170 lines
20 KiB
XML
170 lines
20 KiB
XML
<document id="9A7A7FE6A08F9B83A3DC41557D47054A" ID-DOI="10.11646/zootaxa.5493.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="13330255" ID-ZooBank="16BD6CF7-E0FF-4A04-9ADE-C3DFBF92B7DC" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1723803345649" checkinUser="plazi" docAuthor="Anichtchenko, Alexander & Sciaky, Riccardo" docDate="2024" docId="03B3FE717725FFEB74FD4261A9CFEE23" docLanguage="en" docName="zootaxa.5493.4.2.pdf" docOrigin="Zootaxa 5493 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5493.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mimocolliuris Liebke 1933" docType="treatment" docVersion="3" lastPageNumber="333" masterDocId="FF8A86097720FFEE746A4027AA6FEB24" masterDocTitle="Contribution to the knowledge of the Philippines species of the tribe Odacanthini Laporte, 1834 (Coleoptera: Carabidae)" masterLastPageNumber="344" masterPageNumber="328" pageNumber="333" updateTime="1730486879022" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:titleInfo id="BA5FF67C02CB58A9E581B10855014A23">
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<mods:title id="5A5C530800C16E472942CA983345F111">Contribution to the knowledge of the Philippines species of the tribe Odacanthini Laporte, 1834 (Coleoptera: Carabidae)</mods:title>
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<mods:name id="43AD8FA2168DC37DC29EBE694A7F4AC3" type="personal">
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<mods:namePart id="0584AE5F2993DD5D8BE6F0BCB8209FE8">Anichtchenko, Alexander</mods:namePart>
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<mods:namePart id="98313989D754D21304085F16E2404716">Sciaky, Riccardo</mods:namePart>
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<mods:title id="42D6FB83F4E2BD72D5344B0D8D51BD9D">Zootaxa</mods:title>
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<mods:part id="F3AF231FABF402EEEFAE879205D8938E">
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<mods:date id="6362A9467223933B729B11333700FC5E">2024</mods:date>
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<mods:number id="12DB394C438B0D001287BC3DAA569610">2024-08-13</mods:number>
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<mods:number id="4D9C5A7A7FD3DE6E79AE660CDB5EC55F">5493</mods:number>
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<mods:url id="1B3AB257590F1462347DE6B2971A552F">http://dx.doi.org/10.11646/zootaxa.5493.4.2</mods:url>
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<mods:identifier id="512AF56A743128099622133FFD3C67BC" type="DOI">10.11646/zootaxa.5493.4.2</mods:identifier>
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<treatment id="03B3FE717725FFEB74FD4261A9CFEE23" ID-DOI="http://doi.org/10.5281/zenodo.13330186" ID-Zenodo-Dep="13330186" LSID="urn:lsid:plazi:treatment:03B3FE717725FFEB74FD4261A9CFEE23" httpUri="http://treatment.plazi.org/id/03B3FE717725FFEB74FD4261A9CFEE23" lastPageNumber="333" pageId="5" pageNumber="333">
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<subSubSection id="C3001CEC7725FFEB74FD4261A82EE944" box="[151,577,582,609]" pageId="5" pageNumber="333" type="nomenclature">
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<paragraph id="8BA54F677725FFEB74FD4261A82EE944" blockId="5.[151,577,582,609]" box="[151,577,582,609]" pageId="5" pageNumber="333">
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<heading id="D0EDF80B7725FFEB74FD4261A82EE944" bold="true" box="[151,577,582,609]" fontSize="11" level="1" pageId="5" pageNumber="333" reason="1">
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<emphasis id="B96E93757725FFEB74FD4261A82EE944" bold="true" box="[151,577,582,609]" pageId="5" pageNumber="333">
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Genus
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<taxonomicName id="4C1A34E47725FFEB74844261A82EE944" ID-CoL="8KVVT" authority="Liebke, 1933" authorityName="Liebke" authorityYear="1933" box="[238,577,582,609]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="genus">
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<emphasis id="B96E93757725FFEB74844261ABF6E945" bold="true" box="[238,409,582,609]" italics="true" pageId="5" pageNumber="333">Mimocolliuris</emphasis>
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<bibRefCitation id="EF8B32967725FFEB75CB4261A82EE944" author="Liebke, M." box="[417,577,582,608]" pageId="5" pageNumber="333" pagination="202 - 210" refId="ref9920" refString="Liebke, M. (1933) Neue Colliurinen (Coleopt., Carab.). Stylops, 2 (9), 202 - 210. https: // doi. org / 10.1111 / j. 1365 - 3113.1993. tb 01002. x" type="journal article" year="1933">Liebke, 1933</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3001CEC7725FFEB74FD42ABA9FEE987" box="[151,913,651,676]" pageId="5" pageNumber="333" type="type_taxon">
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<paragraph id="8BA54F677725FFEB74FD42ABA9FEE987" blockId="5.[151,913,651,676]" box="[151,913,651,676]" pageId="5" pageNumber="333">
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<typeStatus id="54A1F1C57725FFEB74FD42ABAAA6E980" box="[151,201,652,676]" pageId="5" pageNumber="333">Type</typeStatus>
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species:
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<taxonomicName id="4C1A34E47725FFEB754C42ACA8F5E987" authority="Boheman, 1858" authorityName="Boheman" authorityYear="1858" box="[294,666,651,675]" class="Insecta" family="Carabidae" genus="Ophionea" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="species" species="chaudoiri">
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<emphasis id="B96E93757725FFEB754C42ACAB9BE987" box="[294,500,651,675]" italics="true" pageId="5" pageNumber="333">Ophionea chaudoiri</emphasis>
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Boheman, 1858
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</taxonomicName>
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by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3001CEC7725FFEB74FD42F5A9CFEE23" pageId="5" pageNumber="333" type="discussion">
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<paragraph id="8BA54F677725FFEB74FD42F5AE76E8E7" blockId="5.[151,1437,721,1288]" pageId="5" pageNumber="333">
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A genus with relatively low diversity, containing 17 species of rather uniform morphology.
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<bibRefCitation id="EF8B32967725FFEB70CA42F6AFF7E9CF" author="Zhao, D. & Tian, M." box="[1184,1432,721,747]" pageId="5" pageNumber="333" pagination="119 - 125" refId="ref10189" refString="Zhao, D. & Tian, M. (2010) Key to species of the genus Mimocolliuris Liebke (Coleoptera, Carabidae, Odacanthini), with description of a new species from southern China. The Pan-Pacific Entomologist, 86 (4), 119 - 125. https: // doi. org / 10.3956 / 2010 - 23" type="journal article" year="2010">Zhao and Tian (2010)</bibRefCitation>
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,
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<bibRefCitation id="EF8B32967725FFEB74FD42D2AB40E82B" author="Baehr, M." box="[151,303,757,783]" pageId="5" pageNumber="333" pagination="25 - 51" refId="ref9386" refString="Baehr, M. (2016) New species and new records of odacanthine carabid beetles from the Oriental, Papuan, and Australian Regions (Coleoptera, Carabidae, Odacanthini). Entomologische Blatter und Coleoptera, 112 (1), 25 - 51." type="journal article" year="2016">Baehr (2016)</bibRefCitation>
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, and more recently Anagha
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<emphasis id="B96E93757725FFEB760542D1A8C6E82B" box="[623,681,757,783]" italics="true" pageId="5" pageNumber="333">et al.</emphasis>
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(2024) have proposed keys to determine the species in this group, all of them, at least partly, inadequate.
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<bibRefCitation id="EF8B32967725FFEB7624433EA92CE817" author="Zhao, D. & Tian, M." box="[590,835,793,819]" pageId="5" pageNumber="333" pagination="119 - 125" refId="ref10189" refString="Zhao, D. & Tian, M. (2010) Key to species of the genus Mimocolliuris Liebke (Coleoptera, Carabidae, Odacanthini), with description of a new species from southern China. The Pan-Pacific Entomologist, 86 (4), 119 - 125. https: // doi. org / 10.3956 / 2010 - 23" type="journal article" year="2010">Zhao and Tian (2010)</bibRefCitation>
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provide a vast distribution of
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<taxonomicName id="4C1A34E47725FFEB70F3433DAF7EE817" baseAuthorityName="Nietner" baseAuthorityYear="1858" box="[1177,1297,793,819]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="species" species="pilifera">
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<emphasis id="B96E93757725FFEB70F3433DAF7EE817" box="[1177,1297,793,819]" italics="true" pageId="5" pageNumber="333">M. pilifera</emphasis>
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</taxonomicName>
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(
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<collectingCountry id="F30D0FF77725FFEB714A433EAFF7E817" box="[1312,1432,793,819]" name="Cambodia" pageId="5" pageNumber="333">Cambodia</collectingCountry>
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,
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<collectingCountry id="F30D0FF77725FFEB74FD4319AABFE87C" box="[151,208,830,856]" name="Laos" pageId="5" pageNumber="333">Laos</collectingCountry>
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,
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<collectingCountry id="F30D0FF77725FFEB74B0431AAB53E873" box="[218,316,829,855]" name="Vietnam" pageId="5" pageNumber="333">Vietnam</collectingCountry>
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,
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<collectingCountry id="F30D0FF77725FFEB752F4319ABDBE87C" box="[325,436,830,856]" name="Myanmar" pageId="5" pageNumber="333">Myanmar</collectingCountry>
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,
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<collectingCountry id="F30D0FF77725FFEB75D7431AA84CE873" box="[445,547,829,855]" name="Thailand" pageId="5" pageNumber="333">Thailand</collectingCountry>
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,
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<collectingCountry id="F30D0FF77725FFEB7644431AA8F2E873" box="[558,669,829,855]" name="Sri Lanka" pageId="5" pageNumber="333">Sri Lanka</collectingCountry>
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,
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<collectingCountry id="F30D0FF77725FFEB76CD431AA88AE873" box="[679,741,829,855]" name="India" pageId="5" pageNumber="333">India</collectingCountry>
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), without justifying it;
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<bibRefCitation id="EF8B32967725FFEB778A431AAE1BE873" author="Baehr, M." box="[992,1140,829,855]" pageId="5" pageNumber="333" pagination="25 - 51" refId="ref9386" refString="Baehr, M. (2016) New species and new records of odacanthine carabid beetles from the Oriental, Papuan, and Australian Regions (Coleoptera, Carabidae, Odacanthini). Entomologische Blatter und Coleoptera, 112 (1), 25 - 51." type="journal article" year="2016">Baehr (2016)</bibRefCitation>
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mentions the characters of the same species on the basis of the original description, but without examining ant specimen. Eventually, Anagha
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<emphasis id="B96E93757725FFEB74FD43A1AAA0E8BB" box="[151,207,901,927]" italics="true" pageId="5" pageNumber="333">et al.</emphasis>
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(2024) combined the two preceding keys without providing new data, but only repeated the distribution of
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<taxonomicName id="4C1A34E47725FFEB711443A1AA86E8E7" baseAuthorityName="Nietner" baseAuthorityYear="1858" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="species" species="pilifera">
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<emphasis id="B96E93757725FFEB711443A1AA86E8E7" italics="true" pageId="5" pageNumber="333">M. pilifera</emphasis>
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</taxonomicName>
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given by
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<bibRefCitation id="EF8B32967725FFEB7533438EA820E8E7" author="Zhao, D. & Tian, M." box="[345,591,937,963]" pageId="5" pageNumber="333" pagination="119 - 125" refId="ref10189" refString="Zhao, D. & Tian, M. (2010) Key to species of the genus Mimocolliuris Liebke (Coleoptera, Carabidae, Odacanthini), with description of a new species from southern China. The Pan-Pacific Entomologist, 86 (4), 119 - 125. https: // doi. org / 10.3956 / 2010 - 23" type="journal article" year="2010">Zhao and Tian (2010)</bibRefCitation>
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, with no more evidence or new material.
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</paragraph>
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<paragraph id="8BA54F677725FFEB74AD43EAA9CFEE23" blockId="5.[151,1437,721,1288]" pageId="5" pageNumber="333">
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<taxonomicName id="4C1A34E47725FFEB74AD43EAA831E8C3" ID-CoL="6RJZT" authority="Habu, 1979" authorityName="Habu" authorityYear="1979" box="[199,606,973,999]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="species" species="insulana">
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<emphasis id="B96E93757725FFEB74AD43EAABA0E8C3" box="[199,463,973,999]" italics="true" pageId="5" pageNumber="333">Mimocolliuris insulana</emphasis>
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<bibRefCitation id="EF8B32967725FFEB75B343EAA831E8C3" author="Habu, A." box="[473,606,973,999]" pageId="5" pageNumber="333" pagination="75 - 79" refId="ref9709" refString="Habu, A. (1979) A new species of Mimocolliuris from southern islands of Japan (Coleoptera, Carabidae). The Entomological Review of Japan, 33, 75 - 79." type="journal article" year="1979">Habu, 1979</bibRefCitation>
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</taxonomicName>
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is the
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<typeStatus id="54A1F1C57725FFEB76C543E9A8B0E8CC" box="[687,735,974,1000]" pageId="5" pageNumber="333">type</typeStatus>
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species of the subgenus
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<taxonomicName id="4C1A34E47725FFEB706B43EAAF09E8C3" ID-CoL="8MGR3" authority="Habu, 1979" authorityName="Habu" authorityYear="1979" box="[1025,1382,973,999]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="subGenus" subGenus="Paramimocolliuris">
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<emphasis id="B96E93757725FFEB706B43EAAEBAE8C3" box="[1025,1237,973,999]" italics="true" pageId="5" pageNumber="333">Paramimocolliuris</emphasis>
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<bibRefCitation id="EF8B32967725FFEB70B443EAAF09E8C3" author="Habu, A." box="[1246,1382,973,999]" pageId="5" pageNumber="333" pagination="75 - 79" refId="ref9709" refString="Habu, A. (1979) A new species of Mimocolliuris from southern islands of Japan (Coleoptera, Carabidae). The Entomological Review of Japan, 33, 75 - 79." type="journal article" year="1979">Habu, 1979</bibRefCitation>
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</taxonomicName>
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, and is characterized by the occurrence of two supernumerary setae on the head; its known distribution includes
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<collectingCountry id="F30D0FF77725FFEB713F43D5AFF7EF28" box="[1365,1432,1010,1036]" name="Japan" pageId="5" pageNumber="333">Japan</collectingCountry>
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,
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<collectingCountry id="F30D0FF77725FFEB74FD4432AAB5EF0B" box="[151,218,1045,1071]" name="China" pageId="5" pageNumber="333">China</collectingCountry>
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and
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<collectingCountry id="F30D0FF77725FFEB757B4432AB08EF0B" box="[273,359,1045,1071]" name="Taiwan" pageId="5" pageNumber="333">Taiwan</collectingCountry>
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. The only other species belonging to this subgenus is
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<taxonomicName id="4C1A34E47725FFEB77A94431AF30EF0B" authority="Zhao & Tian, 2011" authorityName="Zhao & Tian" authorityYear="2011" box="[963,1375,1045,1072]" class="Insecta" family="Carabidae" genus="Colliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="species" species="sinuatiphallus">
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<emphasis id="B96E93757725FFEB77A94431AEECEF0B" box="[963,1155,1045,1072]" italics="true" pageId="5" pageNumber="333">C. sinuatiphallus</emphasis>
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Zhao & Tian, 2011
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</taxonomicName>
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from
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<collectingCountry id="F30D0FF77725FFEB74FD441EAAB5EF77" box="[151,218,1081,1107]" name="China" pageId="5" pageNumber="333">China</collectingCountry>
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(
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<collectingRegion id="49DE81857725FFEB7480441EAB1CEF77" box="[234,371,1081,1107]" country="China" name="Guangdong" pageId="5" pageNumber="333">Guangdong</collectingRegion>
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). This single character seems to us to be insufficient for a subgeneric separation. Therefore, in our opinion both
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<taxonomicName id="4C1A34E47725FFEB750B447AA85AEF53" ID-CoL="8MGR3" authorityName="Habu" authorityYear="1979" box="[353,565,1117,1143]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="subGenus" subGenus="Paramimocolliuris">
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<emphasis id="B96E93757725FFEB750B447AA85AEF53" box="[353,565,1117,1143]" italics="true" pageId="5" pageNumber="333">Paramimocolliuris</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1A34E47725FFEB761B4479A930EF53" authority="Liebke, 1933" authorityName="Liebke" authorityYear="1933" box="[625,863,1117,1143]" class="Insecta" family="Carabidae" genus="Essora" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="genus">
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<emphasis id="B96E93757725FFEB761B4479A8D0EF53" box="[625,703,1118,1143]" italics="true" pageId="5" pageNumber="333">Essora</emphasis>
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<bibRefCitation id="EF8B32967725FFEB76A2447AA930EF53" author="Liebke, M." box="[712,863,1117,1143]" pageId="5" pageNumber="333" pagination="202 - 210" refId="ref9920" refString="Liebke, M. (1933) Neue Colliurinen (Coleopt., Carab.). Stylops, 2 (9), 202 - 210. https: // doi. org / 10.1111 / j. 1365 - 3113.1993. tb 01002. x" type="journal article" year="1933">Liebke, 1933</bibRefCitation>
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</taxonomicName>
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(another low diversity subgenus characterized by very subtle diagnostic features) should be synonymized with
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<taxonomicName id="4C1A34E47725FFEB772844A6A98EEFBF" ID-CoL="8KVVT" authorityName="Liebke" authorityYear="1933" box="[834,993,1153,1179]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B96E93757725FFEB772844A6A98EEFBF" box="[834,993,1153,1179]" italics="true" pageId="5" pageNumber="333">Mimocolliuris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25D2E0E7725FFEB778D44A5AE4BEFB8" box="[999,1060,1154,1180]" pageId="5" pageNumber="333" sensu="stricto">s. str.</taxonomicNameLabel>
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, because all their other characters fit well with those of the nominotypical subgenus. Additionally,
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<bibRefCitation id="EF8B32967725FFEB77FA4482AEF7EF9B" author="Terada, K. & Wu, W." box="[912,1176,1189,1216]" pageId="5" pageNumber="333" pagination="15 - 41" refId="ref10136" refString="Terada, K. & Wu, W. (2014) Notes on Taiwanese Caraboidea (Coleoptera) V. A Review of the Tribe Odacanthini (Carabidae) in Taiwan, with Description of Ophionea bhamoensis taiwanensis subsp. nov. Collection and Research, 27, 15 - 41." type="journal article" year="2014">Terada and Wu (2014)</bibRefCitation>
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give some interesting observations on the extreme affinity of some species of
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<taxonomicName id="4C1A34E47725FFEB776144EEA9C5EFC7" ID-CoL="8KVVT" authorityName="Liebke" authorityYear="1933" box="[779,938,1225,1251]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="genus">
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<emphasis id="B96E93757725FFEB776144EEA9C5EFC7" box="[779,938,1225,1251]" italics="true" pageId="5" pageNumber="333">Mimocolliuris</emphasis>
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</taxonomicName>
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to the known species of
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<emphasis id="B96E93757725FFEB70A844EEAFF2EFC7" box="[1218,1437,1225,1251]" italics="true" pageId="5" pageNumber="333">
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<taxonomicName id="4C1A34E47725FFEB70A844EEAFF6EFC7" ID-CoL="8MGR3" authorityName="Habu" authorityYear="1979" box="[1218,1433,1225,1251]" class="Insecta" family="Carabidae" genus="Mimocolliuris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="333" phylum="Arthropoda" rank="subGenus" subGenus="Paramimocolliuris">Paramimocolliuris</taxonomicName>
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.
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</emphasis>
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To finalize their taxonomic placement additional material is required.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |