199 lines
28 KiB
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199 lines
28 KiB
XML
<document ID-CLB-Dataset="7243" ID-DOI="http://dx.doi.org/10.3897/asp.79.e66978" ID-GBIF-Dataset="a9bfd01a-cc28-46f4-a9ef-d0a41e19eaa9" ID-Pensoft-Pub="1864-8312-79-37" ID-Pensoft-UUID="CE28E31D8CC05ACD8DEEEA5E2094E9D2" ID-ZooBank="6E3AE6524AA549EFB44DC5B99C2AD0C6" ModsDocID="1864-8312-79-37" checkinTime="1618938705165" checkinUser="pensoft" docAuthor="Ward, Philip S. & Boudinot, Brendon E." docDate="2021" docId="4F2F51880B385B9CABB0E36160EC1B8E" docLanguage="en" docName="ArthropodSystPhyl 79: 37-46" docOrigin="Arthropod Systematics & Phylogeny 79" docPubDate="2021-04-19" docSource="http://dx.doi.org/10.3897/asp.79.e66978" docTitle="Camponotus laevigatus" docType="treatment" docVersion="6" id="CE28E31D8CC05ACD8DEEEA5E2094E9D2" lastPageNumber="37" masterDocId="CE28E31D8CC05ACD8DEEEA5E2094E9D2" masterDocTitle="Grappling with homoplasy: taxonomic refinements and reassignments in the ant genera Camponotus and Colobopsis (Hymenoptera: Formicidae)" masterLastPageNumber="56" masterPageNumber="37" pageNumber="37" updateTime="1732574069365" updateUser="ExternalLinkService">
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<mods:mods id="B0FC353EF0C0A7E896E8F53AB118BD4C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B8A53C7D58742A7964873B8524ADAE57">
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<mods:title id="BAD5DD86F41AE2EEC05600ADF93FD0D6">Grappling with homoplasy: taxonomic refinements and reassignments in the ant genera Camponotus and Colobopsis (Hymenoptera: Formicidae)</mods:title>
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<mods:name id="093FDDEF1EFFCAA53A8E545605530262" type="personal">
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<mods:roleTerm id="C1769B7435490B7D4A9175C3EEE2E5B2">Author</mods:roleTerm>
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<mods:namePart id="34B51AB0791DF9205454495FF30514FA">Ward, Philip S.</mods:namePart>
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<mods:affiliation id="224129E768A74EF1FB913D7E0E11442F">Department of Entomology & Nematology, University of California, Davis, One Shields Ave, Davis, CA 95616, USA; P. S. Ward *</mods:affiliation>
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<mods:nameIdentifier id="B78E86047E9FF546ABC877CEE96BF56F" type="email">psward@ucdavis.edu</mods:nameIdentifier>
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<mods:roleTerm id="431FBB2D64DF53B2D25EAB8CFC3AAD64">Author</mods:roleTerm>
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<mods:namePart id="2E18EE7A2533EB7DAD4260CB262563DA">Boudinot, Brendon E.</mods:namePart>
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<mods:nameIdentifier id="C05736574C759218693C0B38217FC52E" type="ORCID">https://orcid.org/0000-0002-4588-0430</mods:nameIdentifier>
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<mods:affiliation id="1A0179D9EEA58E8B0B5BB280CC30B33A">Department of Entomology & Nematology, University of California, Davis, One Shields Ave, Davis, CA 95616, USA; P. S. Ward * & Friedrich-Schiller-Universitaet Jena, Institut fuer Zoologie und Evolutionsforschung, Erberstrasse 1, Jena, Thuringia, 07743, Germany; B. E. Boudinot [boudinotb @ gmail. com]</mods:affiliation>
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<mods:typeOfResource id="89D3A110677B34E28BB3D88634475996">text</mods:typeOfResource>
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<mods:title id="011A1F288EF303A644C7A91661D95486">Arthropod Systematics & Phylogeny</mods:title>
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<mods:part id="D248E407894BD3113AFAB79425C62432">
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<mods:date id="01C9564A4EE310A8ED1F437EE9FD282C">2021</mods:date>
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<mods:number id="5AB89E6EBCF1C1D6F033FAAFE9BC1F5A">2021-04-19</mods:number>
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<mods:number id="795F11F5B6712F0EF7018CA9FDF0FE92">79</mods:number>
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<mods:start id="39C47BCBAA9AF60DB49BD132D10C0456">37</mods:start>
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<mods:end id="F9FA17C67CDB8AB4AD2DBCB215718282">56</mods:end>
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<mods:url id="B0D4B47E65C41498811784AC9A27E45B">http://dx.doi.org/10.3897/asp.79.e66978</mods:url>
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<mods:classification id="C646CDA10137DDCEA86D767429B4D06E">journal article</mods:classification>
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<mods:identifier id="AC7E9703A9CD765EB81425F5B1C13F21" type="DOI">http://dx.doi.org/10.3897/asp.79.e66978</mods:identifier>
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<treatment id="4F2F51880B385B9CABB0E36160EC1B8E" ID-GBIF-Taxon="181520739" LSID="urn:lsid:plazi:treatment:4F2F51880B385B9CABB0E36160EC1B8E" httpUri="http://treatment.plazi.org/id/4F2F51880B385B9CABB0E36160EC1B8E" lastPageNumber="37" pageId="0" pageNumber="37" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="E3C31FAF332E3224C731442680799DD0" pageId="0" pageNumber="37" type="nomenclature">
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<paragraph id="167BB7DA37AC506BA99E3231303D0C7D" pageId="0" pageNumber="37">
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3.2.2.
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<taxonomicName id="E058A511AB7D2A264DBE70870167D40B" ID-CoL="QDX2" LSID="4F2F5188-0B38-5B9C-ABB0-E36160EC1B8E" authority="(F. Smith)" baseAuthorityName="F. Smith" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">Camponotus laevigatus (F. Smith)</taxonomicName>
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<figureCitation id="7F9AC967CB2734B59A77C6254BD51B37" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Camponotus workers, full-face (dorsal) views of head (A, C, E) and lateral views of body (B, D, F); scale bars = 1.0 mm. A, B: Camponotus laevigatus, lectotype (CASENT 0903603). C, D: Camponotus quercicola, paratype (MCZ-ENT 00029334), image from MCZbase (Museum of Comparative Zoology, Harvard University, copyright President and Fellows of Harvard College). E, F: Camponotus laevissimus (CASENT 0280010). Images A, B, E, F from AntWeb (www. antweb. org); photographer Will Ericson." figureDoi="10.3897/asp.79.e66978.figure7" httpUri="https://binary.pensoft.net/fig/533435" pageId="0" pageNumber="37">Fig. 7A-D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4033D74791FA574CB3D65181B47FC3A7" pageId="0" pageNumber="37" type="reference_group">
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<paragraph id="79971FB5B6B7EE2230631BE51B6D56B5" pageId="0" pageNumber="37">
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<taxonomicName id="3AAA69C60A5CA2DF772BF64801DB98FE" authorityName="F.Smith" authorityYear="1858" class="Insecta" family="Formicidae" genus="Formica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Formica laevigata" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigata">Formica laevigata</taxonomicName>
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F. Smith, 1858: 55. Lectotype worker, California, United States (BMNH) [examined via image on AntWeb: CASENT0903603]
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</paragraph>
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<paragraph id="5D24048623892EBF2EE150F7165270B7" pageId="0" pageNumber="37">
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<taxonomicName id="78681B7F4B2412221E88DD2161B25B0A" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">Camponotus laevigatus</taxonomicName>
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(F. Smith); Roger, 1863: 5. Combination in
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<taxonomicName id="1F5C0958E9B11A2F7F7E8A80F660B6B0" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="genus">Camponotus</taxonomicName>
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.
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</paragraph>
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<paragraph id="CB85FAA8B4257EE65C53F32F522FD18D" pageId="0" pageNumber="37">
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<taxonomicName id="0539A4F66606FFEDC411B62519020163" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus (Camponotus) laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus" subGenus="Camponotus">Camponotus (Camponotus) laevigatus</taxonomicName>
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(F. Smith); Forel, 1914: 266. Placement in
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<taxonomicName id="A05FD120A95CFF2ACAC54702E4198875" authority="(Camponotus)" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus (Camponotus)" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="subGenus" subGenus="Camponotus">Camponotus (Camponotus)</taxonomicName>
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.
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</paragraph>
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<paragraph id="08B653A87B0D36D793E1A347BB116826" pageId="0" pageNumber="37">
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<taxonomicName id="AE81488E9199A77AEE0AA898C389D9A7" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus (Camponotus) laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus" subGenus="Camponotus">Camponotus (Camponotus) laevigatus</taxonomicName>
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(F. Smith); Mackay, 2019: 246. Lectotype worker designated.
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</paragraph>
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<paragraph id="6271CCF40F444BD491F641F86689F977" pageId="0" pageNumber="37">
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<taxonomicName id="A45EBE5A8CC83719F06A8CD08C9A95CA" authorityName="Smith" authorityYear="1954" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus (Camponotus) quercicola" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="quercicola" subGenus="Camponotus">Camponotus (Camponotus) quercicola</taxonomicName>
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M. R. Smith, 1954: 211. Holotype worker, Tanbark Flat, Los Angeles County, California (T. C. Lawrence) (USNM) [examined].
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<emphasis id="578CC595E89A7322A0E341D3268ACFCC" bold="true" pageId="0" pageNumber="37">Syn. nov.</emphasis>
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</paragraph>
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<paragraph id="E884EB720A57799D990366B1263F1208" pageId="0" pageNumber="37">
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<taxonomicName id="5843200F509CBA09141FD97DAF676793" authorityName="Smith" authorityYear="1954" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus quercicola" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="quercicola">Camponotus quercicola</taxonomicName>
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M. R. Smith;
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<bibRefCitation id="BEF0DA16B6518D035DEE647EC4031AE2" DOI="https://doi.org/10.1093/aesa/92.4.514" author="Gadau, J" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="37" pagination="514 - 522" refId="B20" refString="Gadau, J, Brady, SG, Ward, PS, 1999. Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae). Annals of the Entomological Society of America 92: 514 - 522, DOI: https://doi.org/10.1093/aesa/92.4.514" title="Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae)." url="https://doi.org/10.1093/aesa/92.4.514" volume="92" year="1999">Gadau et al., 1999</bibRefCitation>
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. Description of male and queen. Biology and distribution.
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</paragraph>
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</subSubSection>
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<subSubSection id="1530638712AF2C03A66EF809915696C0" pageId="0" pageNumber="37" type="">
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<paragraph id="4E5B842B41EDCC5ADCF018350F7E22D6" pageId="0" pageNumber="37">
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<bibRefCitation id="C2EDA739FC1F3D0EE5049BF76478CA74" author="Mackay, WP" journalOrPublisher="Lambert Academic Publishing, Mauritius" pageId="0" pageNumber="37" refId="B30" refString="Mackay, WP, 2019. New World carpenter ants of the hyperdiverse genus Camponotus. Volume 1. Introduction, keys to the subgenera and species complexes and the subgenus Camponotus. Lambert Academic Publishing, Mauritius" title="New World carpenter ants of the hyperdiverse genus Camponotus. Volume 1. Introduction, keys to the subgenera and species complexes and the subgenus Camponotus." year="2019">Mackay (2019)</bibRefCitation>
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discovered that the types of this species in BMNH-a syntype dealate queen and syntype major worker, the latter designated by him as lectotype-do not correspond to the species that has come to be known in the literature as
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<taxonomicName id="0F610FC909BE43CABF90660B2A3B5255" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">
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<emphasis id="F70DF5D403B447E30A2F67F9877B17ED" italics="true" pageId="0" pageNumber="37">Camponotus laevigatus</emphasis>
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</taxonomicName>
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. That species, given the new name
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<taxonomicName id="94DB52C5FB7B618A7E3E1E54C7F3DC44" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ca. laevissimus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevissimus">
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<emphasis id="F0E70BD06201F2A89E0ED019EDE09CCE" italics="true" pageId="0" pageNumber="37">Ca. laevissimus</emphasis>
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</taxonomicName>
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Mackay, is easily recognized by its shiny, iridescent blue-black integument and abundant and bright white standing pilosity on most of the body, including the scapes and tibiae (Fig.
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<figureCitation id="CB9C951EECA52007513BAEB8BAFED74F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Camponotus workers, full-face (dorsal) views of head (A, C, E) and lateral views of body (B, D, F); scale bars = 1.0 mm. A, B: Camponotus laevigatus, lectotype (CASENT 0903603). C, D: Camponotus quercicola, paratype (MCZ-ENT 00029334), image from MCZbase (Museum of Comparative Zoology, Harvard University, copyright President and Fellows of Harvard College). E, F: Camponotus laevissimus (CASENT 0280010). Images A, B, E, F from AntWeb (www. antweb. org); photographer Will Ericson." figureDoi="10.3897/asp.79.e66978.figure7" httpUri="https://binary.pensoft.net/fig/533435" pageId="0" pageNumber="37">7E, F</figureCitation>
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). The real
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<taxonomicName id="7EE1CD8EABFB4B92C7F552555CA5C9ED" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">
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<emphasis id="0D5669258BE188F3925DC56368B37126" italics="true" pageId="0" pageNumber="37">Camponotus laevigatus</emphasis>
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</taxonomicName>
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is a shiny black species, with relatively sparse standing pilosity, inconspicuous pubescence, slender scape base, and ecarinate clypeus. Examination of the lectotype image (Fig.
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<figureCitation id="60C04307AFD188C74225984B7D4016EE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Camponotus workers, full-face (dorsal) views of head (A, C, E) and lateral views of body (B, D, F); scale bars = 1.0 mm. A, B: Camponotus laevigatus, lectotype (CASENT 0903603). C, D: Camponotus quercicola, paratype (MCZ-ENT 00029334), image from MCZbase (Museum of Comparative Zoology, Harvard University, copyright President and Fellows of Harvard College). E, F: Camponotus laevissimus (CASENT 0280010). Images A, B, E, F from AntWeb (www. antweb. org); photographer Will Ericson." figureDoi="10.3897/asp.79.e66978.figure7" httpUri="https://binary.pensoft.net/fig/533435" pageId="0" pageNumber="37">7A, B</figureCitation>
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) shows that
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<taxonomicName id="B273918AA212257A9705B91B3C7B2AC9" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">
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<emphasis id="24DA188ACBA0F0210AEDDC50D43D1F85" italics="true" pageId="0" pageNumber="37">Camponotus laevigatus</emphasis>
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</taxonomicName>
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is conspecific with
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<taxonomicName id="EB8967B68DE692C2BF22037701D2AB00" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ca. quercicola" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="quercicola">
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<emphasis id="B1094D4EF820E8DBBFFE5A2808BC474C" italics="true" pageId="0" pageNumber="37">Ca. quercicola</emphasis>
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</taxonomicName>
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, a widespread California species that nests in the trunks and branches of oak trees (
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<bibRefCitation id="FB0D3BFB28144F9C09253A5020E7B355" DOI="https://doi.org/10.1093/aesa/92.4.514" author="Gadau, J" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="37" pagination="514 - 522" refId="B20" refString="Gadau, J, Brady, SG, Ward, PS, 1999. Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae). Annals of the Entomological Society of America 92: 514 - 522, DOI: https://doi.org/10.1093/aesa/92.4.514" title="Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae)." url="https://doi.org/10.1093/aesa/92.4.514" volume="92" year="1999">Gadau et al. 1999</bibRefCitation>
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) (Fig.
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<figureCitation id="7713CCD05CEB382E36B86B9A781CFB54" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Camponotus workers, full-face (dorsal) views of head (A, C, E) and lateral views of body (B, D, F); scale bars = 1.0 mm. A, B: Camponotus laevigatus, lectotype (CASENT 0903603). C, D: Camponotus quercicola, paratype (MCZ-ENT 00029334), image from MCZbase (Museum of Comparative Zoology, Harvard University, copyright President and Fellows of Harvard College). E, F: Camponotus laevissimus (CASENT 0280010). Images A, B, E, F from AntWeb (www. antweb. org); photographer Will Ericson." figureDoi="10.3897/asp.79.e66978.figure7" httpUri="https://binary.pensoft.net/fig/533435" pageId="0" pageNumber="37">7C, D</figureCitation>
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).
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<bibRefCitation id="11709AD09004C2F4DB172320C71324E7" author="Mackay, WP" journalOrPublisher="Lambert Academic Publishing, Mauritius" pageId="0" pageNumber="37" refId="B30" refString="Mackay, WP, 2019. New World carpenter ants of the hyperdiverse genus Camponotus. Volume 1. Introduction, keys to the subgenera and species complexes and the subgenus Camponotus. Lambert Academic Publishing, Mauritius" title="New World carpenter ants of the hyperdiverse genus Camponotus. Volume 1. Introduction, keys to the subgenera and species complexes and the subgenus Camponotus." year="2019">Mackay (2019)</bibRefCitation>
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claimed that
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<taxonomicName id="DCC568C94D787B5968C55FD43BDC1EB4" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ca. laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">
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<emphasis id="E494CAF191982FA01D08BFE78ABD85E8" italics="true" pageId="0" pageNumber="37">Ca. laevigatus</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="D41A6B0795810BF0760E2F7B94874476" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ca. quercicola" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="quercicola">
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<emphasis id="94609803294DA359C4FBB1396F87E001" italics="true" pageId="0" pageNumber="37">Ca. quercicola</emphasis>
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</taxonomicName>
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in having reduced pilosity on the head, but the lectotype is an old specimen in which the hairs are evidently abraded. Note the asymmetry in presence of hairs on the two sides of the head in the AntWeb image (e.g., short setae present on the left malar region but not on the right side) (Fig.
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<figureCitation id="BCF959C3C356AE7A595E6F355E6E3D43" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Camponotus workers, full-face (dorsal) views of head (A, C, E) and lateral views of body (B, D, F); scale bars = 1.0 mm. A, B: Camponotus laevigatus, lectotype (CASENT 0903603). C, D: Camponotus quercicola, paratype (MCZ-ENT 00029334), image from MCZbase (Museum of Comparative Zoology, Harvard University, copyright President and Fellows of Harvard College). E, F: Camponotus laevissimus (CASENT 0280010). Images A, B, E, F from AntWeb (www. antweb. org); photographer Will Ericson." figureDoi="10.3897/asp.79.e66978.figure7" httpUri="https://binary.pensoft.net/fig/533435" pageId="0" pageNumber="37">7A</figureCitation>
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). Moreover, the amount of standing pilosity shows considerable variation in workers of
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<taxonomicName id="B40A67EC743DD3EC906F0DBCCB8A26AE" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ca. quercicola" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="quercicola">
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<emphasis id="51CDF5209D7846FA0E6AD5E8EA20F78D" italics="true" pageId="0" pageNumber="37">Ca. quercicola</emphasis>
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</taxonomicName>
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, including setation on the malar region (
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<bibRefCitation id="353D059A1C97459C83E646BDADA01BC5" author="Smith, MR" journalOrPublisher="Journal of the New York Entomological Society" pageId="0" pageNumber="37" pagination="211 - 214" refId="B45" refString="Smith, MR, 1954. A new Camponotus in California apparently inhabiting live oak, Quercus sp. (Hymenoptera, Formicidae). Journal of the New York Entomological Society 61: 211 - 214" title="A new Camponotus in California apparently inhabiting live oak, Quercus sp. (Hymenoptera, Formicidae)." volume="61" year="1954">Smith 1954</bibRefCitation>
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;
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<bibRefCitation id="9AC7CE431EEBBB7031E8DB2AB4B56A2E" DOI="https://doi.org/10.1093/aesa/92.4.514" author="Gadau, J" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="37" pagination="514 - 522" refId="B20" refString="Gadau, J, Brady, SG, Ward, PS, 1999. Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae). Annals of the Entomological Society of America 92: 514 - 522, DOI: https://doi.org/10.1093/aesa/92.4.514" title="Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae)." url="https://doi.org/10.1093/aesa/92.4.514" volume="92" year="1999">Gadau et al. 1999</bibRefCitation>
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). We have examined a large series of
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|
<taxonomicName id="DCEF6F7B20E8E7E9707FB210C47C37FE" authorityName="Smith" authorityYear="1954" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus quercicola" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="quercicola">
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<emphasis id="201815B571302C376FA790B0057288ED" italics="true" pageId="0" pageNumber="37">Camponotus quercicola</emphasis>
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|
</taxonomicName>
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|
from throughout California, and we find that the type of
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<taxonomicName id="25FE8439EE0F8A71B51C4623DE34260F" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ca. laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">
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<emphasis id="5348227360E59110B7C71C481493ACE8" italics="true" pageId="0" pageNumber="37">Ca. laevigatus</emphasis>
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</taxonomicName>
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|
falls easily within the range of variation exhibited by this species.
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<bibRefCitation id="B289ABF1232732000B72D11CEB5496B1" author="Mackay, WP" journalOrPublisher="Lambert Academic Publishing, Mauritius" pageId="0" pageNumber="37" refId="B30" refString="Mackay, WP, 2019. New World carpenter ants of the hyperdiverse genus Camponotus. Volume 1. Introduction, keys to the subgenera and species complexes and the subgenus Camponotus. Lambert Academic Publishing, Mauritius" title="New World carpenter ants of the hyperdiverse genus Camponotus. Volume 1. Introduction, keys to the subgenera and species complexes and the subgenus Camponotus." year="2019">Mackay (2019</bibRefCitation>
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: 321) also stated that the male and queen of
|
|
<taxonomicName id="8AA34563CCF1D60897CBAEB4A6D119BB" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ca. quercicola" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="quercicola">
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<emphasis id="355871B55AF83CFB213E4C0DC09DA6DE" italics="true" pageId="0" pageNumber="37">Ca. quercicola</emphasis>
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|
</taxonomicName>
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|
(now
|
|
<taxonomicName id="27D5FE202853613E8373B307BEB17A7A" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ca. laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">
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<emphasis id="A8CACC6EB083D975626C45255CA26211" italics="true" pageId="0" pageNumber="37">Ca. laevigatus</emphasis>
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|
</taxonomicName>
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|
) are unknown, but this is incorrect: they were described and illustrated by
|
|
<bibRefCitation id="604A42AC5F0F5A29EF520107F83A6C51" DOI="https://doi.org/10.1093/aesa/92.4.514" author="Gadau, J" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="37" pagination="514 - 522" refId="B20" refString="Gadau, J, Brady, SG, Ward, PS, 1999. Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae). Annals of the Entomological Society of America 92: 514 - 522, DOI: https://doi.org/10.1093/aesa/92.4.514" title="Systematics, distribution, and ecology of an endemic California Camponotus quercicola (Hymenoptera: Formicidae)." url="https://doi.org/10.1093/aesa/92.4.514" volume="92" year="1999">Gadau et al. (1999)</bibRefCitation>
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and compared with related species.
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</paragraph>
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<caption id="37CE5FC4BEAB9F680FB0F937DB3B5D23" doi="10.3897/asp.79.e66978.figure7" httpUri="https://binary.pensoft.net/fig/533435" pageId="0" pageNumber="37" start="Figure 7" startId="F7">
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<paragraph id="3D9894CF4AC0FCC0427F647DF9854B26" pageId="0" pageNumber="37">
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|
<emphasis id="43862D6D1AC8C9BEE636B63278793F37" bold="true" pageId="0" pageNumber="37">Figure 7.</emphasis>
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|
<taxonomicName id="4299CDC90DE9313E097CC67AC5DF787C" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="genus">
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|
<emphasis id="807EAD4C6D1A4D4C68E7B87BACA721F3" italics="true" pageId="0" pageNumber="37">Camponotus</emphasis>
|
|
</taxonomicName>
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|
workers, full-face (dorsal) views of head (A, C, E) and lateral views of body (B, D, F); scale bars = 1.0 mm.
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|
<emphasis id="AAC4118E91E61A4BE1142905CF2E02B7" bold="true" pageId="0" pageNumber="37">A</emphasis>
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|
,
|
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<emphasis id="8247A97A3AD77993D897415789B042B3" bold="true" pageId="0" pageNumber="37">B</emphasis>
|
|
:
|
|
<taxonomicName id="446408C04E4FB94C41B32F8730782C60" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus laevigatus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevigatus">
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|
<emphasis id="6AAC8C95D66577B07283D94F0E6C6AB0" italics="true" pageId="0" pageNumber="37">Camponotus laevigatus</emphasis>
|
|
</taxonomicName>
|
|
, lectotype (CASENT0903603).
|
|
<emphasis id="8A944CE4E589EF2B939D615A8F6E0660" bold="true" pageId="0" pageNumber="37">C</emphasis>
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|
,
|
|
<emphasis id="C9C273080C8BF8749FD80CAFE974A42E" bold="true" pageId="0" pageNumber="37">D</emphasis>
|
|
:
|
|
<taxonomicName id="6B81FA543F5CE5414A4A161A2393228D" authorityName="Smith" authorityYear="1954" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus quercicola" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="quercicola">
|
|
<emphasis id="EA7C36B8EDBA3B24379FEC9B8D071C0B" italics="true" pageId="0" pageNumber="37">Camponotus quercicola</emphasis>
|
|
</taxonomicName>
|
|
, paratype (MCZ-ENT00029334), image from MCZbase (Museum of Comparative Zoology, Harvard University, copyright President and Fellows of Harvard College).
|
|
<emphasis id="55619442D142A76E38347AC542A8F68D" bold="true" pageId="0" pageNumber="37">E</emphasis>
|
|
,
|
|
<emphasis id="2BB544C57D8343ACAEEF4FDA1394D94B" bold="true" pageId="0" pageNumber="37">F</emphasis>
|
|
:
|
|
<taxonomicName id="5275A61294293DBB616180E100C905F6" baseAuthorityName="Smith" baseAuthorityYear="1859" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus laevissimus" order="Hymenoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="laevissimus">
|
|
<emphasis id="BB58AEBC4AF22613336A70A5A7926768" italics="true" pageId="0" pageNumber="37">Camponotus laevissimus</emphasis>
|
|
</taxonomicName>
|
|
(CASENT0280010). Images A, B, E, F from AntWeb (www.antweb.org); photographer Will Ericson.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
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</document> |