294 lines
34 KiB
XML
294 lines
34 KiB
XML
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<mods:title id="620B11C3A8BCB11AFE2F7510CB492E7C">A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)</mods:title>
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<mods:namePart id="0842D4DEB42F2ADEA4945919F9CD660E">Anistratenko, Vitaliy V.</mods:namePart>
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<mods:affiliation id="E85B95A5A76CDDAF56F27336B259DF36">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine.</mods:affiliation>
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<mods:namePart id="0A336E3569547F34A9BAB38A7638D192">Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation id="50C2F67ADC18BAFAC2E52EBEE1544F80">Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany. tneub @ zo. jlug. de; https: // orcid. org / 0000 - 0002 - 1398 - 9941 & Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.</mods:affiliation>
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<mods:affiliation id="5E6F2447CD957762859BEDAFFA12AB5A">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine. & Department of Cainozoic Deposits, Institute of Geological Sciences, National Academy of Sciences of Ukraine, O. Gontchar Str. 55 b, 01054 Kiev, Ukraine.</mods:affiliation>
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<mods:namePart id="C96F7DC24C3FACCF2DB3DFAB3A05FA69">Kijashko, Pavel V.</mods:namePart>
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<mods:affiliation id="63ED673A12D7C267C3E358A981DF7128">Zoological Institute, Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia.</mods:affiliation>
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<mods:affiliation id="A01ACE0A54FABF302B6628037E428C0C">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands. & Department of Earth Sciences, Utrecht University, Budapestlaan 17, 3584 CD Utrecht, The Netherlands</mods:affiliation>
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<treatment id="03BF87A3C851FFA7FF3DFD0FFE8DA2B5" ID-DOI="http://doi.org/10.5281/zenodo.4559932" ID-GBIF-Taxon="179906620" ID-Zenodo-Dep="4559932" LSID="urn:lsid:plazi:treatment:03BF87A3C851FFA7FF3DFD0FFE8DA2B5" httpUri="http://treatment.plazi.org/id/03BF87A3C851FFA7FF3DFD0FFE8DA2B5" lastPageNumber="167" pageId="16" pageNumber="167">
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<paragraph id="8BA936B5C851FFA7FF3DFD0FFBC0A79C" blockId="16.[151,1125,721,783]" box="[151,1125,721,748]" pageId="16" pageNumber="167">
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<heading id="D0E181D9C851FFA7FF3DFD0FFBC0A79C" bold="true" box="[151,1125,721,748]" fontSize="11" level="1" pageId="16" pageNumber="167" reason="1">
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<emphasis id="B962EAA7C851FFA7FF3DFD0FFBC0A79C" bold="true" box="[151,1125,721,748]" pageId="16" pageNumber="167">
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<taxonomicName id="4C164D36C851FFA7FF3DFD0FFE60A79B" ID-CoL="7QPHF" baseAuthorityName="Starobogatov" baseAuthorityYear="1992" box="[151,453,721,748]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="brotzkajae">
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<emphasis id="B962EAA7C851FFA7FF3DFD0FFE60A79B" bold="true" box="[151,453,721,748]" italics="true" pageId="16" pageNumber="167">Clathrocaspia brotzkajae</emphasis>
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</taxonomicName>
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(Starobogatov in Anistratenko & Prisjazhnjuk, 1992)
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</emphasis>
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</paragraph>
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<subSubSection id="C30C653EC851FFA7FF3DFD2BFF7CA678" box="[151,217,757,783]" pageId="16" pageNumber="167" type="description">
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<paragraph id="8BA936B5C851FFA7FF3DFD2BFF7CA678" blockId="16.[151,1125,721,783]" box="[151,217,757,783]" pageId="16" pageNumber="167">
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<figureCitation id="132D2A30C851FFA7FF3DFD2BFF7CA678" box="[151,217,757,783]" captionStart="FIGURE 8" captionStartId="18.[151,250,1328,1352]" captionTargetBox="[151,1436,181,1304]" captionTargetId="figure-22@18.[151,1436,181,1304]" captionTargetPageId="18" captionText="FIGURE 8. Clathrocaspia brotzkajae (Starobogatov in Anistratenko & Prisjazhnjuk, 1992) from the Middle Caspian Basin. (a, b) Holotype, locality 10 (ZIN #1). (c) Paratype, locality 12 (ZIN, no number). (d, k, l) Paratype, locality 12 (ZIN, no number). (e) Paratype, locality 12 (ZIN, no number). (f) Paratype, locality 12 (ZIN, no number). (g–i) Specimen labelled as “Caspia alighadzhievi, holotype”, locality 6 (ZIN, no number). (j, m) Specimen labelled as “Caspia alighadzhievi, paratype”, locality 11 (ZIN, no number). Scale bars = 1 mm (a–k), 100 μm (l, m)." figureDoi="http://doi.org/10.5281/zenodo.4550168" httpUri="https://zenodo.org/record/4550168/files/figure.png" pageId="16" pageNumber="167">Fig. 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C653EC851FFA7FF3DFCE4FB46A6D9" pageId="16" pageNumber="167" type="reference_group">
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<paragraph id="8BA936B5C851FFA7FF3DFCE4FB46A6D9" blockId="16.[151,1404,826,942]" pageId="16" pageNumber="167">
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<taxonomicName id="4C164D36C851FFA7FF3DFCE4FE71A625" ID-CoL="7PFDQ" authorityName="De Cristofori & Jan" authorityYear="1832" box="[151,468,826,850]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="genus">
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<emphasis id="B962EAA7C851FFA7FF3DFCE4FF4DA625" box="[151,232,826,850]" italics="true" pageId="16" pageNumber="167">Pyrgula</emphasis>
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[(
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<emphasis id="B962EAA7C851FFA7FF55FCE4FEE2A626" box="[255,327,826,849]" italics="true" pageId="16" pageNumber="167">Caspia</emphasis>
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)]
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<emphasis id="B962EAA7C851FFA7FEF4FCE4FE71A625" box="[350,468,826,850]" italics="true" pageId="16" pageNumber="167">knipowitchi</emphasis>
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</taxonomicName>
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[sic] (Mak.)—
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<bibRefCitation id="EF874B44C851FFA7FDC0FCE4FC58A625" author="Logvinenko, B. M. & Starobogatov, Ya. I." box="[618,1021,826,850]" pageId="16" pageNumber="167" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. & Starobogatov, Ya. I. (1969 [" 1968 "]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. & Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko & Starobogatov 1969: 379</bibRefCitation>
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, fig. 367(8) [non
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<bibRefCitation id="EF874B44C851FFA7FB05FCE4FAEBA625" author="Makarov, A. K." box="[1199,1358,826,850]" pageId="16" pageNumber="167" pagination="1055 - 1062" refId="ref32169" refString="Makarov, A. K. (1938) Rasprostraneniye nekotorykh rakoobraznykh (Mysidacea, Cumacea) i limannykh mollyuskov v ust'yakh i otkrytykh limanakh Severnogo Prichernomor'ya. Zoologicheskiy Zhurnal, 17 (6), 1055 - 1062." type="journal article" year="1938">Makarov, 1938</bibRefCitation>
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||
].
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<taxonomicName id="4C164D36C851FFA7FF3DFC87FD21A607" ID-CoL="7QKT4" authority="Starobogatov" authorityName="Starobogatov" authorityYear="2021" box="[151,644,856,881]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="brotzkajae" status="sp. nov." subGenus="Clathrocaspia">
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<emphasis id="B962EAA7C851FFA7FF3DFC87FF7AA607" box="[151,223,857,880]" italics="true" pageId="16" pageNumber="167">Caspia</emphasis>
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(
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<emphasis id="B962EAA7C851FFA7FF44FC86FEDBA607" box="[238,382,856,880]" italics="true" pageId="16" pageNumber="167">Clathrocaspia</emphasis>
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)
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<emphasis id="B962EAA7C851FFA7FE27FC86FE52A607" box="[397,503,856,880]" italics="true" pageId="16" pageNumber="167">brotzkajae</emphasis>
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Starobogatov
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</taxonomicName>
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<taxonomicNameLabel id="A25157DCC851FFA7FD20FC87FD1BA607" box="[650,702,856,880]" pageId="16" pageNumber="167" rank="species">sp. n.</taxonomicNameLabel>
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—Starobogatov in Anistratenko & Prisjazhnjuk 1992: 18–19, fig. 2a.
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<taxonomicName id="4C164D36C851FFA7FF3DFCA6FE78A6F8" ID-CoL="7QKT4" authority="Starobogatov" authorityName="Starobogatov" authorityYear="2021" box="[151,477,887,911]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="brotzkajae">
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<emphasis id="B962EAA7C851FFA7FF3DFCA6FEEAA6F8" box="[151,335,887,911]" italics="true" pageId="16" pageNumber="167">Caspia brotzkajae</emphasis>
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Starobogatov
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</taxonomicName>
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in Anistratenko et Prisjazhnjuk, 1992—
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<bibRefCitation id="EF874B44C851FFA7FCDEFCA9FB23A6F8" author="Kantor, Yu. I. & Sysoev, A. V." box="[884,1158,887,911]" pageId="16" pageNumber="167" refId="ref31309" refString="Kantor, Yu. I. & Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor & Sysoev 2006: 87</bibRefCitation>
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, pl. 41, fig. G.
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<taxonomicName id="4C164D36C851FFA7FF3DFC48FE3DA6D9" ID-CoL="7QPHF" baseAuthorityName="Starobogatov" baseAuthorityYear="1992" box="[151,408,918,942]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="brotzkajae">
|
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<emphasis id="B962EAA7C851FFA7FF3DFC48FE3DA6D9" box="[151,408,918,942]" italics="true" pageId="16" pageNumber="167">Clathrocaspia brotzkajae</emphasis>
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</taxonomicName>
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||
(Starobogatov in Anistratenko & Prisjazhnjuk, 1992)—
|
||
<bibRefCitation id="EF874B44C851FFA7FC67FC48FB7AA6D9" author="Wesselingh, F. P. & Neubauer, T. A. & Anistratenko, V. V. & Vinarski, M. V. & Yanina, T. & ter Poorten, J. J. & Kijashko, P. V. & Albrecht, C. & Anistratenko, O. Yu. & D'Hont, A. & Frolov, P. & Martinez Gandara, A. & Gittenberger, A. & Gogaladze, A. & Karpinsky, M. & Lattuada, M. & Popa, L. & Sands, A. F. & van de Velde, S. & Vandendorpe, J. & Wilke, T." box="[973,1247,918,942]" pageId="16" pageNumber="167" pagination="31 - 124" refId="ref34307" refString="Wesselingh, F. P., Neubauer, T. A., Anistratenko, V. V., Vinarski, M. V., Yanina, T., ter Poorten, J. J., Kijashko, P. V., Albrecht, C., Anistratenko, O. Yu., D'Hont, A., Frolov, P., Martinez Gandara, A., Gittenberger, A., Gogaladze, A., Karpinsky, M., Lattuada, M., Popa, L., Sands, A. F., van de Velde, S., Vandendorpe, J. & Wilke, T. (2019) Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys, 827, 31 - 124. https: // doi. org / 10.3897 / zookeys. 827.31365" type="journal article" year="2019">
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||
Wesselingh
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||
<emphasis id="B962EAA7C851FFA7FBE0FC49FBD2A6D9" box="[1098,1143,918,942]" italics="true" pageId="16" pageNumber="167">et al</emphasis>
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||
. 2019: 69
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||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C30C653EC851FFA7FF3DFC02FB34A061" pageId="16" pageNumber="167" type="materials_examined">
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<paragraph id="8BA936B5C851FFA7FF3DFC02FE15A114" blockId="16.[151,1437,988,1986]" pageId="16" pageNumber="167">
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<emphasis id="B962EAA7C851FFA7FF3DFC02FEEDA681" bold="true" box="[151,328,988,1014]" pageId="16" pageNumber="167">Type material.</emphasis>
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<typeStatus id="54AD8817C851FFA7FEFBFC02FE1CA681" box="[337,441,988,1014]" pageId="16" pageNumber="167" type="holotype">Holotype</typeStatus>
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(
|
||
<collectionCode id="ED07AE70C851FFA7FE61FC02FE5EA681" box="[459,507,988,1014]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="16" pageNumber="167" type="Museum">ZIN</collectionCode>
|
||
#
|
||
<quantity id="4CEE9B50C851FFA7FDB7FC02FDEFA681" box="[541,586,988,1014]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="167" unit="in" value="1.0">1 in</quantity>
|
||
systematic catalogue) and
|
||
<specimenCount id="9D10FD3CC851FFA7FC2EFC02FBB0A680" box="[900,1045,988,1015]" pageId="16" pageNumber="167" type="generic" typeStatus="paratypes">74 paratypes</specimenCount>
|
||
(
|
||
<collectionCode id="ED07AE70C851FFA7FB8CFC02FBF3A681" box="[1062,1110,988,1014]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="16" pageNumber="167" type="Museum">ZIN</collectionCode>
|
||
collection). Four specimens labelled as
|
||
<typeStatus id="54AD8817C851FFA7FEB2FBDFFE26A16C" box="[280,387,1025,1051]" pageId="16" pageNumber="167" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C164D36C851FFA7FE06FBDEFDDCA16D" authorityName="Starobogatov" authorityYear="2021" box="[428,633,1024,1050]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="brotzkajae">
|
||
<emphasis id="B962EAA7C851FFA7FE06FBDEFDDCA16D" box="[428,633,1024,1050]" italics="true" pageId="16" pageNumber="167">Caspia brotzkajae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED07AE70C851FFA7FD23FBDEFD75A16D" box="[649,720,1024,1050]" pageId="16" pageNumber="167">IZAN</collectionCode>
|
||
, unnumbered lot) from a late Holocene drilling borehole taken in the southern part of
|
||
<collectingRegion id="49D2F857C851FFA7FE3CFBFAFD9DA149" box="[406,568,1060,1086]" country="Ukraine" name="Odesa" pageId="16" pageNumber="167">Odessa region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3017625C851FFA7FDE9FBFAFD38A149" box="[579,669,1060,1086]" name="Ukraine" pageId="16" pageNumber="167">Ukraine</collectingCountry>
|
||
(
|
||
<tableCitation id="C694030EC851FFA7FD06FBFAFCA4A149" box="[684,769,1060,1086]" captionStart="TABLE 1" captionStartId="3.[151,239,473,499]" captionTargetPageId="3" captionText="TABLE 1. Locality information of the studied material. Coordinates of collecting sites are given based on data in museum catalogue cards, labels and corresponding publications. Asterisks mark localities with approximate geographic coordinates based on indirect data provided in the ZIN catalogue. Abbreviations: q.—quadrant, st.—station." httpUri="http://table.plazi.org/id/DF69663DC842FFB4FF3DFE07F8C6A760" pageId="16" pageNumber="167" tableUuid="DF69663DC842FFB4FF3DFE07F8C6A760">Table 1</tableCitation>
|
||
, locality 32), do not belong to that species but instead to
|
||
<taxonomicName id="4C164D36C851FFA7FA29FBFAFE89A115" baseAuthorityName="Makarov" baseAuthorityYear="1938" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="knipowitschii">
|
||
<emphasis id="B962EAA7C851FFA7FA29FBFAFE89A115" italics="true" pageId="16" pageNumber="167">C. knipowitschii</emphasis>
|
||
</taxonomicName>
|
||
(see there).
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C851FFA7FF6DFBB2FCBDA1DD" blockId="16.[151,1437,988,1986]" pageId="16" pageNumber="167">
|
||
<emphasis id="B962EAA7C851FFA7FF6DFBB2FEC3A1F1" bold="true" box="[199,358,1132,1158]" pageId="16" pageNumber="167">Type locality.</emphasis>
|
||
<materialsCitation id="3B7E3CE8C851FFA7FEC4FBB2FCB0A1DD" ID-GBIF-Occurrence="3042095301" country="Russia" elevation="68" location="Caspian Sea" pageId="16" pageNumber="167" specimenCount="1" stateProvince="Dagestan" typeStatus="paratype">
|
||
<location id="8EC9606EC851FFA7FEC4FBB2FE52A1F1" LSID="urn:lsid:plazi:treatment:03BF87A3C851FFA7FF3DFD0FFE8DA2B5:8EC9606EC851FFA7FEC4FBB2FE52A1F1" box="[366,503,1132,1158]" country="Russia" name="Caspian Sea" pageId="16" pageNumber="167" stateProvince="Dagestan">Caspian Sea</location>
|
||
shores of
|
||
<collectingRegion id="49D2F857C851FFA7FDC4FBB3FD7CA1F0" box="[622,729,1133,1159]" country="Russia" name="Dagestan" pageId="16" pageNumber="167">Dagestan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3017625C851FFA7FD4FFBB2FC97A1F1" box="[741,818,1132,1158]" name="Russia" pageId="16" pageNumber="167">Russia</collectingCountry>
|
||
, at a depth of ca.
|
||
<quantity id="4CEE9B50C851FFA7FC50FBB2FBFBA1F0" box="[1018,1118,1132,1159]" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="16" pageNumber="167" unit="m" value="67.5" valueMax="75.0" valueMin="60.0">
|
||
<elevation id="003BD186C851FFA7FC50FBB2FBFBA1F0" box="[1018,1118,1132,1159]" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="16" pageNumber="167" unit="m" value="67.5" valueMax="75.0" valueMin="60.0">60–75 m</elevation>
|
||
</quantity>
|
||
(locality 10). The
|
||
<typeStatus id="54AD8817C851FFA7FA9BFBB3FA39A1F0" box="[1329,1436,1133,1159]" pageId="16" pageNumber="167" type="paratype">paratypes</typeStatus>
|
||
were retrieved from locality 12 close by the type locality
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C851FFA7FF6DFB6AFB34A061" blockId="16.[151,1437,988,1986]" pageId="16" pageNumber="167">
|
||
<materialsCitation id="3B7E3CE8C851FFA7FF6DFB6AFB28A061" ID-GBIF-Occurrence="3042095340" country="Russia" county="Caspian Sea" elevation="60" location="Starobogatov's" municipality="Derbent" pageId="16" pageNumber="167" specimenCount="17" stateProvince="Dagestan">
|
||
<emphasis id="B962EAA7C851FFA7FF6DFB6AFE21A1B9" bold="true" box="[199,388,1204,1230]" pageId="16" pageNumber="167">Other material.</emphasis>
|
||
<specimenCount id="9D10FD3CC851FFA7FE27FB6AFD8DA1B9" box="[397,552,1204,1230]" pageId="16" pageNumber="167" type="generic">17 specimens</specimenCount>
|
||
in
|
||
<location id="8EC9606EC851FFA7FDFAFB6AFD5DA1B9" LSID="urn:lsid:plazi:treatment:03BF87A3C851FFA7FF3DFD0FFE8DA2B5:8EC9606EC851FFA7FDFAFB6AFD5DA1B9" box="[592,760,1204,1230]" country="Russia" county="Caspian Sea" municipality="Derbent" name="Starobogatov's" pageId="16" pageNumber="167" stateProvince="Dagestan">Starobogatov’s</location>
|
||
collection, marked as “
|
||
<typeStatus id="54AD8817C851FFA7FBA3FB6AFB99A1B9" box="[1033,1084,1204,1230]" pageId="16" pageNumber="167">type</typeStatus>
|
||
material” of “
|
||
<taxonomicName id="4C164D36C851FFA7FB4EFB6AFF7DA185" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="alighadzhievi">
|
||
<emphasis id="B962EAA7C851FFA7FB4EFB6AFF7DA185" italics="true" pageId="16" pageNumber="167">Caspia alighadzhievi</emphasis>
|
||
</taxonomicName>
|
||
”, an unavailable catalogue name; collected in 1957 and 1960 near
|
||
<collectingMunicipality id="6BCDACCFC851FFA7FC7AFB06FB8FA185" box="[976,1066,1240,1266]" pageId="16" pageNumber="167">Derbent</collectingMunicipality>
|
||
(
|
||
<collectingRegion id="49D2F857C851FFA7FB90FB06FB02A185" box="[1082,1191,1240,1266]" country="Russia" name="Dagestan" pageId="16" pageNumber="167">Dagestan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3017625C851FFA7FB19FB06FB5AA184" box="[1203,1279,1240,1267]" name="Russia" pageId="16" pageNumber="167">Russia</collectingCountry>
|
||
) at a depth of
|
||
<quantity id="4CEE9B50C851FFA7FF3DFB22FF75A060" box="[151,208,1276,1303]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="167" unit="m" value="60.0">
|
||
<elevation id="003BD186C851FFA7FF3DFB22FF75A060" box="[151,208,1276,1303]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="167" unit="m" value="60.0">60 m</elevation>
|
||
</quantity>
|
||
and in the eastern part of the middle
|
||
<collectingCounty id="62C84E39C851FFA7FDD8FB22FD5EA061" box="[626,763,1276,1302]" pageId="16" pageNumber="167">Caspian Sea</collectingCounty>
|
||
at a depth of
|
||
<quantity id="4CEE9B50C851FFA7FC3EFB22FC68A060" box="[916,973,1276,1303]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="16" pageNumber="167" unit="m" value="73.0">
|
||
<elevation id="003BD186C851FFA7FC3EFB22FC68A060" box="[916,973,1276,1303]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="16" pageNumber="167" unit="m" value="73.0">73 m</elevation>
|
||
</quantity>
|
||
(localities 6, 11)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC851FFA7FF6DFAFEFCB9A365" pageId="16" pageNumber="167" type="description">
|
||
<paragraph id="8BA936B5C851FFA7FF6DFAFEFCB9A365" blockId="16.[151,1437,988,1986]" pageId="16" pageNumber="167">
|
||
<emphasis id="B962EAA7C851FFA7FF6DFAFEFEFCA04D" bold="true" box="[199,345,1312,1338]" pageId="16" pageNumber="167">Description.</emphasis>
|
||
Broadly ovoid shell with about 4 whorls and large body whorl (ratio of body whorl height/shell height approximately ¾). Protoconch consists of about 1.2 whorls that measure c. 400 μm, bearing irregular wrin-kles and faint spiral threads. Teleoconch covered by fine but distinct reticulate sculpture, with numerous spiral keels that are equally prominent across whorl profile and have interspaces that are approximately as wide as keels; only uppermost 1–2 keels are usually more prominent and slightly wider spaced; in some shells, uppermost keel forms weak subsutural ramp that results in slightly stepped outline. Aperture wide, sometimes laterally expanded; umbilicus covered in most shells, sometimes present as thin slit.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC851FFA7FF6DF9C2FAFDA20D" pageId="16" pageNumber="167" type="discussion">
|
||
<paragraph id="8BA936B5C851FFA7FF6DF9C2FADBA32D" blockId="16.[151,1437,988,1986]" pageId="16" pageNumber="167">
|
||
<emphasis id="B962EAA7C851FFA7FF6DF9C2FE9EA341" bold="true" box="[199,315,1564,1590]" pageId="16" pageNumber="167">Remarks.</emphasis>
|
||
Compared to other
|
||
<taxonomicName id="4C164D36C851FFA7FD8BF9C2FD65A341" authorityName="Lindholm" authorityYear="1930" box="[545,704,1564,1590]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B962EAA7C851FFA7FD8BF9C2FD65A341" box="[545,704,1564,1590]" italics="true" pageId="16" pageNumber="167">Clathrocaspia</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<taxonomicName id="4C164D36C851FFA7FC83F9C2FC65A341" authorityName="Starobogatov" authorityYear="2021" box="[809,960,1564,1590]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="brotzkajae">
|
||
<emphasis id="B962EAA7C851FFA7FC83F9C2FC65A341" box="[809,960,1564,1590]" italics="true" pageId="16" pageNumber="167">C. brotzkajae</emphasis>
|
||
</taxonomicName>
|
||
is more bulbous, has a larger body whorl and a more expanded aperture. Only
|
||
<taxonomicName id="4C164D36C851FFA7FD9CF99EFD7CA32D" baseAuthorityName="Golikov & Starobogatov" baseAuthorityYear="1966" box="[566,729,1600,1626]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="logvinenkoi">
|
||
<emphasis id="B962EAA7C851FFA7FD9CF99EFD7CA32D" box="[566,729,1600,1626]" italics="true" pageId="16" pageNumber="167">C. logvinenkoi</emphasis>
|
||
</taxonomicName>
|
||
from the Black Sea Basin is even more bulbous but smaller.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C851FFA7FF6DF9BAFD5CA278" blockId="16.[151,1437,988,1986]" pageId="16" pageNumber="167">
|
||
Originally, four specimens from the Holocene of the Danube Delta,
|
||
<collectingCountry id="F3017625C851FFA7FC73F9BAFB93A309" box="[985,1078,1636,1662]" name="Ukraine" pageId="16" pageNumber="167">Ukraine</collectingCountry>
|
||
, were attributed to that species (Anistratenko & Prisjazhnjuk 1992). These shells are more slender and have a smaller aperture; they rather resemble
|
||
<taxonomicName id="4C164D36C851FFA7FF3DF972FEE9A3B1" baseAuthorityName="Makarov" baseAuthorityYear="1938" box="[151,332,1708,1734]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="knipowitschii">
|
||
<emphasis id="B962EAA7C851FFA7FF3DF972FEE9A3B1" box="[151,332,1708,1734]" italics="true" pageId="16" pageNumber="167">C. knipowitschii</emphasis>
|
||
</taxonomicName>
|
||
and are referred to this species herein. In turn, the specimen illustrated as “
|
||
<taxonomicName id="4C164D36C851FFA7FB3BF972FAD1A3B1" box="[1169,1396,1708,1734]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="knipowitchi">
|
||
<emphasis id="B962EAA7C851FFA7FB3BF972FAD1A3B1" box="[1169,1396,1708,1734]" italics="true" pageId="16" pageNumber="167">Pyrgula knipowitchi</emphasis>
|
||
</taxonomicName>
|
||
” in
|
||
<bibRefCitation id="EF874B44C851FFA7FF3DF90EFD83A39C" author="Logvinenko, B. M. & Starobogatov, Ya. I." box="[151,550,1744,1771]" pageId="16" pageNumber="167" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. & Starobogatov, Ya. I. (1969 [" 1968 "]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. & Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko & Starobogatov (1969)</bibRefCitation>
|
||
from the Caspian Sea shows a very bulky shell that is highly reminiscent of and considered conspecific with
|
||
<taxonomicName id="4C164D36C851FFA7FE7DF92AFD57A279" baseAuthorityName="Starobogatov" baseAuthorityYear="1992" box="[471,754,1780,1806]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="brotzkajae">
|
||
<emphasis id="B962EAA7C851FFA7FE7DF92AFD57A279" box="[471,754,1780,1806]" italics="true" pageId="16" pageNumber="167">Clathrocaspia brotzkajae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C851FFA7FF6DF8C7FAFDA20D" blockId="16.[151,1437,988,1986]" pageId="16" pageNumber="167">
|
||
In the collection of Starobogatov we found the “
|
||
<typeStatus id="54AD8817C851FFA7FD48F8C6FCECA245" box="[738,841,1816,1842]" pageId="16" pageNumber="167" type="holotype">holotype</typeStatus>
|
||
” and 16 “
|
||
<typeStatus id="54AD8817C851FFA7FC1CF8C6FB82A245" box="[950,1063,1816,1842]" pageId="16" pageNumber="167" type="paratype">paratypes</typeStatus>
|
||
” of a catalogue taxon Starobogatov named “
|
||
<taxonomicName id="4C164D36C851FFA7FE89F8E2FDB4A221" box="[291,529,1852,1878]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="alighadzhievi">
|
||
<emphasis id="B962EAA7C851FFA7FE89F8E2FDB4A221" box="[291,529,1852,1878]" italics="true" pageId="16" pageNumber="167">Caspia alighadzhievi</emphasis>
|
||
</taxonomicName>
|
||
”. The material shows close similarities with the
|
||
<typeStatus id="54AD8817C851FFA7FB97F8E2FB3AA221" box="[1085,1183,1852,1878]" pageId="16" pageNumber="167" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C164D36C851FFA7FB6DF8E2FAFBA221" baseAuthorityName="Starobogatov" baseAuthorityYear="1992" box="[1223,1374,1852,1878]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="167" phylum="Mollusca" rank="species" species="brotzkajae">
|
||
<emphasis id="B962EAA7C851FFA7FB6DF8E2FAFBA221" box="[1223,1374,1852,1878]" italics="true" pageId="16" pageNumber="167">C. brotzkajae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="132D2A30C851FFA7FAC5F8E2FF01A20D" captionStart="FIGURE 8" captionStartId="18.[151,250,1328,1352]" captionTargetBox="[151,1436,181,1304]" captionTargetId="figure-22@18.[151,1436,181,1304]" captionTargetPageId="18" captionText="FIGURE 8. Clathrocaspia brotzkajae (Starobogatov in Anistratenko & Prisjazhnjuk, 1992) from the Middle Caspian Basin. (a, b) Holotype, locality 10 (ZIN #1). (c) Paratype, locality 12 (ZIN, no number). (d, k, l) Paratype, locality 12 (ZIN, no number). (e) Paratype, locality 12 (ZIN, no number). (f) Paratype, locality 12 (ZIN, no number). (g–i) Specimen labelled as “Caspia alighadzhievi, holotype”, locality 6 (ZIN, no number). (j, m) Specimen labelled as “Caspia alighadzhievi, paratype”, locality 11 (ZIN, no number). Scale bars = 1 mm (a–k), 100 μm (l, m)." figureDoi="http://doi.org/10.5281/zenodo.4550168" httpUri="https://zenodo.org/record/4550168/files/figure.png" pageId="16" pageNumber="167">Fig. 8</figureCitation>
|
||
g–j, m), described from the same region, and we consider Starobogatov’s material to belong to that species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC851FFA7FF6DF85AFE8DA2B5" pageId="16" pageNumber="167" type="distribution">
|
||
<paragraph id="8BA936B5C851FFA7FF6DF85AFE8DA2B5" blockId="16.[151,1437,988,1986]" pageId="16" pageNumber="167">
|
||
<emphasis id="B962EAA7C851FFA7FF6DF85AFEC4A2E9" bold="true" box="[199,353,1924,1950]" pageId="16" pageNumber="167">Distribution.</emphasis>
|
||
Known only from the Caspian Sea shores of
|
||
<collectingRegion id="49D2F857C851FFA7FCC2F85BFC76A2E8" box="[872,979,1925,1951]" country="Russia" name="Dagestan" pageId="16" pageNumber="167">Dagestan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3017625C851FFA7FC4AF85AFB8BA2E9" box="[992,1070,1924,1950]" name="Russia" pageId="16" pageNumber="167">Russia</collectingCountry>
|
||
. No living specimens have been encountered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |