber="40" value="34-05-19">190534</date>
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, n=16; loc. 71-02,
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<collectionCode id="B195F683FFB3933EFC57FF34963B9C4D" box="[998,1062,232,258]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="40">MZB</collectionCode>
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Gst. 12.112, n=14;
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<collectionCode id="B195F683FFB3933EFAA5FF3497499C4D" box="[1300,1364,232,258]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="3" pageNumber="40">ZMB</collectionCode>
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Moll.
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<date id="A33A4886FFB3933EFF26FECC92EF9C65" box="[151,242,272,298]" pageId="3" pageNumber="40" value="35-05-19">190535</date>
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, n=17.
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</paragraph>
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</subSubSection>
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<caption id="83FB3ECEFFB3933EFF26FA1193789B03" httpUri="https://zenodo.org/record/183431/files/figure.png" pageId="3" pageNumber="40" targetBox="[188,1397,355,1450]" targetPageId="3">
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<paragraph id="D73B6E46FFB3933EFF26FA1193789B03" blockId="3.[151,1436,1485,1612]" pageId="3" pageNumber="40">
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<emphasis id="E5F0B254FFB3933EFF26FA11930698AA" bold="true" box="[151,283,1485,1509]" pageId="3" pageNumber="40">FIGURE 2.</emphasis>
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<taxonomicName id="108415C5FFB3933EFE95FA12938598AA" box="[292,408,1486,1509]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="3" pageNumber="40" phylum="Mollusca" rank="species" species="baskasti" status="sp. nov.">
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<emphasis id="E5F0B254FFB3933EFE95FA12938598AA" box="[292,408,1486,1509]" italics="true" pageId="3" pageNumber="40">T. baskasti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FEC30F2FFFB3933EFE13FA1193E798AA" box="[418,506,1485,1509]" pageId="3" pageNumber="40" rank="species">
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<emphasis id="E5F0B254FFB3933EFE13FA1193E798AA" bold="true" box="[418,506,1485,1509]" pageId="3" pageNumber="40">sp. nov.</emphasis>
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</taxonomicNameLabel>
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; A–D. shells, A. holotype, MZB Gst. 12.108 (loc. 71-02), B. paratypes, ZMB Moll. 190535 (loc. 71-02), C. paratypes, ZMB Moll. 190534 (loc. 15-02), D. paratypes, ZMB Moll. 190533 (loc. 14-02). Scale bar = 1 cm. E–F. opercula, E. paratype, ZMB Moll. 190534 (loc. 15-02), F. paratype, ZMB Moll. 190533 (loc. 14-02). Scale bar = 0.5 cm.
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</paragraph>
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</caption>
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<subSubSection id="9F9E3DCDFFB3933EFF77F95C91689B8D" pageId="3" pageNumber="40" type="etymology">
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<paragraph id="D73B6E46FFB3933EFF77F95C91689B8D" blockId="3.[151,1436,1664,2010]" pageId="3" pageNumber="40">
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<emphasis id="E5F0B254FFB3933EFF77F95C934E9BD5" bold="true" box="[198,339,1664,1690]" pageId="3" pageNumber="40">Etymology.</emphasis>
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The new species has been named
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<taxonomicName id="108415C5FFB3933EFD43F95D914F9BD5" box="[754,850,1665,1690]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="3" pageNumber="40" phylum="Mollusca" rank="species" species="baskasti">
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<emphasis id="E5F0B254FFB3933EFD43F95D914F9BD5" box="[754,850,1665,1690]" italics="true" pageId="3" pageNumber="40">baskasti</emphasis>
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</taxonomicName>
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in honour of Bas Kast, who has generously supported research on these snails at the Museum für Naturkunde.
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</paragraph>
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</subSubSection>
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<subSubSection id="9F9E3DCDFFB39339FF77F90C917C9B7E" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40" type="description">
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<paragraph id="D73B6E46FFB3933EFF77F90C91E99A2D" blockId="3.[151,1436,1664,2010]" pageId="3" pageNumber="40">
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<emphasis id="E5F0B254FFB3933EFF77F90C93439BA5" bold="true" box="[198,350,1744,1770]" pageId="3" pageNumber="40">Description.</emphasis>
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Shell (
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<figureCitation id="4FBF72C3FFB3933EFE04F90C93E19BA5" box="[437,508,1744,1770]" captionStart="FIGURE 2. T" captionStartId="3.[151,254,1485,1509]" captionTargetBox="[188,1397,355,1450]" captionTargetId="figure@3.[188,1398,354,1461]" captionTargetPageId="3" captionText="FIGURE 2. T. baskasti sp. nov.; A – D. shells, A. holotype, MZB Gst. 12.108 (loc. 71 - 02), B. paratypes, ZMB Moll. 190535 (loc. 71 - 02), C. paratypes, ZMB Moll. 190534 (loc. 15 - 02), D. paratypes, ZMB Moll. 190533 (loc. 14 - 02). Scale bar = 1 cm. E – F. opercula, E. paratype, ZMB Moll. 190534 (loc. 15 - 02), F. paratype, ZMB Moll. 190533 (loc. 14 - 02). Scale bar = 0.5 cm." httpUri="https://zenodo.org/record/183431/files/figure.png" pageId="3" pageNumber="40">Fig. 2</figureCitation>
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A–D): Medium sized to large, brown, elongate conic, spire angle 13–25°. Top whorls in adult specimens always corroded to a varying degree, 4–9 remaining whorls, can reach up to
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<quantity id="107CC3A3FFB3933EFAD9F92492D89A75" metricMagnitude="-2" metricUnit="m" metricValue="5.47" pageId="3" pageNumber="40" unit="mm" value="54.7">54.7 mm</quantity>
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(
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<tableCitation id="9A065BFDFFB3933EFF67F8FC933C9A75" box="[214,289,1824,1850]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionTargetBox="[159,1390,208,479]" captionTargetPageId="2" captionText="TABLE 1. List of sampling stations with locality details. The locality numbers correspond to those in Fig. 1." httpUri="http://table.plazi.org/id/83FB3ECEFFB2933FFF26FF4B97029DE0" pageId="3" pageNumber="40" tableUuid="83FB3ECEFFB2933FFF26FF4B97029DE0">Tab. 1</tableCitation>
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). With spiral ribs only. Aperture oval, pointed at top and slightly siphonated at base. Columella and interior of aperture brown, in few specimens slightly whitish coating.
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</paragraph>
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<paragraph id="D73B6E46FFB3933EFF77F8AC96549AFD" blockId="3.[151,1436,1664,2010]" pageId="3" pageNumber="40">External morphology: Headfoot black with fine orange dots, sometimes rather dense, foot more intensely pigmented, mantle edge serrated to a varying degree. Body coiled in 3–6 whorls.</paragraph>
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<paragraph id="D73B6E46FFB3933EFF26F81C96F79A95" blockId="3.[151,1436,1664,2010]" box="[151,1258,1984,2010]" pageId="3" pageNumber="40">
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Operculum (
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<figureCitation id="4FBF72C3FFB3933EFE9AF81C936C9A95" box="[299,369,1984,2010]" captionStart="FIGURE 2. T" captionStartId="3.[151,254,1485,1509]" captionTargetBox="[188,1397,355,1450]" captionTargetId="figure@3.[188,1398,354,1461]" captionTargetPageId="3" captionText="FIGURE 2. T. baskasti sp. nov.; A – D. shells, A. holotype, MZB Gst. 12.108 (loc. 71 - 02), B. paratypes, ZMB Moll. 190535 (loc. 71 - 02), C. paratypes, ZMB Moll. 190534 (loc. 15 - 02), D. paratypes, ZMB Moll. 190533 (loc. 14 - 02). Scale bar = 1 cm. E – F. opercula, E. paratype, ZMB Moll. 190534 (loc. 15 - 02), F. paratype, ZMB Moll. 190533 (loc. 14 - 02). Scale bar = 0.5 cm." httpUri="https://zenodo.org/record/183431/files/figure.png" pageId="3" pageNumber="40">Fig. 2</figureCitation>
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E,F): roundish-ovate, last whorl inflated, multispiral with 5–7 whorls (n=3).
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</paragraph>
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<paragraph id="D73B6E46FFB49339FF74FF44918E9C65" blockId="4.[151,1436,152,298]" pageId="4" pageNumber="41">
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Radula (
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<figureCitation id="4FBF72C3FFB49339FE9DFF4493699DFD" box="[300,372,152,178]" captionStart="FIGURE 3. T" captionStartId="4.[151,254,1328,1352]" captionTargetBox="[152,1434,354,1294]" captionTargetId="figure@4.[152,1435,354,1304]" captionTargetPageId="4" captionText="FIGURE 3. T. baskasti sp. nov., ZMB Moll. 190534 (loc. 15 - 02). A – B. radula, A. segment, frontal, B. segment, apical (45 °). Scale bar = 0.1 mm. C – E. embryonic shells, C. lateral view, D. apical whorls, lateral, E. apical view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183432/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
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A,B): 160–194 rows,
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<quantity id="107CC3A3FFB49339FDCCFF4491369DFD" box="[637,811,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" metricValueMax="2.42" metricValueMin="1.64" pageId="4" pageNumber="41" unit="mm" value="20.299999999999997" valueMax="24.2" valueMin="16.4">16.4–24.2 mm</quantity>
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long, on average 8.8 teeth per mm (n=12). Central tooth with pointed and enlarged major denticle. Glabella with very slightly rounded base. Lateral teeth with pointed and enlarged major denticles and 2–3 minor denticles on each side. Marginal teeth shovel-like, inner and outer marginals with three almost equal-sized denticles each.
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</paragraph>
|
||
<caption id="83FB3ECEFFB49339FF26FAEC92D598C2" httpUri="https://zenodo.org/record/183432/files/figure.png" pageId="4" pageNumber="41" targetBox="[152,1434,354,1294]" targetPageId="4">
|
||
<paragraph id="D73B6E46FFB49339FF26FAEC92D598C2" blockId="4.[151,1436,1328,1421]" pageId="4" pageNumber="41">
|
||
<emphasis id="E5F0B254FFB49339FF26FAEC93079807" bold="true" box="[151,282,1328,1352]" pageId="4" pageNumber="41">FIGURE 3.</emphasis>
|
||
<taxonomicName id="108415C5FFB49339FE90FAED93899807" box="[289,404,1329,1352]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="baskasti" status="sp. nov.">
|
||
<emphasis id="E5F0B254FFB49339FE90FAED93899807" box="[289,404,1329,1352]" italics="true" pageId="4" pageNumber="41">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FEC30F2FFFB49339FE2AFAEC93EC9807" box="[411,497,1328,1352]" pageId="4" pageNumber="41" rank="species">
|
||
<emphasis id="E5F0B254FFB49339FE2AFAEC93EC9807" bold="true" box="[411,497,1328,1352]" pageId="4" pageNumber="41">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, ZMB Moll. 190534 (loc. 15-02). A–B. radula, A. segment, frontal, B. segment, apical (45°). Scale bar = 0.1 mm. C–E. embryonic shells, C. lateral view, D. apical whorls, lateral, E. apical view. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D73B6E46FFB49339FF74FA1D979A9894" blockId="4.[151,1437,1473,1985]" box="[197,1415,1473,1499]" pageId="4" pageNumber="41">
|
||
Reproductive biology: Brood pouch contains 4–12 embryos, their size can reach
|
||
<quantity id="107CC3A3FFB49339FBC8FA1D96C99894" box="[1145,1236,1473,1499]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="4" pageNumber="41" unit="mm" value="8.4">8.4 mm</quantity>
|
||
(n=5) (Tab. 3).
|
||
</paragraph>
|
||
<paragraph id="D73B6E46FFB49339FF74FA30917C9B7E" blockId="4.[151,1437,1473,1985]" pageId="4" pageNumber="41">
|
||
Embryonic shells (
|
||
<figureCitation id="4FBF72C3FFB49339FE14FA3093F69B49" box="[421,491,1516,1542]" captionStart="FIGURE 3. T" captionStartId="4.[151,254,1328,1352]" captionTargetBox="[152,1434,354,1294]" captionTargetId="figure@4.[152,1435,354,1304]" captionTargetPageId="4" captionText="FIGURE 3. T. baskasti sp. nov., ZMB Moll. 190534 (loc. 15 - 02). A – B. radula, A. segment, frontal, B. segment, apical (45 °). Scale bar = 0.1 mm. C – E. embryonic shells, C. lateral view, D. apical whorls, lateral, E. apical view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183432/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
|
||
C–E): Elongate-conic, axial ribs emerging on the 2nd to 3rd whorl and fading on the 5th whorl. Spiral striae emerge on 3rd to 4th whorl (Tab. 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9F9E3DCDFFB49339FF77F9E291119A8E" pageId="4" pageNumber="41" type="distribution">
|
||
<paragraph id="D73B6E46FFB49339FF77F9E296B59B16" blockId="4.[151,1437,1473,1985]" box="[198,1192,1598,1625]" pageId="4" pageNumber="41">
|
||
<emphasis id="E5F0B254FFB49339FF77F9E293E39B17" bold="true" box="[198,510,1598,1624]" pageId="4" pageNumber="41">Distribution and habitat.</emphasis>
|
||
South Sulawesi, lower reaches of Larona River (
|
||
<figureCitation id="4FBF72C3FFB49339FB8CF9E3969D9B16" box="[1085,1152,1599,1625]" captionStart="FIGURE 1" captionStartId="1.[151,254,1586,1610]" captionTargetBox="[154,1436,281,1555]" captionTargetId="figure@1.[151,1436,277,1560]" captionTargetPageId="1" captionText="FIGURE 1. Sulawesi and the Malili lake system with sample sites. Locality numbers are the original field numbers and correspond to those in Tab. 1." httpUri="https://zenodo.org/record/183430/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="D73B6E46FFB49339FF77F9BB91499B9E" blockId="4.[151,1437,1473,1985]" pageId="4" pageNumber="41">
|
||
This species was collected in shallow water (
|
||
<quantity id="107CC3A3FFB49339FD66F9BB91529BCE" box="[727,847,1639,1665]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="4" pageNumber="41" unit="m" value="0.3" valueMax="0.5" valueMin="0.1">0.1–0.5 m</quantity>
|
||
depth) in less turbulent zones at the river bank on soft substrate.
|
||
<collectingCountry id="AF932ED6FFB49339FEF6F95393759BE6" box="[327,360,1679,1705]" name="American Samoa" pageId="4" pageNumber="41">As</collectingCountry>
|
||
deeper parts of the river were not accessible for sampling because of strong currents,
|
||
<taxonomicName id="108415C5FFB49339FA37F95392EA9B9F" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="baskasti">
|
||
<emphasis id="E5F0B254FFB49339FA37F95392EA9B9F" italics="true" pageId="4" pageNumber="41">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
must not necessarily be restricted to shallow water.
|
||
</paragraph>
|
||
<paragraph id="D73B6E46FFB49339FF77F90291119A8E" blockId="4.[151,1437,1473,1985]" pageId="4" pageNumber="41">
|
||
<emphasis id="E5F0B254FFB49339FF77F90293D79BB7" bold="true" box="[198,458,1758,1784]" pageId="4" pageNumber="41">Taxonomic remarks.</emphasis>
|
||
Shell shape and radula of
|
||
<taxonomicName id="108415C5FFB49339FCA5F90391899BB7" box="[788,916,1759,1784]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="baskasti">
|
||
<emphasis id="E5F0B254FFB49339FCA5F90391899BB7" box="[788,916,1759,1784]" italics="true" pageId="4" pageNumber="41">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
are similar to that of
|
||
<taxonomicName id="108415C5FFB49339FB13F90392C69A6E" authority="Kruimel, 1913" authorityName="Kruimel" authorityYear="1913" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="lalemae">
|
||
<emphasis id="E5F0B254FFB49339FB13F90397029BB7" box="[1186,1311,1759,1784]" italics="true" pageId="4" pageNumber="41">T. lalemae</emphasis>
|
||
(
|
||
<bibRefCitation id="B31513B7FFB49339FA80F90392CE9A6E" author="Kruimel" pageId="4" pageNumber="49" refString="Kruimel, J. H. (1913) Verzeichnis der von Herrn E. C. Abendanon in Celebes gesammelten Susswasser-Mollusken. Bijdragen tot de Dierkunde, 19, 217 - 235." type="journal article" year="1913">Kruimel, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from Lake Towuti. The shell of
|
||
<taxonomicName id="108415C5FFB49339FDE4F8DB90CD9A6F" box="[597,720,1799,1824]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="baskasti">
|
||
<emphasis id="E5F0B254FFB49339FDE4F8DB90CD9A6F" box="[597,720,1799,1824]" italics="true" pageId="4" pageNumber="41">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
is more fragile and always lacks the conspicuous white aperture of
|
||
<taxonomicName id="108415C5FFB49339FF5DF8F393799A07" box="[236,356,1839,1864]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="lalemae">
|
||
<emphasis id="E5F0B254FFB49339FF5DF8F393799A07" box="[236,356,1839,1864]" italics="true" pageId="4" pageNumber="41">T. lalemae</emphasis>
|
||
</taxonomicName>
|
||
, though. The radula of
|
||
<taxonomicName id="108415C5FFB49339FDCBF8F390EA9A07" box="[634,759,1839,1864]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="baskasti">
|
||
<emphasis id="E5F0B254FFB49339FDCBF8F390EA9A07" box="[634,759,1839,1864]" italics="true" pageId="4" pageNumber="41">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
also closely resembles that of all other described riverine species of Sulawesi such as e.g.
|
||
<taxonomicName id="108415C5FFB49339FDA3F88B917F9A3E" authority="Mousson, 1849" authorityName="Mousson" authorityYear="1849" box="[530,866,1879,1905]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="perfecta">
|
||
<emphasis id="E5F0B254FFB49339FDA3F88B90919A3F" box="[530,652,1879,1904]" italics="true" pageId="4" pageNumber="41">T. perfecta</emphasis>
|
||
(Mousson, 1849)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="108415C5FFB49339FCDDF88B91F49A3F" box="[876,1001,1879,1904]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="4" pageNumber="41" phylum="Mollusca" rank="species" species="baskasti">
|
||
<emphasis id="E5F0B254FFB49339FCDDF88B91F49A3F" box="[876,1001,1879,1904]" italics="true" pageId="4" pageNumber="41">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
can be unambiguously distinguished from those taxa by its characteristic shell and the molecular data clearly confirm its relationship to the Malili lakes species flock (compare also below,
|
||
<emphasis id="E5F0B254FFB49339FDCAF87B90E19A8F" box="[635,764,1959,1984]" italics="true" pageId="4" pageNumber="41">Discussion</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |