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<document ID-DOI="http://dx.doi.org/10.3897/zse.100.119297" ID-Pensoft-Pub="1860-0743-2-405" ID-Pensoft-UUID="EE3B0B141BBF5B9BABB2CA9EE36EC7B1" ID-ZooBank="6B07BF7A9ADA4959A65459E747E200DE" ModsDocID="1860-0743-100-2-405" checkinTime="1712716951600" checkinUser="pensoft" docAuthor="Jonishi, Taro &amp; Nakano, Takafumi" docDate="2024" docId="3631F7E75C12572785B661E1056EA4EB" docLanguage="en" docName="ZoosystEvol 100(2): 405-423" docOrigin="Zoosystematics and Evolution 100 (2)" docPubDate="2024-04-09" docSource="http://dx.doi.org/10.3897/zse.100.119297" docTitle="Scolopocryptops quadristriatus" docType="treatment" docVersion="1" id="EE3B0B141BBF5B9BABB2CA9EE36EC7B1" lastPageNumber="405" masterDocId="EE3B0B141BBF5B9BABB2CA9EE36EC7B1" masterDocTitle="Taxonomic re-appraisal of Scolopocryptops quadristriatus (Verhoeff, 1934) and a description of a new species from Japan and Taiwan (Chilopoda, Scolopendromorpha, Scolopocryptopidae)" masterLastPageNumber="423" masterPageNumber="405" pageNumber="405" updateTime="1712716951600" updateUser="pensoft">
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<mods:title id="1A8B2621D41F7D3C0B7BD389C7F9AAA4">Taxonomic re-appraisal of Scolopocryptops quadristriatus (Verhoeff, 1934) and a description of a new species from Japan and Taiwan (Chilopoda, Scolopendromorpha, Scolopocryptopidae)</mods:title>
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<mods:namePart id="E86AD39F72576173E0DDA7675E15B3AF">Jonishi, Taro</mods:namePart>
<mods:nameIdentifier id="244FFBAA947BF5153C81044747105363" type="ORCID">https://orcid.org/0000-0002-9620-0618</mods:nameIdentifier>
<mods:affiliation id="A2594B17913FDC9BEC6A6D184436E520">Department of Zoology, Graduate School of Science, Kyoto University, Kyoto 606 - 8502, Japan</mods:affiliation>
<mods:nameIdentifier id="AAA6A692B485AB57A63CE1A54AB79961" type="email">ykn347635@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5803C4D3E3BC6691CE8A62369EDC6032">Nakano, Takafumi</mods:namePart>
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<mods:affiliation id="1FBF544F6EDD86326A251A29604607F8">Department of Zoology, Graduate School of Science, Kyoto University, Kyoto 606 - 8502, Japan</mods:affiliation>
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<subSubSection id="05C4253D48C1E77A9034FA7BA77651CC" pageId="0" pageNumber="405" type="nomenclature">
<paragraph id="A2CE9AA17A06CA5906E4164F734F3822" pageId="0" pageNumber="405">
<taxonomicName id="46AE1B9AB67D9FEEA5FBDA1BDE1B28C8" LSID="3631F7E7-5C12-5727-85B6-61E1056EA4EB" authority="(Verhoeff, 1934)" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">Scolopocryptops quadristriatus (Verhoeff, 1934)</taxonomicName>
</paragraph>
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<subSubSection id="B147C6CE60E1CDAF5AAC340FB5F1E202" pageId="0" pageNumber="405" type="description">
<paragraph id="BB262E7CFA20B0791C6830DC50E2E48F" pageId="0" pageNumber="405">
<figureCitation id="C5D90EFF87113A5200899F6E47F5674C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scolopocryptops quadristriatus (Verhoeff, 1934), the likely holotype, ♀ (ZSM A 20051244). A. Cephalic capsule, maxillae, legs and left ultimate leg; B. Forcipular segment, leg-bearing segments and legs. Scale bars: 5 mm." figureDoi="10.3897/zse.100.119297.figure2" httpUri="https://binary.pensoft.net/fig/1022272" pageId="0" pageNumber="405">Figs 2</figureCitation>
<figureCitation id="1638FD62200158074962560F027ED307" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopocryptops quadristriatus (Verhoeff, 1934), collected from near the type locality, ♀ (KUZ Z 5104) and the habitat near the type locality. A. Habitat at Ome City, Tokyo; B. Live specimen, dorsal view. Scale bar: 10 mm." figureDoi="10.3897/zse.100.119297.figure3" httpUri="https://binary.pensoft.net/fig/1022273" pageId="0" pageNumber="405">, 3</figureCitation>
<figureCitation id="F5CB7AB4DD00AAD5045761E2C9A5BB5C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Basal articles of left antenna, dorsal view; B. Cephalic plate, dorsal view. Scale bars: 0.5 mm (A); 1 mm (B)." figureDoi="10.3897/zse.100.119297.figure4" httpUri="https://binary.pensoft.net/fig/1022274" pageId="0" pageNumber="405">, 4</figureCitation>
<figureCitation id="ACE6C122C138A373B7C35676578B9264" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergite 1, dorsal view; B. Distal part of article 2, article 3 and pretarsus of left second maxilla, medial view; C. Article 3 and pretarsus of left second maxilla, lateral view; D. Head, ventral view. Abbreviations: am - anterior margin of forcipular coxosternite; bs - basal suture on forcipular trochanteroprefemoral process; db - dorsal brush on article 3 of second maxilla; ds - dorsal spur on article 2 of second maxilla; pt - pretarsus of second maxilla; ptr - process of forcipular trochanteroprefemur. Scale bars: 1 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure5" httpUri="https://binary.pensoft.net/fig/1022275" pageId="0" pageNumber="405">, 5</figureCitation>
<figureCitation id="F4BD509752BF57A44415C6D66884120E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">, 6</figureCitation>
<figureCitation id="4B82B7A49BB37F75E1AA3C67DC3E9433" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality. A. KUZ Z 5095; B. KUZ Z 5094; C. KUZ Z 5104. A. Left ultimate leg, lateral view; B. Male genital segments, penis and anal valves, ventral view; C. Female genital segment and anal valves, ventral view. Abbreviations: av - anal valve; pn - penis; sgs I - sternite of genital segment 1; sgs II - sternite of genital segment 2; tg - tergite of genital segment. Scale bars: 2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure7" httpUri="https://binary.pensoft.net/fig/1022277" pageId="0" pageNumber="405">, 7 Japanese name: Yosuji-akamukade</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DB00CB21F460EFEC5629908304223047" pageId="0" pageNumber="405" type="reference_group">
<paragraph id="F22E4FDCDFCC7729A7E047BD6B53970E" pageId="0" pageNumber="405">
<taxonomicName id="E0846AEE5C3ED1170827B325C10E6C3D" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops sexspinosus subsp. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="sexspinosus" subSpecies="quadristriatus">Otocryptops sexspinosus quadristriatus</taxonomicName>
Verhoeff, 1934: 54;
<bibRefCitation id="0DCAF09B97BB9E4551D94893FAC4ABDC" author="Takakuwa, Y" journalOrPublisher="Dobutsugaku Zasshi" pageId="0" pageNumber="405" pagination="698 - 700" refId="B63" refString="Takakuwa, Y, 1939. Eine neue (dritte) art von Otocryptops aus Japan. Dobutsugaku Zasshi 51 (10): 698 - 700" title="Eine neue (dritte) art von Otocryptops aus Japan." volume="51" year="1939">Takakuwa (1939</bibRefCitation>
: 699), fig. 3 (as
<taxonomicName id="6DCF8CCDC5BA83314B31828FADBDEFB8" family="Scolopocryptopidae" genus="Octocryptops" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Octocryptops" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="genus">Octocryptops</taxonomicName>
[sic]
<taxonomicName id="363E8A7AB0B927B0F7E69DCBC5C0C3D8" authorityName="Say" authorityYear="1821" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops sexspinosus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="sexspinosus">Scolopocryptops sexspinosus</taxonomicName>
<taxonomicName id="9860442C66C39185646BF2E7095DB314" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">Scolopocryptops quadristriatus</taxonomicName>
);
<bibRefCitation id="83B931DC255020448A8CCA898CAB2B26" author="Takakuwa, Y" journalOrPublisher="Sanseido, Tokyo" pageId="0" pageNumber="405" refId="B64" refString="Takakuwa, Y, 1940. Scolopendromorpha (Class Chilopoda: Epimorpha). Fauna Nipponica, Vol. IX, Fas. VIII, No. II. Sanseido, Tokyo" title="Scolopendromorpha (Class Chilopoda: Epimorpha). Fauna Nipponica, Vol. IX, Fas. VIII, No. II." year="1940">Takakuwa (1940</bibRefCitation>
: 73), fig. 77.
</paragraph>
<paragraph id="35C7921BF6F54E25663A89C21392BE66" pageId="0" pageNumber="405">
<taxonomicName id="11C428413AEA6086780659DF2F2AA57A" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">Scolopocryptops quadristriatus</taxonomicName>
:
<bibRefCitation id="9DC3596E33B2A7F51A7228684BCBA88C" author="Shinohara, K" journalOrPublisher="Edaphologia" pageId="0" pageNumber="405" pagination="39 - 42" publicationUrl="https://agriknowledge.affrc.go.jp/RN/2010842561" refId="B55" refString="Shinohara, K, 1984. Two new species of the Scolopocryptops from Japan (Chilopoda: Cryptopidae). Edaphologia 31: 39 - 42, https://agriknowledge.affrc.go.jp/RN/2010842561" title="Two new species of the Scolopocryptops from Japan (Chilopoda: Cryptopidae)." url="https://agriknowledge.affrc.go.jp/RN/2010842561" volume="31" year="1984">Shinohara (1984</bibRefCitation>
: 41);
<bibRefCitation id="F8F52D13751DC7A599B13272BBB4FD67" author="Shinohara, K" editor="Aoki, J-I" journalOrPublisher="Tokai University Press, Hadano" pageId="0" pageNumber="405" pagination="873 - 910" refId="B57" refString="Shinohara, K, Takano, M, Ishii, K, 2015. Chilopoda. In: Aoki, J-I, Ed., Pictorial Keys to Soil Animals of Japan, Second Edition. Tokai University Press, Hadano: 873 - 910" title="Chilopoda." volumeTitle="Pictorial Keys to Soil Animals of Japan, Second Edition." year="2015">Shinohara et al. (2015</bibRefCitation>
: 880, 906);
<bibRefCitation id="6B6938B6E4328AC887F6B7685CE47630" DOI="https://doi.org/10.11646/zootaxa.5228.4.3" author="Le, SX" journalOrPublisher="Zootaxa" pageId="0" pageNumber="405" pagination="411 - 447" refId="B30" refString="Le, SX, Schileyko, AA, Nguyen, AD, 2023. A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 411 - 447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3" title="A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species." url="https://doi.org/10.11646/zootaxa.5228.4.3" volume="5228" year="2023">Le et al. (2023</bibRefCitation>
: 437, 442).
</paragraph>
<paragraph id="07BF5943728DBF1C678DC595BB279B1A" pageId="0" pageNumber="405">
<taxonomicName id="45BAAEE868766BDD43B9E6586D11356B" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">Scolopocryptops quadristriatus</taxonomicName>
Not
<taxonomicName id="076A5FF25592938E88884EAA006112CC" class="Chilopoda" family="Scolopocryptopidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="S. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="quadristriatus">S. quadristriatus</taxonomicName>
. &quot;
<taxonomicName id="2B107F9E7219E5DEAE18EF9461BFB007" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops sexspinosus subsp. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="sexspinosus" subSpecies="quadristriatus">Otocryptops sexspinosus quadristriatus</taxonomicName>
&quot;:
<bibRefCitation id="CAE446344AD22C615D334F602CEEECBF" author="Takakuwa, Y" journalOrPublisher="Shokubutsu oyobi Dobutsu" pageId="0" pageNumber="405" pagination="1457 - 1464" refId="B60" refString="Takakuwa, Y, 1933b. Scolopendromorpha of Japan. Shokubutsu oyobi Dobutsu 1 (10): 1457 - 1464" title="Scolopendromorpha of Japan." volume="1" year="1933 b">Takakuwa (1933b</bibRefCitation>
: 1457, 1459) (
<emphasis id="ABA8F7DDB579E4C0002FC8A0C94AC6C8" italics="true" pageId="0" pageNumber="405">nomen nudum</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="886A9446D14B52EA84BB87A37EECA4D3" pageId="0" pageNumber="405" type="type specimen">
<paragraph id="8E2BD3519ACDEED758900EF1F1EBEEC8" pageId="0" pageNumber="405">Type specimen.</paragraph>
<paragraph id="C40A1E91E7292A02F46C78C948D47B53" pageId="0" pageNumber="405">
<taxonomicName id="E76F9501689B7BA38D414292AAC2245D" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops s. subsp. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="B458CAC18C151340FF440AD1FF4F8827" italics="true" pageId="0" pageNumber="405">Otocryptops s. quadristriatus</emphasis>
</taxonomicName>
was described, based on a single specimen from the vicinity of Tokyo, without any information on the collector, collecting date or deposition of the specimen (
<bibRefCitation id="A57E330FD35C1B8D39403CBB5684FB36" author="Verhoeff, KW" journalOrPublisher="Zoologische Jahrbuecher Abteilung fuer Systematik" pageId="0" pageNumber="405" pagination="1 - 112" refId="B70" refString="Verhoeff, KW, 1934. Beitraege zur Systematik und Geographie der Chilopoden. Zoologische Jahrbuecher Abteilung fuer Systematik 66 (1-2): 1 - 112" title="Beitraege zur Systematik und Geographie der Chilopoden." volume="66" year="1934">Verhoeff 1934</bibRefCitation>
). Most of the
<normalizedToken id="7AD76B29B762E4A7C67FFAD222FD8518" originalValue="Verhoeff’s">Verhoeff's</normalizedToken>
specimens are now kept at the Zoologische Staatssammlung
<normalizedToken id="5412510E777FCA9B873BA8BB32EF6105" originalValue="München">Muenchen</normalizedToken>
(ZSM), the Museum
<normalizedToken id="6EE51B75E020AEBDFCC1575FBB076AEA" originalValue="für">fuer</normalizedToken>
Naturkunde (MfN) and the Naturhistorischen Museums Wien (NHMW) (
<bibRefCitation id="0BC4E97C5539D20ACFD9B8F444B69AE9" author="Melzer, RR" journalOrPublisher="Spixiana" pageId="0" pageNumber="405" pagination="11 - 20" publicationUrl="https://pfeil-verlag.de/publikationen/spixiana-zeitschrift-fuer-zoologie-inhalt-band-34/" refId="B34" refString="Melzer, RR, Friedrich, S, Ritzerfeld, M, Bohn, J, Spelda, J, 2011. GLOMYRIS and TYMUNAC: Myriapoda and Acari databases of the GBIF-D node invertebrates II. Spixiana 34 (1): 11 - 20, https://pfeil-verlag.de/publikationen/spixiana-zeitschrift-fuer-zoologie-inhalt-band-34/" title="GLOMYRIS and TYMUNAC: Myriapoda and Acari databases of the GBIF-D node invertebrates II." url="https://pfeil-verlag.de/publikationen/spixiana-zeitschrift-fuer-zoologie-inhalt-band-34/" volume="34" year="2011">Melzer et al. 2011</bibRefCitation>
). A specimen of
<taxonomicName id="CAEC05533508B88D83D9BE38CC05B75C" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. s. subsp. quadristriatus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="6652683E52A1D13B5F9F152CEC9512CB" italics="true" pageId="0" pageNumber="405">O. s. quadristriatus</emphasis>
</taxonomicName>
is deposited at the ZSM, but neither MfN nor NHMW have specimens labelled as
<taxonomicName id="390D86A542371C0DB38844B0DD56BF40" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. s. subsp. quadristriatus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="F53D91821316609480B185A534A2BD1D" italics="true" pageId="0" pageNumber="405">O. s. quadristriatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="27275D45508DFB8D63A20D3FBA1EB3FF" DOI="https://doi.org/10.1002/mmnz.4830550215" author="Moritz, M" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="0" pageNumber="405" pagination="297 - 352" refId="B37" refString="Moritz, M, Fischer, SC, 1979. Die Typen der Myriapoden-Sammlung des Zoologischen Museums Berlin. II. Chilopoda. Mitteilungen aus dem Zoologischen Museum in Berlin 55: 297 - 352, DOI: https://doi.org/10.1002/mmnz.4830550215" title="Die Typen der Myriapoden-Sammlung des Zoologischen Museums Berlin. II. Chilopoda." url="https://doi.org/10.1002/mmnz.4830550215" volume="55" year="1979">Moritz and Fischer 1979</bibRefCitation>
;
<bibRefCitation id="82FB4FED22F541C7DADCCD7EAE790B7B" author="Schileyko, A" journalOrPublisher="Journal of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="405" refId="B50" refString="Schileyko, A, Stagl, V, 2004. The collection of the scolopendromorph centipedes (Chilopoda) in the Natural History Museum in Vienna: A critical re-evaluation of former taxonomic identifications. Annalen des Naturhistorischen Museums in Wien 105B: 67-137." title="The collection of the scolopendromorph centipedes (Chilopoda) in the Natural History Museum in Vienna: A critical re-evaluation of former taxonomic identifications. Annalen des Naturhistorischen Museums in Wien 105 B: 67 - 137." year="2004">Schileyko and Stagl 2004</bibRefCitation>
). ZSM A20051244 (Fig.
<figureCitation id="82F680E52243D746592042D8A5900D11" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scolopocryptops quadristriatus (Verhoeff, 1934), the likely holotype, ♀ (ZSM A 20051244). A. Cephalic capsule, maxillae, legs and left ultimate leg; B. Forcipular segment, leg-bearing segments and legs. Scale bars: 5 mm." figureDoi="10.3897/zse.100.119297.figure2" httpUri="https://binary.pensoft.net/fig/1022272" pageId="0" pageNumber="405">2</figureCitation>
) is, thus, the only specimen of
<taxonomicName id="5185E91FFC70FEE06317878206225740" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. s. subsp. quadristriatus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="8F108E7E614D8B014ED18D29AEBF9C38" italics="true" pageId="0" pageNumber="405">O. s. quadristriatus</emphasis>
</taxonomicName>
remaining within the
<normalizedToken id="663EC944D89041BD32E1CC2E3E182498" originalValue="Verhoeff’s">Verhoeff's</normalizedToken>
collection and is supposed to be the holotype of this nominal taxon, although the original label and collection data of this specimen are unavailable (Stefan Friedrich, personal communication). Its cephalic capsule, maxillae and left ultimate leg had been preserved in a separate vial (Fig.
<figureCitation id="FAB1B9AF1C268AEF470A428698AC0511" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scolopocryptops quadristriatus (Verhoeff, 1934), the likely holotype, ♀ (ZSM A 20051244). A. Cephalic capsule, maxillae, legs and left ultimate leg; B. Forcipular segment, leg-bearing segments and legs. Scale bars: 5 mm." figureDoi="10.3897/zse.100.119297.figure2" httpUri="https://binary.pensoft.net/fig/1022272" pageId="0" pageNumber="405">2A</figureCitation>
); the body had been cut into two parts on leg-bearing segment 8 (Fig.
<figureCitation id="59A4C6EA993A6841DA589546F5F96BC5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Scolopocryptops quadristriatus (Verhoeff, 1934), the likely holotype, ♀ (ZSM A 20051244). A. Cephalic capsule, maxillae, legs and left ultimate leg; B. Forcipular segment, leg-bearing segments and legs. Scale bars: 5 mm." figureDoi="10.3897/zse.100.119297.figure2" httpUri="https://binary.pensoft.net/fig/1022272" pageId="0" pageNumber="405">2B</figureCitation>
); its ultimate leg-bearing segment had been dissected. Additionally, its left legs 6, 8, 12, 14, 15, 17, 18, 20 and 22; right legs 3, 8, 12-18 and 20-22; and right ultimate leg had been lost or loosened; its left leg 16 was loosened during observation by the first author. Morphological features of the likely holotype are consistent with the original description and specimens newly obtained in this study. Thus, a description, based on both ZSM A20051244 and our specimens, is provided below.
</paragraph>
<caption id="7A4A737C0554FCE1EB2ABA709997E1A1" doi="10.3897/zse.100.119297.figure2" httpUri="https://binary.pensoft.net/fig/1022272" pageId="0" pageNumber="405" start="Figure 2" startId="F2">
<paragraph id="F6809B522AD14907C97B74E5AEA964B6" pageId="0" pageNumber="405">
<emphasis id="DECEF172F89BE0139209965F73685196" bold="true" pageId="0" pageNumber="405">Figure 2.</emphasis>
<taxonomicName id="EBA6276BEE547F64EC48259F622A2AFA" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="1F166A3AA5C53E94DAE3EFD4E05533E2" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
(Verhoeff, 1934), the likely holotype, ♀ (ZSM A20051244).
<emphasis id="05299ADC18E68EC1BC184260BEF54E1F" bold="true" pageId="0" pageNumber="405">A.</emphasis>
Cephalic capsule, maxillae, legs and left ultimate leg;
<emphasis id="EE3A4499F9E3DF33912C080A9101E8BC" bold="true" pageId="0" pageNumber="405">B.</emphasis>
Forcipular segment, leg-bearing segments and legs. Scale bars: 5 mm.
</paragraph>
</caption>
<paragraph id="F61B7B987398BCDCB4EAB5C2FF589A5B" pageId="0" pageNumber="405">
<bibRefCitation id="B2810EA8DF8D7A912957BAE375AE25F2" author="Shinohara, K" journalOrPublisher="Atypus" pageId="0" pageNumber="405" pagination="25 - 28" refId="B54" refString="Shinohara, K, 1982. Organizing Japanese names of centipedes. Atypus 80: 25 - 28" title="Organizing Japanese names of centipedes." volume="80" year="1982">Shinohara (1982)</bibRefCitation>
stated that a Japanese myriapodologist, the late Dr Yoshioki Takakuwa, sent a specimen of
<taxonomicName id="E32D4ED516DBE64C10DE82A3574DDE7E" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. s. subsp. quadristriatus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="B1E7BB2890AC33ECA84C3EC57559CBAC" italics="true" pageId="0" pageNumber="405">O. s. quadristriatus</emphasis>
</taxonomicName>
to Verhoeff. Given that Verhoeff also received other chilopod specimens from Takakuwa (e.g.
<bibRefCitation id="CFB3343961B99E0ABB97BAB504FB431D" author="Verhoeff, KW" journalOrPublisher="Zoologische Jahrbuecher Abteilung fuer Systematik" pageId="0" pageNumber="405" pagination="1 - 112" refId="B70" refString="Verhoeff, KW, 1934. Beitraege zur Systematik und Geographie der Chilopoden. Zoologische Jahrbuecher Abteilung fuer Systematik 66 (1-2): 1 - 112" title="Beitraege zur Systematik und Geographie der Chilopoden." volume="66" year="1934">Verhoeff (1934</bibRefCitation>
,
<bibRefCitation id="5C23305E463F6A88B65763ACCB2B4051" author="Verhoeff, KW" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="405" pagination="10 - 23" refId="B71" refString="Verhoeff, KW, 1935. Ueber Scolioplanes (Chilopoda). Zoologischer Anzeiger 111: 10 - 23" title="Ueber Scolioplanes (Chilopoda)." volume="111" year="1935">1935</bibRefCitation>
,
<bibRefCitation id="C0336326E69C955FB511465549BF3690" author="Verhoeff, KW" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="405" refId="B72" refString="Verhoeff, KW, 1937. Chilopoden-studien. Zur Kenntnis der Lithobiiden. Archiv fuer Naturgeschichte N.F.6: 171-257." title="Chilopoden-studien. Zur Kenntnis der Lithobiiden. Archiv fuer Naturgeschichte N. F. 6: 171 - 257." year="1937">1937</bibRefCitation>
)), it is plausible that the likely holotype was also dispatched from Takakuwa to Verhoeff.
</paragraph>
</subSubSection>
<subSubSection id="0A68426081A0A616F315E3EE4BD5775E" pageId="0" pageNumber="405" type="materials_examined">
<paragraph id="BE44C2D14BF30DB0CC38A5DC2ABC32D8" pageId="0" pageNumber="405">Material examined.</paragraph>
<paragraph id="03C0EF3DE7E1C02D822DB8DA2BA7B90A" pageId="0" pageNumber="405">
<materialsCitation id="0960F0CE568D5BC4765118448439AF89" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="Japan" location="Japan" specimenCount="♀" specimenCount-female="1" typeStatus="Holotype">
<emphasis id="EAF24610B769BCF925F131D8FECF994D" bold="true" pageId="0" pageNumber="405">
<emphasis id="4A5BEF0947E2282DD20D097A02603621" italics="true" pageId="0" pageNumber="405">
<typeStatus id="753B37406703ACAC37188AAE5523C0A3" type="holotype">Holotype</typeStatus>
</emphasis>
(?):
<collectingCountry id="D9FE5B9C0702145BA8F346C136EA232F" name="Japan">Japan</collectingCountry>
</emphasis>
•
<specimenCount id="9DF39A919981C725099AD6A1F59FF85E" count="1" type="female">♀</specimenCount>
(approx.
<quantity id="BE39855574378C0C49FC6530ABDBB6DF" metricMagnitude="-2" metricUnit="m" metricValue="3.35" unit="mm" value="33.5">33.5 mm</quantity>
); ZSM A
<collectingDate id="758B06DF83EE9D3F4A71091EFBBD2C86" value="1244-05-20">20051244</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4C9FEB7CEEC229D8946B607407F0EB57" pageId="0" pageNumber="405" type="materials_examined">
<paragraph id="D8B15D0D4F6683B2A103CC40BF771395" pageId="0" pageNumber="405">Additional material.</paragraph>
<paragraph id="9DABDFECE9DDB0DFA8A420EB96EA117E" pageId="0" pageNumber="405">
<materialsCitation id="89F5C6B7F582FA67F46FEBEA17A15CD3" collectingDate="2017-04-15" collectorName="Taiga Kato" country="Japan" location="Uratakao-machi" municipality="Hachioji City" specimenCount="1" stateProvince="Tokyo">
<collectingCountry id="F167615F5CE7DAE64BB1A6360993D178" name="Japan">Japan</collectingCountry>
-
<emphasis id="B1DDBF0C03166EC0798F49E5D1F90006" bold="true" pageId="0" pageNumber="405">
<collectingRegion id="2E345D93480B8442EC0DC35417906504" country="Japan" name="Tokyo">Tokyo</collectingRegion>
</emphasis>
• 1,
<quantity id="8730E0317C92931BBE8B0FFC11CDE767" metricMagnitude="-2" metricUnit="m" metricValue="3.3899999999999997" unit="mm" value="33.9">33.9 mm</quantity>
(KUZ Z4083);
<collectingMunicipality id="C5EC8C992AB7620FAC4D765D07FEFC02">Hachioji City</collectingMunicipality>
,
<location id="BA2E5238658E842639B43FC4D3A9487E" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:BA2E5238658E842639B43FC4D3A9487E" country="Japan" municipality="Hachioji City" name="Uratakao-machi" stateProvince="Tokyo">Uratakao-machi</location>
;
<collectingDate id="30A9C88668EE60B8D54A4A3584592A03" value="2017-04-15">15 Apr 2017</collectingDate>
;
<collectorName id="D9F80C6F0078B78A00F63BFBF2CF0DD1">Taiga Kato</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="EFC2771BE700482120FCE9E0E67E1502" collectingDate="2017-11-18" collectorName="T. Kato" country="Japan" location="Mt. Takao-san" municipality="Hachioji City" specimenCount="1" stateProvince="Tokyo">
1,
<quantity id="589C0FF9513D961F3C4589472FB5B3BD" metricMagnitude="-2" metricUnit="m" metricValue="3.32" unit="mm" value="33.2">33.2 mm</quantity>
(KUZ Z5091); same data as for preceding • 1, 33.0 mm (KUZ Z5092);
<collectingMunicipality id="F97ED7B8604029D8CEBF3AB1B6A8BB74">Hachioji City</collectingMunicipality>
,
<location id="A1C2F120FAE345832AFC2A651F12A987" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:A1C2F120FAE345832AFC2A651F12A987" country="Japan" municipality="Hachioji City" name="Mt. Takao-san" stateProvince="Tokyo">Mt. Takao-san</location>
;
<collectingDate id="D8411BA665756EC77D758F4436BF69FD" value="2017-11-18">18 Nov 2017</collectingDate>
;
<collectorName id="094C66915E12CEE35731E4921E279031">T. Kato</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="9E2FDCADDA2D506EDADBBDFC7B980F1D" collectingDate="2021-10-18" collectorName="T. Jonishi" country="Japan" elevation="250" latitude="35.644833" location="Uratakao-machi" longLatPrecision="11" longitude="139.23933" municipality="Hachioji City" specimenCount="1" stateProvince="Tokyo">
1,
<quantity id="512D8F32513CCAB29C2D78EBAB4EB1B3" metricMagnitude="-2" metricUnit="m" metricValue="2.86" unit="mm" value="28.6">28.6 mm</quantity>
(KUZ Z5093);
<collectingMunicipality id="15320E1CFA8D199A1FFE2EC6F565F303">Hachioji City</collectingMunicipality>
,
<location id="505AFFB321F796FD7BEAE1DC8FB6209D" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:505AFFB321F796FD7BEAE1DC8FB6209D" country="Japan" latitude="35.644833" longLatPrecision="11" longitude="139.23933" municipality="Hachioji City" name="Uratakao-machi" stateProvince="Tokyo">Uratakao-machi</location>
;
<geoCoordinate id="95D0D97114928F8B7CEE428E6C45C586" degrees="35" direction="north" minutes="38.69" orientation="latitude" precision="9" value="35.644833">35°38.69'N</geoCoordinate>
,
<geoCoordinate id="8490CCACEA235E113885BC04A7765E3F" degrees="139" direction="east" minutes="14.36" orientation="longitude" precision="9" value="139.23933">139°14.36'E</geoCoordinate>
; approx.
<quantity id="ED2ADC0CE362B7A12C686DCF0CF407FC" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">
<elevation id="931C8DD210CEEBC84A3DC3A6F36A2983" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">250 m</elevation>
</quantity>
alt.;
<collectingDate id="8EE930D52D5BDBF676D8DF2FE5B5B9C9" value="2021-10-18">18 Oct 2021</collectingDate>
;
<collectorName id="347127BD996F660E8DC1F7BB9C0D0E8B">T. Jonishi</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="90E76D89473E6EC25E180216E5120A7B" collectingDate="2021-10-18" collectorName="T. Jonishi" country="Japan" elevation="260" latitude="35.645332" location="Uratakao-machi" longLatPrecision="11" longitude="139.23883" municipality="Hachioji City" specimenCount="1" specimenCount-male="1" stateProvince="Tokyo">
<specimenCount id="587F82CD2B4BF6A6D9977D8A3AB578CE" count="1" type="male">1 ♂</specimenCount>
,
<quantity id="E097FC8CE924A909B328DC1B93EC1673" metricMagnitude="-2" metricUnit="m" metricValue="3.21" unit="mm" value="32.1">32.1 mm</quantity>
(KUZ Z5094) and 3,
<quantity id="6B3B904D90770C7A1BBA15201B55D749" metricMagnitude="-2" metricUnit="m" metricValue="3.31" unit="mm" value="33.1">33.1 mm</quantity>
(KUZ Z5095),
<quantity id="84954EC746A95B3BFC49308FA5EEF526" metricMagnitude="-2" metricUnit="m" metricValue="3.62" unit="mm" value="36.2">36.2 mm</quantity>
(KUZ Z5096),
<quantity id="93D262EEB4F2602BA66D4B6DF49E904D" metricMagnitude="-2" metricUnit="m" metricValue="2.77" unit="mm" value="27.7">27.7 mm</quantity>
(KUZ Z5097); same locality as for preceding;
<geoCoordinate id="64981802D280D4CD1F55D463AFD73538" degrees="35" direction="north" minutes="38.72" orientation="latitude" precision="9" value="35.645332">35°38.72'N</geoCoordinate>
,
<geoCoordinate id="0EB9612ADB8B01C2488910597B4DB5B7" degrees="139" direction="east" minutes="14.33" orientation="longitude" precision="9" value="139.23883">139°14.33'E</geoCoordinate>
; approx.
<quantity id="25F1964937D829DDD6ECD67D57F12408" metricMagnitude="2" metricUnit="m" metricValue="2.6" unit="m" value="260.0">
<elevation id="885EA3205E935E5E0CB830108742886A" metricMagnitude="2" metricUnit="m" metricValue="2.6" unit="m" value="260.0">260 m</elevation>
</quantity>
alt.;
<collectingDate id="A6D71AEBFA2D3BC62E67B0C941E9821F" value="2021-10-18">18 Oct 2021</collectingDate>
;
<collectorName id="54D4A114D3087551D0A40C5B6D081C42">T. Jonishi</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="460D8E0773D9E41C75444845105AED60" collectingDate="2021-10-19" collectorName="T. Jonishi" country="Japan" elevation="230" latitude="35.797832" location="Futamatao" longLatPrecision="11" longitude="139.226" municipality="Ome City" specimenCount="1" stateProvince="Tokyo">
1,
<quantity id="5AA3A6D592C4A4D883506B2400F61CEC" metricMagnitude="-2" metricUnit="m" metricValue="3.86" unit="mm" value="38.6">38.6 mm</quantity>
(KUZ Z5098);
<collectingMunicipality id="797F7BA3AED8B7F57FC57B4429C8010D">Ome City</collectingMunicipality>
,
<location id="BE96817D10833EFF5088581E6B9DC511" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:BE96817D10833EFF5088581E6B9DC511" country="Japan" latitude="35.797832" longLatPrecision="11" longitude="139.226" municipality="Ome City" name="Futamatao" stateProvince="Tokyo">Futamatao</location>
;
<geoCoordinate id="12B138D781205AD9709EF5360518FB34" degrees="35" direction="north" minutes="47.87" orientation="latitude" precision="9" value="35.797832">35°47.87'N</geoCoordinate>
,
<geoCoordinate id="0E956C84DBE58F63169CA80E6BFAEF65" degrees="139" direction="east" minutes="13.56" orientation="longitude" precision="9" value="139.226">139°13.56'E</geoCoordinate>
; approx.
<quantity id="8EB37453B93BAD31B62EC2FC723877BC" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">
<elevation id="D89ED0CB6784A523D6882049978005BF" metricMagnitude="2" metricUnit="m" metricValue="2.3" unit="m" value="230.0">230 m</elevation>
</quantity>
alt.;
<collectingDate id="5F42D1A32290C416911E9D9E96ABA230" value="2021-10-19">19 Oct 2021</collectingDate>
;
<collectorName id="43738C2D52717F273726E2030DCDE5B7">T. Jonishi</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="2022FC75B535D63E6C70C52CB37EE7DD" collectingDate="2021-10-19" collectorName="T. Jonishi" country="Japan" elevation="350" latitude="35.811333" location="Sawai" longLatPrecision="11" longitude="139.18184" municipality="Ome City" specimenCount="1" specimenCount-male="1" stateProvince="Tokyo">
<specimenCount id="602E71EDDB95259380483EDDC9B3FB5E" count="1" type="male">1 ♂</specimenCount>
,
<quantity id="20DAEADA67B161D6E54AFB8955C82FF0" metricMagnitude="-2" metricUnit="m" metricValue="3.82" unit="mm" value="38.2">38.2 mm</quantity>
(KUZ Z5099);
<collectingMunicipality id="E874C0F18FA1917CC3036F51070EC34F">Ome City</collectingMunicipality>
,
<location id="CB987B531B2DC0FBF64C63EF0ABEB5DC" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:CB987B531B2DC0FBF64C63EF0ABEB5DC" country="Japan" latitude="35.811333" longLatPrecision="11" longitude="139.18184" municipality="Ome City" name="Sawai" stateProvince="Tokyo">Sawai</location>
;
<geoCoordinate id="8948943CE4422CDF3A3F41EA29B0109E" degrees="35" direction="north" minutes="48.68" orientation="latitude" precision="9" value="35.811333">35°48.68'N</geoCoordinate>
,
<geoCoordinate id="D27439CC64CBE4F4A3D771FA2216631A" degrees="139" direction="east" minutes="10.91" orientation="longitude" precision="9" value="139.18184">139°10.91'E</geoCoordinate>
; approx.
<quantity id="0C63B1C6EDEEA747AEF444D6E9755C37" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
<elevation id="63391AD3EC29654BD5EA901F6428BE01" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</elevation>
</quantity>
alt.;
<collectingDate id="D8255E79DBC6BA004AD1B223CBB0722B" value="2021-10-19">19 Oct 2021</collectingDate>
;
<collectorName id="71C1C85003CD603660893F534A76AEB4">T. Jonishi</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="21EC9D52B6E788E9B1D068DD7EF10FDE" collectingDate="2022-04-12" collectorName="Futaro Okuyama" country="Japan" elevation="250" latitude="35.738" location="Yokosawa" longLatPrecision="11" longitude="139.238" municipality="Akiruno City" specimenCount="1" specimenCount-juvenile="1" stateProvince="Tokyo">
2,
<quantity id="7B0AAF5FFD62129E763A76EC66BC966C" metricMagnitude="-2" metricUnit="m" metricValue="3.36" unit="mm" value="33.6">33.6 mm</quantity>
(KUZ Z5100),
<quantity id="5DE462CC85F9E90EE7E3CF5F4819B3DA" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" unit="mm" value="29.7">29.7 mm</quantity>
(KUZ Z5101) and
<specimenCount id="AF2EFCFB0FF465B5A1B8DFE80099E9C3" count="1" type="juvenile">1 juvenile</specimenCount>
,
<quantity id="9348EF3CAC3F7B450C3E456F11A3E1CE" metricMagnitude="-2" metricUnit="m" metricValue="1.28" unit="mm" value="12.8">12.8 mm</quantity>
(KUZ Z5102);
<collectingMunicipality id="33BE84B38021746AC5E51305B68E969E">Akiruno City</collectingMunicipality>
,
<location id="5F1AA2864EC55EFFDD1B7ABD03139798" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:5F1AA2864EC55EFFDD1B7ABD03139798" country="Japan" latitude="35.738" longLatPrecision="11" longitude="139.238" municipality="Akiruno City" name="Yokosawa" stateProvince="Tokyo">Yokosawa</location>
;
<geoCoordinate id="78D487E88FB155DC50A36C03E4B716EE" degrees="35" direction="north" minutes="44.28" orientation="latitude" precision="9" value="35.738">35°44.28'N</geoCoordinate>
,
<geoCoordinate id="7DCEE9254B0C718858D0EE8E5FB91BBB" degrees="139" direction="east" minutes="14.28" orientation="longitude" precision="9" value="139.238">139°14.28'E</geoCoordinate>
; approx.
<quantity id="1E027448ABEAE7622C38786C4F31F1C0" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">
<elevation id="3F45D94568FC9DD5F486AA184BA5E7E3" metricMagnitude="2" metricUnit="m" metricValue="2.5" unit="m" value="250.0">250 m</elevation>
</quantity>
alt.;
<collectingDate id="23D2DCAE5B61416C2546549084A28BFD" value="2022-04-12">12 Apr 2022</collectingDate>
;
<collectorName id="03291AFC9D37E15F4B3558D80CE9E205">Futaro Okuyama</collectorName>
leg.
</materialsCitation>
•
<materialsCitation id="AC23618B28E7F56445CEDF9351717353" collectingDate="2022-11-06" collectorName="T. Kato" country="Japan" elevation="270" latitude="35.645832" location="Uratakao-machi" longLatPrecision="11" longitude="139.236" municipality="Hachioji City" specimenCount="1" stateProvince="Tokyo">
1,
<quantity id="7859F8FF0A12375A7A5A3965CE74FFA9" metricMagnitude="-2" metricUnit="m" metricValue="2.96" unit="mm" value="29.6">29.6 mm</quantity>
(KUZ Z5103);
<collectingMunicipality id="AF4836B3B70A444C68D995043FAEF9D9">Hachioji City</collectingMunicipality>
,
<location id="6F9E8771043AF8CA12AAC700B7AAD33F" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:6F9E8771043AF8CA12AAC700B7AAD33F" country="Japan" latitude="35.645832" longLatPrecision="11" longitude="139.236" municipality="Hachioji City" name="Uratakao-machi" stateProvince="Tokyo">Uratakao-machi</location>
;
<geoCoordinate id="8B9C5FE03011034A668E617D6F52325E" degrees="35" direction="north" minutes="38.75" orientation="latitude" precision="9" value="35.645832">35°38.75'N</geoCoordinate>
,
<geoCoordinate id="1986A96CE9895ED49FC5AB1A293A0C2F" degrees="139" direction="east" minutes="14.16" orientation="longitude" precision="9" value="139.236">139°14.16'E</geoCoordinate>
; approx.
<quantity id="1B95C021D2B9E50722801654B7B7A43E" metricMagnitude="2" metricUnit="m" metricValue="2.7" unit="m" value="270.0">
<elevation id="74CEDBE548BE939E0BB8F32EDE71CB36" metricMagnitude="2" metricUnit="m" metricValue="2.7" unit="m" value="270.0">270 m</elevation>
</quantity>
alt.;
<collectingDate id="89D966DD6D6EE24A1137E86F25F0B1E5" value="2022-11-06">6 Nov 2022</collectingDate>
;
<collectorName id="0CC1BAF57836C8E3C756EE4EDA50FFC5">T. Kato</collectorName>
leg.
</materialsCitation>
-
<materialsCitation id="5AE1D9B237AE7F99B6CC20C0EBF7AC5A" collectingDate="2021-10-19" collectorName="T. Jonishi" country="Japan" county="Hiki County" elevation="110" latitude="36.000168" location="Hatoyama-machi" longLatPrecision="11" longitude="139.357" specimenCount="1" specimenCount-female="1" stateProvince="Saitama Prefecture">
<emphasis id="43299FD2C665F40DE31699092D40F1D8" bold="true" pageId="0" pageNumber="405">
<collectingRegion id="CFB7C269423C8FCDFF650B7B0AC82942" country="Japan" name="Saitama">Saitama Prefecture</collectingRegion>
</emphasis>
•
<specimenCount id="268781419F921C5D43F966180F6788E9" count="1" type="female">1 ♀</specimenCount>
, 39.0 mm (KUZ Z5104);
<collectingCounty id="AA63E4591689AEEF22BAD98EF7BCED34">Hiki County</collectingCounty>
,
<location id="3DCAB851A85C0D4CA14BE83C13BC2A8B" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:3DCAB851A85C0D4CA14BE83C13BC2A8B" country="Japan" county="Hiki County" latitude="36.000168" longLatPrecision="11" longitude="139.357" name="Hatoyama-machi" stateProvince="Saitama Prefecture">Hatoyama-machi</location>
;
<geoCoordinate id="97F17023160CF6C5B65F5EBD7FD4AE98" degrees="36" direction="north" minutes="0.01" orientation="latitude" precision="9" value="36.000168">36°0.01'N</geoCoordinate>
,
<geoCoordinate id="7449E93012E9742DAD3A55A2DCF0A108" degrees="139" direction="east" minutes="21.42" orientation="longitude" precision="9" value="139.357">139°21.42'E</geoCoordinate>
; approx.
<quantity id="EB19557ABD64083D1D871A3F0C41942F" metricMagnitude="2" metricUnit="m" metricValue="1.1" unit="m" value="110.0">
<elevation id="48C5F0FEB98F311635DB842B7AA8FABF" metricMagnitude="2" metricUnit="m" metricValue="1.1" unit="m" value="110.0">110 m</elevation>
</quantity>
alt.;
<collectingDate id="3CFD3DA80BAFC98C35235A054289D199" value="2021-10-19">19 Oct 2021</collectingDate>
;
<collectorName id="9D4E6D487624DC1B72453D8A0112A6CB">T. Jonishi</collectorName>
leg.
</materialsCitation>
-
<materialsCitation id="1F1FCA30F9704B8506DBC824AFF7BACA" collectingDate="2022-04-09" collectorName="Eitaro Matsushita" country="Japan" county="Minamitsuru County" elevation="670" latitude="35.513668" location="Nishikatsura-cho" longLatPrecision="11" longitude="138.84067" specimenCount="1" stateProvince="Yamanashi Prefecture">
<emphasis id="A32ABCC1A2E54EFD9B99723743778039" bold="true" pageId="0" pageNumber="405">
<collectingRegion id="E501DEF0226125B6AC4B2CDE681F8F12" country="Japan" name="Yamanashi">Yamanashi Prefecture</collectingRegion>
</emphasis>
• 1,
<quantity id="5053055DFCE8D69AE8372B0F3D76E49C" metricMagnitude="-2" metricUnit="m" metricValue="3.05" unit="mm" value="30.5">30.5 mm</quantity>
(KUZ Z5105);
<collectingCounty id="06DBF792700765AAA7C5866892E67A78">Minamitsuru County</collectingCounty>
,
<location id="D98C01C0A9854AA0F95D8022185E090A" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:D98C01C0A9854AA0F95D8022185E090A" country="Japan" county="Minamitsuru County" latitude="35.513668" longLatPrecision="11" longitude="138.84067" name="Nishikatsura-cho" stateProvince="Yamanashi Prefecture">Nishikatsura-cho</location>
;
<geoCoordinate id="02FD3C301CBBEBA4C235978EA0666296" degrees="35" direction="north" minutes="30.82" orientation="latitude" precision="9" value="35.513668">35°30.82'N</geoCoordinate>
,
<geoCoordinate id="241BFC030302F7FBD7E9B64D02FA43D7" degrees="138" direction="east" minutes="50.44" orientation="longitude" precision="9" value="138.84067">138°50.44'E</geoCoordinate>
; approx.
<quantity id="7AF2B8AF2EA9A7EE07230B1104402C64" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">
<elevation id="950AF62823AED2B3E81A56FCA30EBE6F" metricMagnitude="2" metricUnit="m" metricValue="6.7" unit="m" value="670.0">670 m</elevation>
</quantity>
alt.;
<collectingDate id="7C4B8C6CC2BBA75C380F2A29FA7D720D" value="2022-04-09">9 Apr 2022</collectingDate>
;
<collectorName id="2722C4C337DAD64D4FA3D0420EAC00B5">Eitaro Matsushita</collectorName>
leg.
</materialsCitation>
-
<materialsCitation id="36B4BDDD8BD4A0E3841971E7464B45A3" collectingDate="2022-06-02" collectorName="T. Nakano" country="Japan" elevation="910" latitude="35.265335" location="Suyama" longLatPrecision="11" longitude="138.79916" municipality="Susono City" specimenCount="1" stateProvince="Shizuoka Prefecture">
<emphasis id="ABA90A69FB27E8558E7C75CAE84AA458" bold="true" pageId="0" pageNumber="405">
<collectingRegion id="AF7B36F8DB0B020056FA7015D0F6A1D4" country="Japan" name="Shizuoka">Shizuoka Prefecture</collectingRegion>
</emphasis>
• 1,
<quantity id="6741611355F26577134F095F5C6760FF" metricMagnitude="-2" metricUnit="m" metricValue="3.04" unit="mm" value="30.4">30.4 mm</quantity>
(KUZ Z5106);
<collectingMunicipality id="EA7B496BF77A2961C7C7E61A071445B9">Susono City</collectingMunicipality>
,
<location id="55E09982305C639C7319E98E7ACA7713" LSID="urn:lsid:plazi:treatment:3631F7E75C12572785B661E1056EA4EB:55E09982305C639C7319E98E7ACA7713" country="Japan" latitude="35.265335" longLatPrecision="11" longitude="138.79916" municipality="Susono City" name="Suyama" stateProvince="Shizuoka Prefecture">Suyama</location>
;
<geoCoordinate id="C543C9634459D587ABFB643DBA2F08B0" degrees="35" direction="north" minutes="15.92" orientation="latitude" precision="9" value="35.265335">35°15.92'N</geoCoordinate>
,
<geoCoordinate id="39FFF11FC90E14A121261B62DE0EA538" degrees="138" direction="east" minutes="47.95" orientation="longitude" precision="9" value="138.79916">138°47.95'E</geoCoordinate>
; approx.
<quantity id="02863A0910272549E5C7D9A88BC69D02" metricMagnitude="2" metricUnit="m" metricValue="9.1" unit="m" value="910.0">
<elevation id="81C3DDE2195B270E769C01490AD2F8EB" metricMagnitude="2" metricUnit="m" metricValue="9.1" unit="m" value="910.0">910 m</elevation>
</quantity>
alt.;
<collectingDate id="557E31FDBA0D0F2AE097682C1EB4D129" value="2022-06-02">2 June 2022</collectingDate>
;
<collectorName id="6141B93D84C5F8CF65EE7B2F3F8F8429">T. Nakano</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="BDE62A864A9D4942BB03786D5E6A6E3A" pageId="0" pageNumber="405" type="diagnosis">
<paragraph id="F57E070F9307A62FE68FCACEE4C2F88F" pageId="0" pageNumber="405">Diagnosis.</paragraph>
<paragraph id="0BA082C6F48B59C07347727D03DB810B" pageId="0" pageNumber="405">Antenna with sparse short hairs and setae on dorsal surface of two basal articles, subsequent articles densely covered with short setae. Cephalic plate with complete lateral marginal sulci. Tergites lacking paramedian sutures, tergites 6-20 with four longitudinal keels and median depression bordered by paramedian keels.</paragraph>
</subSubSection>
<subSubSection id="C9EA3B792F8AC149DE565ABC25749C41" pageId="0" pageNumber="405" type="description">
<paragraph id="CE8B12BE7670887B1EB6D23638539204" pageId="0" pageNumber="405">Description</paragraph>
<paragraph id="5DB54B83EA2EBB7446559DB83C435D0D" pageId="0" pageNumber="405">
[
<emphasis id="8064E7AC45ADF6AFE37FF4F6DF25B68F" bold="true" pageId="0" pageNumber="405">variations given in square brackets].</emphasis>
Body length approx. 27.7-39.0 mm in 75% ethanol. Colour in life yellowish-brown with dark pigment on two basal antennal articles, purplish on subsequent articles; reddish-brown on forcipules; reddish-brown with dark pigment on anterior, lateral and posterior margins of cephalic plate, tergites 1, 22 and 23; purplish dark brown on tergites 2-21; legs and ultimate legs brownish-yellow or orange with bluish dark pigment (Fig.
<figureCitation id="ED021528DD8444BDA54D946E1391923F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scolopocryptops quadristriatus (Verhoeff, 1934), collected from near the type locality, ♀ (KUZ Z 5104) and the habitat near the type locality. A. Habitat at Ome City, Tokyo; B. Live specimen, dorsal view. Scale bar: 10 mm." figureDoi="10.3897/zse.100.119297.figure3" httpUri="https://binary.pensoft.net/fig/1022273" pageId="0" pageNumber="405">3B</figureCitation>
). Colour in ethanol slightly greenish on tergites and legs (Fig.
<figureCitation id="84A94B0EB17E8CEFB7D9B49365A4B698" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6A</figureCitation>
).
</paragraph>
<caption id="F4FD091562251DEA9C3335C67714D810" doi="10.3897/zse.100.119297.figure3" httpUri="https://binary.pensoft.net/fig/1022273" pageId="0" pageNumber="405" start="Figure 3" startId="F3">
<paragraph id="06017095ADD6F15B4774C3C5F6FC0C6E" pageId="0" pageNumber="405">
<emphasis id="9D00E958645E554F497498B221674D83" bold="true" pageId="0" pageNumber="405">Figure 3.</emphasis>
<taxonomicName id="84A072F2C53C7D6CA5D4A89E2E160166" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="81236BE04E6BFC1FFF4864A6C71CD8AC" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
(Verhoeff, 1934), collected from near the type locality, ♀ (KUZ Z5104) and the habitat near the type locality.
<emphasis id="2C33BB3F42327833E9BCB622FFC0CAB3" bold="true" pageId="0" pageNumber="405">A.</emphasis>
Habitat at Ome City, Tokyo;
<emphasis id="3E11D0E7A45076355624CBE6A0A3C46B" bold="true" pageId="0" pageNumber="405">B.</emphasis>
Live specimen, dorsal view. Scale bar: 10 mm.
</paragraph>
</caption>
<paragraph id="20606EA18CC96AD9192EBF4EC6CBC0B3" pageId="0" pageNumber="405">
Antennae 7.7-13.4 mm in length, approx. 0.2-0.35
<normalizedToken id="EE88D02E8C5E25CCE2BCA3FC19FF6CF3" originalValue="×">x</normalizedToken>
as long as body, composed of 17 articles; two basal articles with sparse short hairs and setae (sensu
<bibRefCitation id="D4882F6941DE3E9EE26BEC21F6BC299A" DOI="https://doi.org/10.3897/zookeys.69.737" author="Bonato, L" journalOrPublisher="ZooKeys" pageId="0" pageNumber="405" pagination="17 - 51" refId="B2" refString="Bonato, L, Edgecombe, GD, Lewis, JGE, Minelli, A, Pereira, LA, Shelley, RM, Zapparoli, M, 2010. A common terminology for the external anatomy of centipedes (Chilopoda). ZooKeys 69: 17 - 51, DOI: https://doi.org/10.3897/zookeys.69.737" title="A common terminology for the external anatomy of centipedes (Chilopoda)." url="https://doi.org/10.3897/zookeys.69.737" volume="69" year="2010">Bonato et al. (2010)</bibRefCitation>
) dorsally, subsequent articles densely covered with short setae (Fig.
<figureCitation id="B46A8CB1CFF8746094E54ECECBCAA5E3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Basal articles of left antenna, dorsal view; B. Cephalic plate, dorsal view. Scale bars: 0.5 mm (A); 1 mm (B)." figureDoi="10.3897/zse.100.119297.figure4" httpUri="https://binary.pensoft.net/fig/1022274" pageId="0" pageNumber="405">4A</figureCitation>
); each seta emerging from small collar. Cephalic plate as long as wide; its surface sparsely punctate with sparse minute hairs, with complete lateral margination (Figs
<figureCitation id="82514B76DFC60AF6F54139F736CE3406" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Basal articles of left antenna, dorsal view; B. Cephalic plate, dorsal view. Scale bars: 0.5 mm (A); 1 mm (B)." figureDoi="10.3897/zse.100.119297.figure4" httpUri="https://binary.pensoft.net/fig/1022274" pageId="0" pageNumber="405">4B</figureCitation>
,
<figureCitation id="BC50B506EE73015350609BAF13047FC6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergite 1, dorsal view; B. Distal part of article 2, article 3 and pretarsus of left second maxilla, medial view; C. Article 3 and pretarsus of left second maxilla, lateral view; D. Head, ventral view. Abbreviations: am - anterior margin of forcipular coxosternite; bs - basal suture on forcipular trochanteroprefemoral process; db - dorsal brush on article 3 of second maxilla; ds - dorsal spur on article 2 of second maxilla; pt - pretarsus of second maxilla; ptr - process of forcipular trochanteroprefemur. Scale bars: 1 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure5" httpUri="https://binary.pensoft.net/fig/1022275" pageId="0" pageNumber="405">5A</figureCitation>
).
</paragraph>
<caption id="B6B6961185DA86532D902EF8B89C8843" doi="10.3897/zse.100.119297.figure4" httpUri="https://binary.pensoft.net/fig/1022274" pageId="0" pageNumber="405" start="Figure 4" startId="F4">
<paragraph id="539F3EDA513301068109116F7D9C0AD0" pageId="0" pageNumber="405">
<emphasis id="73233C78322CD84A9092E99C060D3A1A" bold="true" pageId="0" pageNumber="405">Figure 4.</emphasis>
<taxonomicName id="3B3CD01ED4EB6D3301390909CB5D3CA7" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="92630627DC0F4677A2270EAED4A436CC" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
(Verhoeff, 1934) from near the type locality (KUZ Z5095).
<emphasis id="ECC17167826F14C0892C863C52DF9EC8" bold="true" pageId="0" pageNumber="405">A.</emphasis>
Basal articles of left antenna, dorsal view;
<emphasis id="4A31D8ABD40658F79E9EAD0E7E777B66" bold="true" pageId="0" pageNumber="405">B.</emphasis>
Cephalic plate, dorsal view. Scale bars: 0.5 mm (
<emphasis id="540EC3C53D4742201A02A191368881FF" bold="true" pageId="0" pageNumber="405">A</emphasis>
); 1 mm (
<emphasis id="DEDBFCEEF89978D8457EF2592848A4BA" bold="true" pageId="0" pageNumber="405">B</emphasis>
).
</paragraph>
</caption>
<caption id="681671EA5E6B620AA7BE3E40498668C4" doi="10.3897/zse.100.119297.figure5" httpUri="https://binary.pensoft.net/fig/1022275" pageId="0" pageNumber="405" start="Figure 5" startId="F5">
<paragraph id="3351D47D3BCCBF88AFE955E922482984" pageId="0" pageNumber="405">
<emphasis id="76C9A1202359766A48258B99BDABD1D4" bold="true" pageId="0" pageNumber="405">Figure 5.</emphasis>
<taxonomicName id="533DB9D1181AC9F70E4BE0D5A04EFD1F" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="CC65A91AFA92380B961E9EC3BC6F6B6A" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
(Verhoeff, 1934) from near the type locality (KUZ Z5095).
<emphasis id="974C445F50807C1350BF31D69B61726F" bold="true" pageId="0" pageNumber="405">A.</emphasis>
Cephalic plate and tergite 1, dorsal view;
<emphasis id="0F716BD3F6585AEB805648B30FA37C94" bold="true" pageId="0" pageNumber="405">B.</emphasis>
Distal part of article 2, article 3 and pretarsus of left second maxilla, medial view;
<emphasis id="C68DA282358E0467CE006CAF65FB4F40" bold="true" pageId="0" pageNumber="405">C.</emphasis>
Article 3 and pretarsus of left second maxilla, lateral view;
<emphasis id="DD7289233D40A6B85C4852BEE4777ECB" bold="true" pageId="0" pageNumber="405">D.</emphasis>
Head, ventral view. Abbreviations:
<emphasis id="72FD8F393133F14A9E4803749F00E2C6" bold="true" pageId="0" pageNumber="405">am</emphasis>
- anterior margin of forcipular coxosternite;
<emphasis id="BA1DBD57AEE4EDDF55C521BBD5B7DD44" bold="true" pageId="0" pageNumber="405">bs</emphasis>
- basal suture on forcipular trochanteroprefemoral process;
<emphasis id="130CD5DC48B14D0C528AEEA03BACA12B" bold="true" pageId="0" pageNumber="405">db</emphasis>
- dorsal brush on article 3 of second maxilla;
<emphasis id="CC643ECA1EA42B31E1E6CC8EF49B6554" bold="true" pageId="0" pageNumber="405">ds</emphasis>
- dorsal spur on article 2 of second maxilla;
<emphasis id="473520935CBF0E60177B514D7CA2A184" bold="true" pageId="0" pageNumber="405">pt</emphasis>
- pretarsus of second maxilla;
<emphasis id="D73C06EFCECCC0DB5AADABAA59254982" bold="true" pageId="0" pageNumber="405">ptr</emphasis>
- process of forcipular trochanteroprefemur. Scale bars: 1 mm (
<emphasis id="DED81A1CEA0CB70DEE633E98664E79E3" bold="true" pageId="0" pageNumber="405">A, D</emphasis>
); 0.2 mm (
<emphasis id="546EB821A55DE32F7AF1D920D0DF3149" bold="true" pageId="0" pageNumber="405">B, C</emphasis>
).
</paragraph>
</caption>
<paragraph id="1542CEE4388E3407FC54DF955047801C" pageId="0" pageNumber="405">
Second maxillae article 2 with elongated and semi-transparent [dark brown] dorsal spur distally; dorsal brush with transparent margination; pretarsus consisting of dark brown basal and semi-transparent short apical parts (Fig.
<figureCitation id="280ED946641D7F59D91E0FBB0125101C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergite 1, dorsal view; B. Distal part of article 2, article 3 and pretarsus of left second maxilla, medial view; C. Article 3 and pretarsus of left second maxilla, lateral view; D. Head, ventral view. Abbreviations: am - anterior margin of forcipular coxosternite; bs - basal suture on forcipular trochanteroprefemoral process; db - dorsal brush on article 3 of second maxilla; ds - dorsal spur on article 2 of second maxilla; pt - pretarsus of second maxilla; ptr - process of forcipular trochanteroprefemur. Scale bars: 1 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure5" httpUri="https://binary.pensoft.net/fig/1022275" pageId="0" pageNumber="405">5B, C</figureCitation>
). Forcipular coxosternite and trochanteroprefemora sparsely punctate, coxosternite with median suture and transverse sutures cross median one on anterior third of coxosternite (Fig.
<figureCitation id="57D1CA77328B440950BC46E354B9043A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergite 1, dorsal view; B. Distal part of article 2, article 3 and pretarsus of left second maxilla, medial view; C. Article 3 and pretarsus of left second maxilla, lateral view; D. Head, ventral view. Abbreviations: am - anterior margin of forcipular coxosternite; bs - basal suture on forcipular trochanteroprefemoral process; db - dorsal brush on article 3 of second maxilla; ds - dorsal spur on article 2 of second maxilla; pt - pretarsus of second maxilla; ptr - process of forcipular trochanteroprefemur. Scale bars: 1 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure5" httpUri="https://binary.pensoft.net/fig/1022275" pageId="0" pageNumber="405">5D</figureCitation>
); trochanteroprefemur with small and blunt black process with basal suture (Fig.
<figureCitation id="5D97844C3CE7830EE1E5D63BE5E9F897" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergite 1, dorsal view; B. Distal part of article 2, article 3 and pretarsus of left second maxilla, medial view; C. Article 3 and pretarsus of left second maxilla, lateral view; D. Head, ventral view. Abbreviations: am - anterior margin of forcipular coxosternite; bs - basal suture on forcipular trochanteroprefemoral process; db - dorsal brush on article 3 of second maxilla; ds - dorsal spur on article 2 of second maxilla; pt - pretarsus of second maxilla; ptr - process of forcipular trochanteroprefemur. Scale bars: 1 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure5" httpUri="https://binary.pensoft.net/fig/1022275" pageId="0" pageNumber="405">5D</figureCitation>
); anterior margin of coxosternite strongly sclerotised and slightly convex, divided into two low lobes by median diastema (Fig.
<figureCitation id="2644AE445C70E999C46C7EC2A8CE59AD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergite 1, dorsal view; B. Distal part of article 2, article 3 and pretarsus of left second maxilla, medial view; C. Article 3 and pretarsus of left second maxilla, lateral view; D. Head, ventral view. Abbreviations: am - anterior margin of forcipular coxosternite; bs - basal suture on forcipular trochanteroprefemoral process; db - dorsal brush on article 3 of second maxilla; ds - dorsal spur on article 2 of second maxilla; pt - pretarsus of second maxilla; ptr - process of forcipular trochanteroprefemur. Scale bars: 1 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure5" httpUri="https://binary.pensoft.net/fig/1022275" pageId="0" pageNumber="405">5D</figureCitation>
).
</paragraph>
<paragraph id="9FA3E4755B3DAA2F13C6239DD60260E5" pageId="0" pageNumber="405">
Tergites sparsely punctate; tergite 1 with anterior transverse suture; anterior margin overlapped by cephalic plate (Figs
<figureCitation id="DE2A3EB0A667AD84D1F09ECD271CC42D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergite 1, dorsal view; B. Distal part of article 2, article 3 and pretarsus of left second maxilla, medial view; C. Article 3 and pretarsus of left second maxilla, lateral view; D. Head, ventral view. Abbreviations: am - anterior margin of forcipular coxosternite; bs - basal suture on forcipular trochanteroprefemoral process; db - dorsal brush on article 3 of second maxilla; ds - dorsal spur on article 2 of second maxilla; pt - pretarsus of second maxilla; ptr - process of forcipular trochanteroprefemur. Scale bars: 1 mm (A, D); 0.2 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure5" httpUri="https://binary.pensoft.net/fig/1022275" pageId="0" pageNumber="405">5A</figureCitation>
,
<figureCitation id="5B5321733C21375C7C8AE97876038D58" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6A</figureCitation>
). All tergites lacking paramedian sutures; tergites 6 [5-7]-21 with longitudinal median depression bordered by paramedian keels, lateral keels present on tergites 6 [6-8]-20; median depression and keels unapparent on anterior and posterior tergites [depression and keels unapparent on all tergites in adult KUZ Z5099] (Fig.
<figureCitation id="8272238710AFB6CB885CF642951F9D4A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6A, B</figureCitation>
); tergites 6 [5-7]-21 [22] with complete or nearly complete lateral marginations (Fig.
<figureCitation id="C0549464ED343C7EB99331714288163E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6A, B</figureCitation>
); 2 or 3 short longitudinal sulci present on posterior margin of several tergites.
</paragraph>
<caption id="0E8663948C155845FC1CBABF9B16BDDA" doi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405" start="Figure 6" startId="F6">
<paragraph id="D02AD8B55BBFFD09F96D97D96CAA47B7" pageId="0" pageNumber="405">
<emphasis id="31578C5B3D221A4BC41DD43A3266C544" bold="true" pageId="0" pageNumber="405">Figure 6.</emphasis>
<taxonomicName id="5EF4E736D22581589C916FFBA2500493" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="E24CE21F9F050D97D25444C576760BA2" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
(Verhoeff, 1934) from near the type locality (KUZ Z5095).
<emphasis id="E308DA3151FA6FE8A45C7C87E9F2812F" bold="true" pageId="0" pageNumber="405">A.</emphasis>
Cephalic plate and tergites 1-13, dorso-lateral view;
<emphasis id="EC42DD45FB32A090169028D1F266E30B" bold="true" pageId="0" pageNumber="405">B.</emphasis>
Tergites 14-16, dorso-lateral view;
<emphasis id="470BF011BD30BA1E37EB0468F478B390" bold="true" pageId="0" pageNumber="405">C.</emphasis>
Sternites 8-13, ventral view;
<emphasis id="A2B98F346D1F04F4812DFF0217E92A8B" bold="true" pageId="0" pageNumber="405">D.</emphasis>
Leg-bearing segments 3-8, lateral view;
<emphasis id="BDD65801E4FB8CE11323E20981FB0C2E" bold="true" pageId="0" pageNumber="405">E.</emphasis>
Sternite of ultimate leg-bearing segment and coxopleuron, ventral view;
<emphasis id="71295EE8F4180757195283E0B561F5C2" bold="true" pageId="0" pageNumber="405">F.</emphasis>
Left coxopleuron, lateral view;
<emphasis id="E15D934F96BBAA2E8F02AC68F22178BC" bold="true" pageId="0" pageNumber="405">G.</emphasis>
Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations:
<emphasis id="B152CFAFF3B09E749EC71D2AEF79BC4C" bold="true" pageId="0" pageNumber="405">cxp</emphasis>
- coxopleural process;
<emphasis id="8D5C7626E6DEFDE7F6EA44F2D5DB94A3" bold="true" pageId="0" pageNumber="405">dm</emphasis>
- dorsal margin of ultimate pleuron;
<emphasis id="6DDFA2BBF032B507975AA3CC3383DE79" bold="true" pageId="0" pageNumber="405">mds</emphasis>
- minute dark spine on ultimate pleuron;
<emphasis id="7E270DFC366A36ECCFFDC9A1C25A74AB" bold="true" pageId="0" pageNumber="405">sp</emphasis>
- spiracle;
<emphasis id="AA62B3DBEDA7AB47608C26EC3A0F44EA" bold="true" pageId="0" pageNumber="405">tlk</emphasis>
- tergal lateral keel;
<emphasis id="761433C8D9B92234ED5B9C3E1595DFCF" bold="true" pageId="0" pageNumber="405">tm</emphasis>
- tergal margination;
<emphasis id="E277A881D49964D346A3D29127905FDF" bold="true" pageId="0" pageNumber="405">tpk</emphasis>
- tergal paramedian keel. Scale bars: 5 mm (
<emphasis id="DA91671939842CC95246EBA61541AA44" bold="true" pageId="0" pageNumber="405">A</emphasis>
); 2 mm (
<emphasis id="D105AE1DD5A1E810C21B06D853FFDB42" bold="true" pageId="0" pageNumber="405">B-D</emphasis>
); 0.5 mm (
<emphasis id="F9D5CEF58732B1C9232A6C35115E27BF" bold="true" pageId="0" pageNumber="405">E-G</emphasis>
).
</paragraph>
</caption>
<paragraph id="D0A7C4C904B3D3F93C84ECC182809C58" pageId="0" pageNumber="405">
Sternites sparsely punctate, lacking paramedian sutures (Fig.
<figureCitation id="6A3F7D985D67F94E26862C33733BE9DE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6C</figureCitation>
). Sides of sternite of ultimate leg-bearing segment converging posteriorly, posterior margin slightly concave [slightly convex in KUZ Z4083] (Fig.
<figureCitation id="663A534FA533316CFDBC158E920EDE31" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6E</figureCitation>
).
</paragraph>
<paragraph id="344F53FCE5D60D5396BBB165E2093FD6" pageId="0" pageNumber="405">
Ovoid spiracles present on leg-bearing segments 3, 5, 8, 10, 12, 14, 16, 18, 20 and 22 (Fig.
<figureCitation id="C68A017EDFC964E22293F1B9F02D81C7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6D</figureCitation>
).
</paragraph>
<paragraph id="B84F223C5D5FC98745C3CCA8E5E72C48" pageId="0" pageNumber="405">Legs almost lacking setae [sparse minute setae present in several specimens]; tarsi of legs 1-21 undivided; legs 1-20 with lateral and ventral tibial spurs and tarsal spur, leg 21 with tibial spur and tarsal spur; leg 22 with tarsal spur only; all legs with two accessory spines.</paragraph>
<paragraph id="682715D6DAFC459FBDCB2C78569CE6D3" pageId="0" pageNumber="405">
Coxopleuron approx. 1.5-1.7
<normalizedToken id="9F4DE1C75AEEAD0E1A2983D848FB479E" originalValue="×">x</normalizedToken>
as long as sternite of ultimate leg-bearing segment (Fig.
<figureCitation id="8091E1C90F66CB1753C2D0A0B27CC655" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6E, F</figureCitation>
). Dorsal margin of ultimate pleuron protruding from lateral side of tergite of ultimate leg-bearing segment, dorso-posterior margin with minute dark spine (Fig.
<figureCitation id="F25FD3856F939F54848738DD9D0BE258" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6F, G</figureCitation>
). Posterior and ventral margins of coxopleuron converging posteriorly, forming approx. 60-75° angle; coxopleural process short, tip of process pointed, slightly directed dorsally (Fig.
<figureCitation id="716026702178256DF9851883020E3AD9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6F</figureCitation>
). Surface of coxopleuron without setae, covered with various-sized coxal pores (Fig.
<figureCitation id="1A5795BFF1B5A33BA7139D84A7D9A7D7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6F</figureCitation>
). Pore-free area present on coxopleural process and dorso-posterior area of coxopleuron (Fig.
<figureCitation id="0788233CC82C00DD27E10BEF2A21E2E6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality (KUZ Z 5095). A. Cephalic plate and tergites 1 - 13, dorso-lateral view; B. Tergites 14 - 16, dorso-lateral view; C. Sternites 8 - 13, ventral view; D. Leg-bearing segments 3 - 8, lateral view; E. Sternite of ultimate leg-bearing segment and coxopleuron, ventral view; F. Left coxopleuron, lateral view; G. Tergite of ultimate leg-bearing segment, dorso-lateral view. Abbreviations: cxp - coxopleural process; dm - dorsal margin of ultimate pleuron; mds - minute dark spine on ultimate pleuron; sp - spiracle; tlk - tergal lateral keel; tm - tergal margination; tpk - tergal paramedian keel. Scale bars: 5 mm (A); 2 mm (B-D); 0.5 mm (E-G)." figureDoi="10.3897/zse.100.119297.figure6" httpUri="https://binary.pensoft.net/fig/1022276" pageId="0" pageNumber="405">6F</figureCitation>
).
</paragraph>
<paragraph id="3F4EA55BC7D10D9AB717CBD6BCD3BA4B" pageId="0" pageNumber="405">
Ultimate leg 9.0-12.4 mm in length, approx. 0.3
<normalizedToken id="4EE78336BD93DB04324A1633A2CCBE5C" originalValue="×">x</normalizedToken>
as long as body; all articles almost lacking setae [tarsi with sparse minute setae]; prefemur with two conical and pointed spinous processes, ventral process larger than dorso-medial one; pretarsus with two accessory spines (Fig.
<figureCitation id="968FE022BE4E884FFCF60CDAD2972471" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality. A. KUZ Z 5095; B. KUZ Z 5094; C. KUZ Z 5104. A. Left ultimate leg, lateral view; B. Male genital segments, penis and anal valves, ventral view; C. Female genital segment and anal valves, ventral view. Abbreviations: av - anal valve; pn - penis; sgs I - sternite of genital segment 1; sgs II - sternite of genital segment 2; tg - tergite of genital segment. Scale bars: 2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure7" httpUri="https://binary.pensoft.net/fig/1022277" pageId="0" pageNumber="405">7A</figureCitation>
).
</paragraph>
<caption id="B7837D86174BCAD65052FC92F8591228" doi="10.3897/zse.100.119297.figure7" httpUri="https://binary.pensoft.net/fig/1022277" pageId="0" pageNumber="405" start="Figure 7" startId="F7">
<paragraph id="D6FF05D8C84704CCBC308FE0C6F739C2" pageId="0" pageNumber="405">
<emphasis id="FB29B6A696D7F6ACD7E065C3523C22BE" bold="true" pageId="0" pageNumber="405">Figure 7.</emphasis>
<taxonomicName id="A93D25CB7AD7F98119F641B0485D543C" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="597CA29AC5E1763DD70444D5247AFF8C" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
(Verhoeff, 1934) from near the type locality.
<emphasis id="214DAC5A69658E73ED84AB26B3AC7945" bold="true" pageId="0" pageNumber="405">A.</emphasis>
KUZ Z5095;
<emphasis id="305E51352CDF1AD68CCFAECC2CD7F459" bold="true" pageId="0" pageNumber="405">B.</emphasis>
KUZ Z5094;
<emphasis id="D2B4BE0BEC5380C3A1B615B6A97C6429" bold="true" pageId="0" pageNumber="405">C.</emphasis>
KUZ Z5104.
<emphasis id="85C550C211EE6C73F54E90C868DB05AA" bold="true" pageId="0" pageNumber="405">A.</emphasis>
Left ultimate leg, lateral view;
<emphasis id="38AD09F2EC161ABFD1557088B91AB166" bold="true" pageId="0" pageNumber="405">B.</emphasis>
Male genital segments, penis and anal valves, ventral view;
<emphasis id="4C1E23C5443A5F3E7F1F1C777BC6EFE5" bold="true" pageId="0" pageNumber="405">C.</emphasis>
Female genital segment and anal valves, ventral view. Abbreviations:
<emphasis id="6701995E17A7D9722BAEB487C7FB06BC" bold="true" pageId="0" pageNumber="405">av</emphasis>
- anal valve;
<emphasis id="2F4D0534E2BAD7FB76920E978DA341CB" bold="true" pageId="0" pageNumber="405">pn</emphasis>
- penis;
<emphasis id="A4CFEDE0E7089BF1437FC7335993E36D" bold="true" pageId="0" pageNumber="405">sgs I</emphasis>
- sternite of genital segment 1;
<emphasis id="11E73250953B09CEC0A40A1CDE5AF5AA" bold="true" pageId="0" pageNumber="405">sgs II</emphasis>
- sternite of genital segment 2;
<emphasis id="4484C76ECBCF589BF6430C43B214ADB9" bold="true" pageId="0" pageNumber="405">tg</emphasis>
- tergite of genital segment. Scale bars: 2 mm (
<emphasis id="07D36292EBECFB459E12EE3BC5983251" bold="true" pageId="0" pageNumber="405">A</emphasis>
); 0.5 mm (
<emphasis id="A572FD62DDFAD33F7357E35B75BE5C97" bold="true" pageId="0" pageNumber="405">B, C</emphasis>
).
</paragraph>
</caption>
<paragraph id="8B1A58F0FFB9B6E5A6DA0706BBBB7ABE" pageId="0" pageNumber="405">
Genital segments occupying approx. 0.7-0.8
<normalizedToken id="93144042E9BCF432D23D4C9AEA406FF2" originalValue="×">x</normalizedToken>
length of sternite of ultimate leg-bearing segment; tergite of genital segment covered with sparse minute setae (Fig.
<figureCitation id="1010C73F9DFF7942A40E3ED6EDA63292" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality. A. KUZ Z 5095; B. KUZ Z 5094; C. KUZ Z 5104. A. Left ultimate leg, lateral view; B. Male genital segments, penis and anal valves, ventral view; C. Female genital segment and anal valves, ventral view. Abbreviations: av - anal valve; pn - penis; sgs I - sternite of genital segment 1; sgs II - sternite of genital segment 2; tg - tergite of genital segment. Scale bars: 2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure7" httpUri="https://binary.pensoft.net/fig/1022277" pageId="0" pageNumber="405">7B, C</figureCitation>
). Sternite of genital segment 1 covered with sparse short setae, posterior margin weakly convex (Fig.
<figureCitation id="BBD78B720C58EC24DF1BE5A4B4C16181" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality. A. KUZ Z 5095; B. KUZ Z 5094; C. KUZ Z 5104. A. Left ultimate leg, lateral view; B. Male genital segments, penis and anal valves, ventral view; C. Female genital segment and anal valves, ventral view. Abbreviations: av - anal valve; pn - penis; sgs I - sternite of genital segment 1; sgs II - sternite of genital segment 2; tg - tergite of genital segment. Scale bars: 2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure7" httpUri="https://binary.pensoft.net/fig/1022277" pageId="0" pageNumber="405">7B, C</figureCitation>
). Sternite of genital segment 2 well developed in male, covered with sparse short setae; posterior part of genital segment 2 overlapped by lamina subanalis, penis not visible in ventral view [penis visible in KUZ Z5094] (Fig.
<figureCitation id="5DDF1257A529C8B2F5A1C0A55B0D3EED" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality. A. KUZ Z 5095; B. KUZ Z 5094; C. KUZ Z 5104. A. Left ultimate leg, lateral view; B. Male genital segments, penis and anal valves, ventral view; C. Female genital segment and anal valves, ventral view. Abbreviations: av - anal valve; pn - penis; sgs I - sternite of genital segment 1; sgs II - sternite of genital segment 2; tg - tergite of genital segment. Scale bars: 2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure7" httpUri="https://binary.pensoft.net/fig/1022277" pageId="0" pageNumber="405">7B</figureCitation>
); genital segment 2 not visible in female (Fig.
<figureCitation id="BC491E1CFD2805D9065BED18563F14CC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality. A. KUZ Z 5095; B. KUZ Z 5094; C. KUZ Z 5104. A. Left ultimate leg, lateral view; B. Male genital segments, penis and anal valves, ventral view; C. Female genital segment and anal valves, ventral view. Abbreviations: av - anal valve; pn - penis; sgs I - sternite of genital segment 1; sgs II - sternite of genital segment 2; tg - tergite of genital segment. Scale bars: 2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure7" httpUri="https://binary.pensoft.net/fig/1022277" pageId="0" pageNumber="405">7C</figureCitation>
). Anal valves covered with sparse short setae (Fig.
<figureCitation id="9DE8C1405D31E751C6070A3F023199E9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scolopocryptops quadristriatus (Verhoeff, 1934) from near the type locality. A. KUZ Z 5095; B. KUZ Z 5094; C. KUZ Z 5104. A. Left ultimate leg, lateral view; B. Male genital segments, penis and anal valves, ventral view; C. Female genital segment and anal valves, ventral view. Abbreviations: av - anal valve; pn - penis; sgs I - sternite of genital segment 1; sgs II - sternite of genital segment 2; tg - tergite of genital segment. Scale bars: 2 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/zse.100.119297.figure7" httpUri="https://binary.pensoft.net/fig/1022277" pageId="0" pageNumber="405">7B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7A0FAE3125D2FB1A822EE09BD151583E" pageId="0" pageNumber="405" type="distribution">
<paragraph id="26EFF21818214E92F4B22CA7D5AB914B" pageId="0" pageNumber="405">Distribution.</paragraph>
<paragraph id="7580AAC19FA65F9D8544008D699DC15D" pageId="0" pageNumber="405">
This species has been recorded from Honshu and the Izu Islands and is abundant in Tokyo and adjacent areas (
<bibRefCitation id="C83CA8A72176B18658286588EB493A16" author="Takakuwa, Y" journalOrPublisher="Dobutsugaku Zasshi" pageId="0" pageNumber="405" pagination="698 - 700" refId="B63" refString="Takakuwa, Y, 1939. Eine neue (dritte) art von Otocryptops aus Japan. Dobutsugaku Zasshi 51 (10): 698 - 700" title="Eine neue (dritte) art von Otocryptops aus Japan." volume="51" year="1939">Takakuwa 1939</bibRefCitation>
,
<bibRefCitation id="582C379304C6787953DB82F8F24F4BCA" author="Takakuwa, Y" journalOrPublisher="Sanseido, Tokyo" pageId="0" pageNumber="405" refId="B64" refString="Takakuwa, Y, 1940. Scolopendromorpha (Class Chilopoda: Epimorpha). Fauna Nipponica, Vol. IX, Fas. VIII, No. II. Sanseido, Tokyo" title="Scolopendromorpha (Class Chilopoda: Epimorpha). Fauna Nipponica, Vol. IX, Fas. VIII, No. II." year="1940">1940</bibRefCitation>
;
<bibRefCitation id="0A0C794EA18D81CA1990965E23711C6C" DOI="https://doi.org/10.2476/asjaa.11.80" author="Shinohara, K" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="405" pagination="80 - 85" refId="B53" refString="Shinohara, K, 1949. Notes on centipedes collected by Mr. Fujiyama in Hachijo Island. Acta Arachnologica 11 (3-4): 80 - 85, DOI: https://doi.org/10.2476/asjaa.11.80" title="Notes on centipedes collected by Mr. Fujiyama in Hachijo Island." url="https://doi.org/10.2476/asjaa.11.80" volume="11" year="1949">Shinohara 1949</bibRefCitation>
;
<bibRefCitation id="6B82D75E820E5A6FC69474ABCA1E9A21" DOI="https://doi.org/10.2476/asjaa.13.3" author="Takashima, H" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="405" pagination="3 - 17" refId="B66" refString="Takashima, H, Shinohara, K, 1952. The centipede-fauna of the Tokyo District. Acta Arachnologica 13 (1): 3 - 17, DOI: https://doi.org/10.2476/asjaa.13.3" title="The centipede-fauna of the Tokyo District." url="https://doi.org/10.2476/asjaa.13.3" volume="13" year="1952">Takashima and Shinohara 1952</bibRefCitation>
;
<bibRefCitation id="AEAD11CA6C4BB858370E22BDAC08EEB7" author="Takano, M" journalOrPublisher="Takakuwaia" pageId="0" pageNumber="405" pagination="3 - 4" refId="B65" refString="Takano, M, 1973. Diplopoda and Chilopoda of Nii-jima island, Japan. Takakuwaia 6: 3 - 4" title="Diplopoda and Chilopoda of Nii-jima island, Japan." volume="6" year="1973">Takano 1973</bibRefCitation>
).
<bibRefCitation id="40965A5878C556946CA94FE11F507896" author="Miyosi, Y" journalOrPublisher="Collecting and Breeding" pageId="0" pageNumber="405" pagination="73 - 75" refId="B35" refString="Miyosi, Y, 1953. Myriapods of Siebold's residence (Narutaki, Nagasaki City). Supplement: myriapods from several localities in northern Kyushu. Collecting and Breeding 15 (3): 73 - 75" title="Myriapods of Siebold's residence (Narutaki, Nagasaki City). Supplement: myriapods from several localities in northern Kyushu." volume="15" year="1953">Miyosi (1953)</bibRefCitation>
recorded
<taxonomicName id="1A0DCE1C290C3A7FEE8ED58384CD763C" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" lsidName="S. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="quadristriatus">
<emphasis id="E532BCCF140FC75AAE1722A647D994F1" italics="true" pageId="0" pageNumber="405">S. quadristriatus</emphasis>
</taxonomicName>
from Nagasaki in Kyushu, but this species was not obtained during the survey conducted by the first author in Nagasaki and adjacent localities (25-27 July 2023).
</paragraph>
</subSubSection>
<subSubSection id="B7EF9F7A5409C6B9781A85BB40EA926F" pageId="0" pageNumber="405" type="availability of “quadristriatus” based on takakuwa’s works">
<paragraph id="EF1EC1EB552C0F9B001ED6A600ACF88F" pageId="0" pageNumber="405">
Availability of &quot;
<emphasis id="E5B5D94FE3905ADC419A411E42332E51" italics="true" pageId="0" pageNumber="405">quadristriatus</emphasis>
&quot; based on
<normalizedToken id="1BFAF763A75161976158922E2749ED80" originalValue="Takakuwa’s">Takakuwa's</normalizedToken>
works.
</paragraph>
<paragraph id="532095A658540033AFC940B7538D6F53" pageId="0" pageNumber="405">
The name &quot;
<taxonomicName id="3F2D3A99C5EDEFD8933906E3A0E68AFF" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops sexspinosus subsp. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="sexspinosus" subSpecies="quadristriatus">
<emphasis id="996B5D0B5278E144CD8321AD9D33D375" italics="true" pageId="0" pageNumber="405">Otocryptops sexspinosus quadristriatus</emphasis>
</taxonomicName>
&quot;, which was attributed to Verhoeff, was introduced by
<normalizedToken id="62DB23A98CC3B82BE1B54F92616D7A99" originalValue="Takakuwa’s">Takakuwa's</normalizedToken>
two works (
<bibRefCitation id="827675EDEDD7CF3D643EE8FE67FDC6C4" author="Takakuwa, Y" journalOrPublisher="Hakubutsugaku Zasshi" pageId="0" pageNumber="405" pagination="11 - 22" refId="B59" refString="Takakuwa, Y, 1933a. External morphology of Otocryptops (Miscellaneous notes on centipedes. IX.). Hakubutsugaku Zasshi 31 (49): 11 - 22" title="External morphology of Otocryptops (Miscellaneous notes on centipedes. IX.)." volume="31" year="1933 a">Takakuwa 1933a</bibRefCitation>
,
<bibRefCitation id="F139AD4B0651AF041469584D75EF7857" author="Takakuwa, Y" journalOrPublisher="Shokubutsu oyobi Dobutsu" pageId="0" pageNumber="405" pagination="1457 - 1464" refId="B60" refString="Takakuwa, Y, 1933b. Scolopendromorpha of Japan. Shokubutsu oyobi Dobutsu 1 (10): 1457 - 1464" title="Scolopendromorpha of Japan." volume="1" year="1933 b">b</bibRefCitation>
) before its formal description by Verhoeff in 1934. However, we herein decide that neither
<bibRefCitation id="A54579D734B6A79DD159ABC148EABC88" author="Takakuwa, Y" journalOrPublisher="Hakubutsugaku Zasshi" pageId="0" pageNumber="405" pagination="11 - 22" refId="B59" refString="Takakuwa, Y, 1933a. External morphology of Otocryptops (Miscellaneous notes on centipedes. IX.). Hakubutsugaku Zasshi 31 (49): 11 - 22" title="External morphology of Otocryptops (Miscellaneous notes on centipedes. IX.)." volume="31" year="1933 a">Takakuwa (1933a)</bibRefCitation>
nor
<bibRefCitation id="BE320857A582C64A4A7D8B6DB6DC9081" author="Takakuwa, Y" journalOrPublisher="Shokubutsu oyobi Dobutsu" pageId="0" pageNumber="405" pagination="1457 - 1464" refId="B60" refString="Takakuwa, Y, 1933b. Scolopendromorpha of Japan. Shokubutsu oyobi Dobutsu 1 (10): 1457 - 1464" title="Scolopendromorpha of Japan." volume="1" year="1933 b">Takakuwa (1933b)</bibRefCitation>
made the species-group name &quot;
<emphasis id="5B385DC0178674693C5AF16D1C00C9AF" italics="true" pageId="0" pageNumber="405">quadristriatus</emphasis>
&quot; available. &quot;
<taxonomicName id="03C890B8FF9CDF6415C373A98F4986AB" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops sexspinosus subsp. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="sexspinosus" subSpecies="quadristriatus">
<emphasis id="E8CD09FE261E1299CF07FFFD320A274E" italics="true" pageId="0" pageNumber="405">Otocryptops sexspinosus quadristriatus</emphasis>
</taxonomicName>
Verh&quot; first appeared in
<bibRefCitation id="CEBC2F0AFD0BAEDB76E494D377758D9F" author="Takakuwa, Y" journalOrPublisher="Hakubutsugaku Zasshi" pageId="0" pageNumber="405" pagination="11 - 22" refId="B59" refString="Takakuwa, Y, 1933a. External morphology of Otocryptops (Miscellaneous notes on centipedes. IX.). Hakubutsugaku Zasshi 31 (49): 11 - 22" title="External morphology of Otocryptops (Miscellaneous notes on centipedes. IX.)." volume="31" year="1933 a">Takakuwa (1933a</bibRefCitation>
: 11), who intended to provide general anatomical features of
<taxonomicName id="CCB70E49C888CEA547D834E994EFADFE" authorityName="Newport" authorityYear="1844" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="FFC77589BB0183ED1A5686377468D8C2" italics="true" pageId="0" pageNumber="405">Scolopocryptops</emphasis>
</taxonomicName>
(originally &quot;
<taxonomicName id="3256AE0ABC87720E74300CFC9B276B0F" authorityName="Haase" authorityYear="1887" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="B6D7AA5AD69CB6DE31A71BCD6845F22C" italics="true" pageId="0" pageNumber="405">Otocryptops</emphasis>
</taxonomicName>
&quot;). Nonetheless, the detailed morphological features and figures provided in this work were unambiguously based on
<taxonomicName id="F39A4F00E19B5F9991FA623B9AAA0697" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" lsidName="S. rubiginosus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="rubiginosus">
<emphasis id="FCC63E3635771146F7AF3E1089C8C4C6" italics="true" pageId="0" pageNumber="405">S. rubiginosus</emphasis>
</taxonomicName>
L. Koch, 1878 (referred as &quot;
<taxonomicName id="820E64BF709DBFAE1BB2831CB2653E6D" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops ruliginosus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="ruliginosus">
<emphasis id="FD360E4164FB7E31DDD5589AC50D3307" italics="true" pageId="0" pageNumber="405">Otocryptops ruliginosus</emphasis>
</taxonomicName>
&quot; [sic]). Therefore, the name
<emphasis id="B82F857B1122ABD77C44418075D584D7" italics="true" pageId="0" pageNumber="405">quadristriatus</emphasis>
in the combination of
<taxonomicName id="E4127AE32CA42056765ADABAA7429206" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops sexspinosus subsp. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="sexspinosus" subSpecies="quadristriatus">
<emphasis id="EC8F885A56146065CC6B73F997B2CB4A" italics="true" pageId="0" pageNumber="405">Otocryptops sexspinosus quadristriatus</emphasis>
</taxonomicName>
in
<bibRefCitation id="5CC9623109CAEE32A94F0EEDC9CEFCE0" author="Takakuwa, Y" journalOrPublisher="Hakubutsugaku Zasshi" pageId="0" pageNumber="405" pagination="11 - 22" refId="B59" refString="Takakuwa, Y, 1933a. External morphology of Otocryptops (Miscellaneous notes on centipedes. IX.). Hakubutsugaku Zasshi 31 (49): 11 - 22" title="External morphology of Otocryptops (Miscellaneous notes on centipedes. IX.)." volume="31" year="1933 a">Takakuwa (1933a)</bibRefCitation>
did not satisfy the provision of Article 13.1 of the International Code of Zoological Nomenclature (hereinafter, Code;
<bibRefCitation id="1B7F64A3EE32C1990B77485B45CB8D42" author="Hoang, DT" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="405" refId="B18" refString="International Commission on Zoological Nomenclature (1999) International code of zoological nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London." year="2017">International Commission on Zoological Nomenclature 1999</bibRefCitation>
) and thus is unavailable.
</paragraph>
<paragraph id="BF5424646B576B7B39DDC5F913D54D23" pageId="0" pageNumber="405">
In a synopsis of the Japanese centipedes,
<bibRefCitation id="AF2F81DB53D069F2348AFB04A39EC8A5" author="Takakuwa, Y" journalOrPublisher="Shokubutsu oyobi Dobutsu" pageId="0" pageNumber="405" pagination="1457 - 1464" refId="B60" refString="Takakuwa, Y, 1933b. Scolopendromorpha of Japan. Shokubutsu oyobi Dobutsu 1 (10): 1457 - 1464" title="Scolopendromorpha of Japan." volume="1" year="1933 b">Takakuwa (1933b</bibRefCitation>
: 1459) provided a brief taxonomic account and morphological descriptions of the subspecies referred as &quot;
<taxonomicName id="0B2A7195505C579E4D14B84AA54A4C9D" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops sexspinosus subsp. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="sexspinosus" subSpecies="quadristriatus">
<emphasis id="49C42C668B22531A385B9165B43F049C" italics="true" pageId="0" pageNumber="405">Otocryptops sexspinosus quadristriatus</emphasis>
</taxonomicName>
Verhoeff&quot;. However, Takakuwa considered that &quot;
<taxonomicName id="20DFB34B6D890C010F5968EAB3FA4333" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. s. subsp. quadristriatus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="D688DCCF338FF8F04693B68B8B09EA53" italics="true" pageId="0" pageNumber="405">O. s. quadristriatus</emphasis>
</taxonomicName>
&quot; sensu Verhoeff was indistinguishable from the nominotypical subspecies &quot;
<taxonomicName id="26D98B6C132A40F0718571FB10755DD0" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. S. subsp. sexspinosus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="S." subSpecies="sexspinosus">
<emphasis id="C027C1EECE8C73FFAA845ABDE3499F81" italics="true" pageId="0" pageNumber="405">O. S. sexspinosus</emphasis>
</taxonomicName>
&quot;; thus, he did not provide any description or definition that are purported to differentiate &quot;
<taxonomicName id="D736D6B8ED265FF42B7C8D24AB1958B7" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. s. subsp. quadristriatus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="29863C81BA38D44B87280D23F1930608" italics="true" pageId="0" pageNumber="405">O. s. quadristriatus</emphasis>
</taxonomicName>
&quot; (see Article 13.1.1 of the Code). Therefore, we conclude that the species-group name
<emphasis id="F3328F10BDA4985165EF9BB6C30B3300" italics="true" pageId="0" pageNumber="405">quadristriatus</emphasis>
in the combination of
<taxonomicName id="D22FF95AD9D7CE9BB1A65E74FE78A8B6" authorityName="Verhoeff" authorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Otocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Otocryptops sexspinosus subsp. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="sexspinosus" subSpecies="quadristriatus">
<emphasis id="6EEAFFA60E0C674450CA5C781F534038" italics="true" pageId="0" pageNumber="405">Otocryptops sexspinosus quadristriatus</emphasis>
</taxonomicName>
in
<bibRefCitation id="F8E80340EC0DC4D73EFAB55ED95FA36B" author="Takakuwa, Y" journalOrPublisher="Shokubutsu oyobi Dobutsu" pageId="0" pageNumber="405" pagination="1457 - 1464" refId="B60" refString="Takakuwa, Y, 1933b. Scolopendromorpha of Japan. Shokubutsu oyobi Dobutsu 1 (10): 1457 - 1464" title="Scolopendromorpha of Japan." volume="1" year="1933 b">Takakuwa (1933b)</bibRefCitation>
is also unavailable and the authorship of this nominal taxon is attributed to
<bibRefCitation id="91FBADF42A8BA2CAC86C330853C84A53" author="Verhoeff, KW" journalOrPublisher="Zoologische Jahrbuecher Abteilung fuer Systematik" pageId="0" pageNumber="405" pagination="1 - 112" refId="B70" refString="Verhoeff, KW, 1934. Beitraege zur Systematik und Geographie der Chilopoden. Zoologische Jahrbuecher Abteilung fuer Systematik 66 (1-2): 1 - 112" title="Beitraege zur Systematik und Geographie der Chilopoden." volume="66" year="1934">Verhoeff (1934)</bibRefCitation>
, who established
<taxonomicName id="240FBB623D46BD60071C724FA8EBCF16" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. s. subsp. quadristriatus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="E5BA8686457B3F10025D2F005C3D9E9B" italics="true" pageId="0" pageNumber="405">O. s. quadristriatus</emphasis>
</taxonomicName>
explicitly as a new subspecies. Moreover, according to
<bibRefCitation id="3DC51589C64141667A1C846046476C60" author="Shinohara, K" journalOrPublisher="Atypus" pageId="0" pageNumber="405" pagination="25 - 28" refId="B54" refString="Shinohara, K, 1982. Organizing Japanese names of centipedes. Atypus 80: 25 - 28" title="Organizing Japanese names of centipedes." volume="80" year="1982">Shinohara (1982</bibRefCitation>
,
<bibRefCitation id="3812EB3CB112E83C4E4A4FEC9F1B9BD0" DOI="https://doi.org/10.19004/pjssz.41.0_62" author="Shinohara, K" journalOrPublisher="Proceedings of the Japanese Society of Systematic Zoology" pageId="0" pageNumber="405" pagination="62 - 65" refId="B56" refString="Shinohara, K, 1990. A new species of the genus Scolopocryptops (Chilopoda: Cryptopidae) from Japan. Proceedings of the Japanese Society of Systematic Zoology 41: 62 - 65, DOI: https://doi.org/10.19004/pjssz.41.0_62" title="A new species of the genus Scolopocryptops (Chilopoda: Cryptopidae) from Japan." url="https://doi.org/10.19004/pjssz.41.0_62" volume="41" year="1990">1990</bibRefCitation>
),
<bibRefCitation id="19BF242BF1A0CA7DD598ACA0AD3542BE" author="Takakuwa, Y" journalOrPublisher="Shokubutsu oyobi Dobutsu" pageId="0" pageNumber="405" pagination="1457 - 1464" refId="B60" refString="Takakuwa, Y, 1933b. Scolopendromorpha of Japan. Shokubutsu oyobi Dobutsu 1 (10): 1457 - 1464" title="Scolopendromorpha of Japan." volume="1" year="1933 b">
<normalizedToken id="77B72D04E11956D8D7A655CE05046775" originalValue="Takakuwa’s">Takakuwa's</normalizedToken>
(1933b)
</bibRefCitation>
description of this taxon was based on specimen(s) misidentified as &quot;
<taxonomicName id="139E747CD9A00B2676B8D0F7C45ACA23" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. sexspinosus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="sexspinosus">
<emphasis id="8B324B2F7A3244AA5695498B52C5D9DF" italics="true" pageId="0" pageNumber="405">O. sexspinosus</emphasis>
</taxonomicName>
&quot;;
<taxonomicName id="FE2242CC6AAFC1BBBA008A2ECA6E3579" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. s. subsp. quadristriatus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="s." subSpecies="quadristriatus">
<emphasis id="1CDE4DFD22700C740B14D80352691535" italics="true" pageId="0" pageNumber="405">O. s. quadristriatus</emphasis>
</taxonomicName>
sensu
<bibRefCitation id="24C27108E3268D755B169E9B310EBA96" author="Takakuwa, Y" journalOrPublisher="Shokubutsu oyobi Dobutsu" pageId="0" pageNumber="405" pagination="1457 - 1464" refId="B60" refString="Takakuwa, Y, 1933b. Scolopendromorpha of Japan. Shokubutsu oyobi Dobutsu 1 (10): 1457 - 1464" title="Scolopendromorpha of Japan." volume="1" year="1933 b">Takakuwa (1933b)</bibRefCitation>
was later described as
<taxonomicName id="D5EBEBFF93BD9A7A0DC3B62A3D94C76E" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" lsidName="S. nipponicus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="nipponicus">
<emphasis id="1DFE1AAE178A50FAE66152840BCF4155" italics="true" pageId="0" pageNumber="405">S. nipponicus</emphasis>
</taxonomicName>
Shinohara, 1990 (placed in synonymy with
<taxonomicName id="DCFB09973038DE2B721AA2548BB28A8D" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" lsidName="S. spinicaudus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="spinicaudus">
<emphasis id="FA2F6420EA3925D1A490A711477D86AC" italics="true" pageId="0" pageNumber="405">S. spinicaudus</emphasis>
</taxonomicName>
by
<bibRefCitation id="5206E4F6C2BE594DFBF0ACD07A748A29" author="Shelley, RM" journalOrPublisher="Virginia Museum of Natural History Memoir 5, Virginia Museum of Natural History, Virginia" pageId="0" pageNumber="405" refId="B52" refString="Shelley, RM, 2002. A Synopsis of the North American Centipedes of the Order Scolopendromorpha (Chilopoda). Virginia Museum of Natural History Memoir 5, Virginia Museum of Natural History, Virginia" title="A Synopsis of the North American Centipedes of the Order Scolopendromorpha (Chilopoda)." year="2002">Shelley 2002</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7573B9424BA136E30DF1703999F4CDB4" pageId="0" pageNumber="405" type="remarks">
<paragraph id="852C538CB9DA17C688966D7A54E7BDAF" pageId="0" pageNumber="405">Remarks.</paragraph>
<paragraph id="71C232225008060D22B023BB238FDF25" pageId="0" pageNumber="405">
<bibRefCitation id="940DAEBC1FD5DE38C1945876AF4C5D3D" author="Verhoeff, KW" journalOrPublisher="Zoologische Jahrbuecher Abteilung fuer Systematik" pageId="0" pageNumber="405" pagination="1 - 112" refId="B70" refString="Verhoeff, KW, 1934. Beitraege zur Systematik und Geographie der Chilopoden. Zoologische Jahrbuecher Abteilung fuer Systematik 66 (1-2): 1 - 112" title="Beitraege zur Systematik und Geographie der Chilopoden." volume="66" year="1934">Verhoeff (1934)</bibRefCitation>
established this taxon as a subspecies of the North American
<taxonomicName id="E932F042276F91053E417FE5BBA6835F" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" lsidName="S. sexspinosus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="sexspinosus">
<emphasis id="FFCC06F3382E0CC87643B99581656B63" italics="true" pageId="0" pageNumber="405">S. sexspinosus</emphasis>
</taxonomicName>
, based on brief taxonomic accounts. He only described the absence of tergal paramedian sutures, the presence of four longitudinal keels on tergites and the colouration of head and leg-bearing segments.
</paragraph>
<paragraph id="F86E676FB132A4950EB5EFC559119162" pageId="0" pageNumber="405">
<bibRefCitation id="8DA6A5C7BE6C34F2BD12BD2F5FC0E5A2" author="Shinohara, K" journalOrPublisher="Edaphologia" pageId="0" pageNumber="405" pagination="39 - 42" publicationUrl="https://agriknowledge.affrc.go.jp/RN/2010842561" refId="B55" refString="Shinohara, K, 1984. Two new species of the Scolopocryptops from Japan (Chilopoda: Cryptopidae). Edaphologia 31: 39 - 42, https://agriknowledge.affrc.go.jp/RN/2010842561" title="Two new species of the Scolopocryptops from Japan (Chilopoda: Cryptopidae)." url="https://agriknowledge.affrc.go.jp/RN/2010842561" volume="31" year="1984">Shinohara (1984)</bibRefCitation>
elevated
<taxonomicName id="E9AADF7E79F3F6034C64A9118F66C43D" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="041E3BF24D7605E676C9C3B5E9E17913" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
to full species status, based on the following features: 1) cephalic marginal sulci reaching from postero-lateral margin of cephalic plate to antennae; 2) tergites without paramedian sutures and with four longitudinal keels; 3) arrangement of tibial and tarsal spurs on legs 19-23; and 4) a slightly slender &quot;general form&quot; compared with other species of the genus. It is unclear whether Shinohara compared
<taxonomicName id="89B020D544A981BF95088A28289819F8" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="AD16E4A0FB400D732AF599014423FEE8" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
with the North American
<taxonomicName id="30C6AD82FCF2377313277E5B236BC2FE" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. sexspinosus subsp. sexspinosus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="subSpecies" species="sexspinosus" subSpecies="sexspinosus">
<emphasis id="53620FD6C05B4470D71FB4803C55CCE5" italics="true" pageId="0" pageNumber="405">O. sexspinosus sexspinosus</emphasis>
</taxonomicName>
or the Japanese &quot;
<taxonomicName id="CA9A47DDA789BB37699EB8208474B28D" class="Chilopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="O. sexspinosus" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="sexspinosus">
<emphasis id="08DB376D5AACD1ADB5F2ABF3EEE56FEA" italics="true" pageId="0" pageNumber="405">O. sexspinosus</emphasis>
</taxonomicName>
&quot; (see above) and the characters of 1), 3) and 4) cannot conclusively distinguish
<taxonomicName id="5FFCE56235DC718C68B4E61F92211466" baseAuthorityName="Verhoeff" baseAuthorityYear="1934" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops quadristriatus" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="species" species="quadristriatus">
<emphasis id="4A12D8386AB055E2A0206A79D43FF864" italics="true" pageId="0" pageNumber="405">Scolopocryptops quadristriatus</emphasis>
</taxonomicName>
from other species. Nonetheless, the presence of four longitudinal keels on tergites distinguishes
<taxonomicName id="29690B9EF20D7DB00959C4397D848812" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" lsidName="S. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="quadristriatus">
<emphasis id="EC42719871221D552F910EAB6C5FC8AC" italics="true" pageId="0" pageNumber="405">S. quadristriatus</emphasis>
</taxonomicName>
from all other
<taxonomicName id="EC21160A4B3C3D11F69E679B6B350456" authorityName="Newport" authorityYear="1844" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolopocryptops" order="Diptera" pageId="0" pageNumber="405" phylum="Arthropoda" rank="genus">
<emphasis id="417942AD5CCA9B057DED3B7DB647DC21" italics="true" pageId="0" pageNumber="405">Scolopocryptops</emphasis>
</taxonomicName>
(except
<taxonomicName id="F98032634B605E92DA48380F4E88C4CF" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" lsidName="S. longisetosus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="longisetosus">
<emphasis id="2736AAFD5CCAE35CFA2688B210EE40C2" italics="true" pageId="0" pageNumber="405">S. longisetosus</emphasis>
</taxonomicName>
sp. nov.; see below). The distinctness of
<taxonomicName id="0E5A4EC59EE0D42DBBC4CAA847B23FF6" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" lsidName="S. quadristriatus" order="Scolopendromorpha" pageId="0" pageNumber="405" rank="species" species="quadristriatus">
<emphasis id="DD8A486972806C8208B2298D7747AA0F" italics="true" pageId="0" pageNumber="405">S. quadristriatus</emphasis>
</taxonomicName>
is also supported by the molecular phylogenetic analyses (Fig.
<figureCitation id="420CC8FBFDF95FA47D285495531A76E3" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Bayesian Inference tree for 2510 bp-aligned positions of the ITS 2, 28 S, 16 S and COI sequences. Real branch length is shown on the upper left. Numbers on nodes indicate ultrafast bootstrap values (UFBoot) and Bayesian posterior probabilities (BPP). An asterisk denotes the node with UFBoot = 100 % and BPP = 1.0. Locality numbers (Q 1 - Q 3 and L 1 - L 6) are shown in Fig. 1 and Table 1." figureDoi="10.3897/zse.100.119297.figure13" httpUri="https://binary.pensoft.net/fig/1022283" pageId="0" pageNumber="405">13</figureCitation>
).
</paragraph>
<paragraph id="19C8015923DD417CAD3318AF23C6D795" pageId="0" pageNumber="405">
This species is absent from, but should be added to Chilobase 2.0 (
<bibRefCitation id="DC25FA8CC7AF616F593E1D4FA6DF1FD8" author="Bonato, L" journalOrPublisher="ZooKeys" pageId="0" pageNumber="405" publicationUrl="https://chilobase.biologia.unipd.it/searches/result_genres/129" refId="B3" refString="Bonato, L, Chagas-Jr, A, Edgecombe, GD, Lewis, JGE, Minelli, A, Pereira, LA, Shelley, RM, Stoev, P, Zapparoli, M, 2016. ChiloBase 2.0-A World Catalogue of Centipedes (Chilopoda). Scolopocryptops Newport, 1844. https://chilobase.biologia.unipd.it/searches/result_genres/129" title="ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Scolopocryptops Newport, 1844." url="https://chilobase.biologia.unipd.it/searches/result_genres/129" year="2016">Bonato et al. 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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