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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1154.98273" ID-Pensoft-Pub="1313-2970-1154-17" ID-Pensoft-UUID="4086AC35C39554CDB72F84E3FB82AE71" ID-ZooBank="B1C06376F17541F4B772DB50278C5C76" ModsDocID="1313-2970-1154-17" checkinTime="1679118257375" checkinUser="pensoft" docAuthor="Zhang, Yan, Chen, Zhaoyang, Li, Daiqin &amp; Xu, Xin" docDate="2023" docId="0190F05535D751A7ACAAB0178136A982" docLanguage="en" docName="ZooKeys 1154: 17-31" docOrigin="ZooKeys 1154" docPubDate="2023-03-17" docSource="http://dx.doi.org/10.3897/zookeys.1154.98273" docTitle="Songthela anhua Zhang &amp; Xu 2023, sp. nov." docType="treatment" docUuid="508E6FAF-B31C-4680-AFEA-7935331007AC" docUuidSource="ZooBank" docVersion="1" id="4086AC35C39554CDB72F84E3FB82AE71" lastPageNumber="17" masterDocId="4086AC35C39554CDB72F84E3FB82AE71" masterDocTitle="Three new species of the primitively segmented spider genus Songthela (Mesothelae, Liphistiidae, Heptathelinae) from Hunan Province, China" masterLastPageNumber="31" masterPageNumber="17" pageNumber="17" updateTime="1679118257375" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of the primitively segmented spider genus Songthela (Mesothelae, Liphistiidae, Heptathelinae) from Hunan Province, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Yan</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha 410081, Hunan Province, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Zhaoyang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3657-974X</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha 410081, Hunan Province, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Daiqin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8269-7734</mods:nameIdentifier>
<mods:affiliation>Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, 117543, Singapore</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Xin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5632-6622</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha 410081, Hunan Province, China</mods:affiliation>
<mods:nameIdentifier type="email">xuxin_09@163.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1154</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>17</mods:start>
<mods:end>31</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1154.98273</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1154.98273</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1154-17</mods:identifier>
<mods:identifier type="ZooBank">B1C06376F17541F4B772DB50278C5C76</mods:identifier>
<mods:identifier type="Pensoft-UUID">4086AC35C39554CDB72F84E3FB82AE71</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:508E6FAF-B31C-4680-AFEA-7935331007AC" httpUri="http://treatment.plazi.org/id/0190F05535D751A7ACAAB0178136A982" lastPageNumber="17" pageId="0" pageNumber="17">
<subSubSection pageId="0" pageNumber="17" type="nomenclature">
<paragraph pageId="0" pageNumber="17">
<taxonomicName LSID="https://zoobank.org/508E6FAF-B31C-4680-AFEA-7935331007AC" authority="Zhang &amp; Xu" authorityName="Zhang &amp; Xu" authorityYear="2023" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela anhua" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="anhua" status="sp. nov.">Songthela anhua Zhang &amp; Xu</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="17">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="description">
<paragraph pageId="0" pageNumber="17">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="type material">
<paragraph pageId="0" pageNumber="17">Type material.</paragraph>
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" italics="true" pageId="0" pageNumber="17">Holotype</emphasis>
: China · 1 ♂; Hunan Province, Yiyang City, Anhua County, Moon Hill Park;
<geoCoordinate degrees="28.39" direction="north" orientation="latitude" precision="555" value="28.39">28.39°N</geoCoordinate>
;
<geoCoordinate degrees="111.22" direction="east" orientation="longitude" precision="555" value="111.22">111.22°E</geoCoordinate>
; alt. 125 m; 7 September 2021; Z.Y. Chen, X. Xu, Y. Zhan, Y. Zhang leg.; XUX-2021-007 (matured on 25 August 2022).
<emphasis bold="true" italics="true" pageId="0" pageNumber="17">Paratypes</emphasis>
: China · 1 ♂ 3 ♀; same data as for the holotype, alt. 127-144 m; XUX-2021-008, 012, 018 (matured on 25 August 2022), 019.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="diagnosis">
<paragraph pageId="0" pageNumber="17">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="17">
Male of
<taxonomicName lsidName="S. anhua" pageId="0" pageNumber="17" rank="species" species="anhua">
<emphasis italics="true" pageId="0" pageNumber="17">S. anhua</emphasis>
</taxonomicName>
sp. nov. resembles those of
<taxonomicName lsidName="S. tianzhu" pageId="0" pageNumber="17" rank="species" species="tianzhu">
<emphasis italics="true" pageId="0" pageNumber="17">S. tianzhu</emphasis>
</taxonomicName>
Chen, Li, Li &amp; Xu, 2021,
<taxonomicName lsidName="S. yuping" pageId="0" pageNumber="17" rank="species" species="yuping">
<emphasis italics="true" pageId="0" pageNumber="17">S. yuping</emphasis>
</taxonomicName>
Chen, Li, Li &amp; Xu, 2021, and
<taxonomicName lsidName="S. xiangnan" pageId="0" pageNumber="17" rank="species" species="xiangnan">
<emphasis italics="true" pageId="0" pageNumber="17">S. xiangnan</emphasis>
</taxonomicName>
Li, Liu, Li &amp; Xu, 2020 by conductor with blade-shaped apical spine (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3C-E, H, I</figureCitation>
), but can be distinguished from
<taxonomicName lsidName="S. tianzhu" pageId="0" pageNumber="17" rank="species" species="tianzhu">
<emphasis italics="true" pageId="0" pageNumber="17">S. tianzhu</emphasis>
</taxonomicName>
by tegulum with smaller terminal apophysis and distinctly helicoid marginal apophysis (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3F, G</figureCitation>
), by conductor with wider apical spine and several short teeth in prolateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3B, C, E, H, I</figureCitation>
); from
<taxonomicName lsidName="S. xiangnan" pageId="0" pageNumber="17" rank="species" species="xiangnan">
<emphasis italics="true" pageId="0" pageNumber="17">S. xiangnan</emphasis>
</taxonomicName>
by wider apical spine of conductor lacking of bifid apex distally (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3C, E, H, I</figureCitation>
), and by contrategulum with irregular dense dentate margin (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3C, D</figureCitation>
); from
<taxonomicName lsidName="S. yuping" pageId="0" pageNumber="17" rank="species" species="yuping">
<emphasis italics="true" pageId="0" pageNumber="17">S. yuping</emphasis>
</taxonomicName>
by tegulum with slightly narrower dorsal extension of terminal apophysis (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3F, G</figureCitation>
); from those of
<taxonomicName lsidName="S. longhui" pageId="0" pageNumber="17" rank="species" species="longhui">
<emphasis italics="true" pageId="0" pageNumber="17">S. longhui</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="S. zhongpo" pageId="0" pageNumber="17" rank="species" species="zhongpo">
<emphasis italics="true" pageId="0" pageNumber="17">S. zhongpo</emphasis>
</taxonomicName>
sp. nov. by conductor with slightly shorter and blade-shaped apical spine (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3B-E, H, I</figureCitation>
), by tegulum with slightly narrower dorsal extension of terminal apophysis (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3F, G</figureCitation>
); from those of other species of
<taxonomicName authorityName="De.-Q.Li, Chen, Liu, Da.-Q.Li &amp; Xu" authorityYear="2022" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela multidentata" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="multidentata">
<emphasis italics="true" pageId="0" pageNumber="17">Songthela multidentata</emphasis>
</taxonomicName>
-group by conductor with blade-shaped apical spine (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3C-E, H, I</figureCitation>
); from those of other
<taxonomicName authorityName="Ono" authorityYear="2000" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="17">Songthela</emphasis>
</taxonomicName>
species by middle part of conductor with several short teeth (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3B-E, H, I</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">Figure 3.</emphasis>
Male and female genital anatomy of
<taxonomicName authorityName="Zhang &amp; Xu" authorityYear="2023" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela anhua" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="anhua">
<emphasis italics="true" pageId="0" pageNumber="17">Songthela anhua</emphasis>
</taxonomicName>
Zhang &amp; Xu, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="17">A-C</emphasis>
left palp
<emphasis bold="true" pageId="0" pageNumber="17">A</emphasis>
prolateral view
<emphasis bold="true" pageId="0" pageNumber="17">B</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="17">C</emphasis>
retrolateral view
<emphasis bold="true" pageId="0" pageNumber="17">D-G</emphasis>
right palp
<emphasis bold="true" pageId="0" pageNumber="17">D</emphasis>
prolateral view
<emphasis bold="true" pageId="0" pageNumber="17">E</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="17">F</emphasis>
retrolateral view
<emphasis bold="true" pageId="0" pageNumber="17">G</emphasis>
distal view
<emphasis bold="true" pageId="0" pageNumber="17">H</emphasis>
conductor of left palp
<emphasis bold="true" pageId="0" pageNumber="17">I</emphasis>
conductor of right palp
<emphasis bold="true" pageId="0" pageNumber="17">J, K</emphasis>
vulva dorsal view
<emphasis bold="true" pageId="0" pageNumber="17">L, M</emphasis>
vulva ventral view
<emphasis bold="true" pageId="0" pageNumber="17">A-C, H</emphasis>
XUX-2021-007 (holotype)
<emphasis bold="true" pageId="0" pageNumber="17">D-G, I</emphasis>
XUX-2021-018
<emphasis bold="true" pageId="0" pageNumber="17">J, L</emphasis>
XUX-2021-008
<emphasis bold="true" pageId="0" pageNumber="17">K, M</emphasis>
XUX-2021-012. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="17">A-G, J-M</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="17">H, I</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="17">
Females of
<taxonomicName lsidName="S. anhua" pageId="0" pageNumber="17" rank="species" species="anhua">
<emphasis italics="true" pageId="0" pageNumber="17">S. anhua</emphasis>
</taxonomicName>
sp. nov. can be distinguished from those of
<taxonomicName lsidName="S. longhui" pageId="0" pageNumber="17" rank="species" species="longhui">
<emphasis italics="true" pageId="0" pageNumber="17">S. longhui</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="S. pluma" pageId="0" pageNumber="17" rank="species" species="pluma">
<emphasis italics="true" pageId="0" pageNumber="17">S. pluma</emphasis>
</taxonomicName>
Yu, Li &amp; Zhang, 2018 by median receptacular clusters slightly larger than lateral ones, and middle genital stalks separated from each other basally (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3J, K</figureCitation>
); from
<taxonomicName lsidName="S. zhongpo" pageId="0" pageNumber="17" rank="species" species="zhongpo">
<emphasis italics="true" pageId="0" pageNumber="17">S. zhongpo</emphasis>
</taxonomicName>
sp. nov. by trapeziform anterior margin of bursa copulatrix, median receptacular clusters with slightly longer genital stalks (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3J-K</figureCitation>
); from those of other species of
<taxonomicName authorityName="De.-Q.Li, Chen, Liu, Da.-Q.Li &amp; Xu" authorityYear="2022" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela multidentata" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="multidentata">
<emphasis italics="true" pageId="0" pageNumber="17">Songthela multidentata</emphasis>
</taxonomicName>
-group by median receptacular clusters separated from each other basally and posterior margin of genital area wider and flat (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3J-M</figureCitation>
); from those of other
<taxonomicName authorityName="Ono" authorityYear="2000" class="Arachnida" family="Heptathelidae" genus="Songthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Songthela" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="17">Songthela</emphasis>
</taxonomicName>
species by four receptacular clusters situated at the dorsal side of bursa copulatrix (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3J-M</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="description">
<paragraph pageId="0" pageNumber="17">Description.</paragraph>
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">Male</emphasis>
(holotype; Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Microhabitat (A-D) and general somatic morphology (E-J) of three new Songthela species E-G female H-J male E, H Songthela zhongpo Zhang &amp; Xu, sp. nov. F, I Songthela longhui Zhang &amp; Xu, sp. nov. G, J Songthela anhua Zhang &amp; Xu, sp. nov. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1154.98273.figure1" httpUri="https://binary.pensoft.net/fig/826618" pageId="0" pageNumber="17">1J</figureCitation>
). Carapace black brown; opisthosoma yellow brown, with 12 black brown tergites attached a pair of thick bristles, the second to fifth larger than others and the fourth largest; sternum narrow, much longer than wide; a few pointed hairs running over ocular area; chelicerae robust with promargin of cheliceral groove with 14 denticles of variable size; legs with sturdy hairs and spines; 8 spinnerets. Measurements: BL 13.28, CL 6.42, CW 5.70, OL 6.19, OW 4.85; ALE&gt; PLE&gt; PME&gt; AME; leg I 20.58 (5.80 + 2.53 + 4.25 + 5.27 + 2.73), leg II 19.69 (5.54 + 2.52 + 4.18 + 4.88 + 2.57), leg III 22.95 (5.56 + 2.69 + 4.28 + 6.95 + 3.47), leg IV 28.29 (7.32 + 3.03 + 5.69 + 7.97 + 4.28).
</paragraph>
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">
<emphasis italics="true" pageId="0" pageNumber="17">Palp</emphasis>
.
</emphasis>
Prolateral portion of paracymbium unpigmented and unsclerotised, with several setae and spines on the tip (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3A-C</figureCitation>
). Contrategulum with an obviously triangular apophysis proximally and two irregular dentate edges distally (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3C, D</figureCitation>
). Tegulum with a helicoid marginal apophysis, a dentate dorsal extension of the terminal apophysis, and a thumb-like terminal apophysis retrolaterally (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3F, G</figureCitation>
). Conductor lamellar, fused with embolus ventroproximally, with a blade-shaped apical spine pointed to the one-third of opening of embolus proximally, and the middle portion inserted with several teeth (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3B-E, H, I</figureCitation>
). Embolus largely sclerotized with a wide opening, several ribbed ridges, and with a twisted top in ventral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3B-E, G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">Female</emphasis>
(XUX-2021-008; Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Microhabitat (A-D) and general somatic morphology (E-J) of three new Songthela species E-G female H-J male E, H Songthela zhongpo Zhang &amp; Xu, sp. nov. F, I Songthela longhui Zhang &amp; Xu, sp. nov. G, J Songthela anhua Zhang &amp; Xu, sp. nov. Scale bar: 5 mm." figureDoi="10.3897/zookeys.1154.98273.figure1" httpUri="https://binary.pensoft.net/fig/826618" pageId="0" pageNumber="17">1G</figureCitation>
). Carapace yellow brown and opisthosoma dark brown in alcohol, with 12 dark brown tergites attached a pair of thick bristles, the second to fifth larger than others and the fourth largest; sternum narrow, much longer than wide; a few pointed hairs running over ocular area; chelicerae robust with promargin of cheliceral groove with 12 denticles of variable size; legs with sturdy hairs and spines; 7 spinnerets. Measurements: BL 10.18, CL 5.16, CW 4.52, OL 4.43, OW 3.57; ALE&gt; PLE&gt; PME&gt; AME; palp 9.82 (3.37 + 1.81 + 2.41 + 2.23), leg I 10.97 (3.66 + 1.83 + 2.16 + 1.96 + 1.36), leg II 10.29 (3.21 + 1.79 + 1.86 + 2.06 + 1.37), leg III 9.71 (2.91 + 1.74 + 1.22 + 2.27 + 1.57), leg IV 14.37 (4.22 + 1.60 + 2.49 + 3.72 + 2.34).
</paragraph>
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">
<emphasis italics="true" pageId="0" pageNumber="17">Female genitalia</emphasis>
.
</emphasis>
Two pairs of receptacular clusters situated on the dorsal side of the bursa copulatrix; the middle pair of receptacular clusters with long genital stalks and larger than the lateral ones, the middle stalks separated from each other; the posterior margin of the bursa copulatrix sclerotized; the posterior margin of the genital area wide (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3J-M</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="variation">
<paragraph pageId="0" pageNumber="17">Variation.</paragraph>
<paragraph pageId="0" pageNumber="17">
Males and females vary in body size, cheliceral teeth, and spinnerets. Range of measurements in males (
<emphasis italics="true" pageId="0" pageNumber="17">N</emphasis>
= 2): BL 11.94-13.28, CL 5.59-6.42, CW 5.07-5.70, OL 6.08-6.19, OW 4.80-4.85; the number of cheliceral teeth varies from 9-14 (
<emphasis italics="true" pageId="0" pageNumber="17">N</emphasis>
= 2); there are 7 or 8 spinnerets. Females (
<emphasis italics="true" pageId="0" pageNumber="17">N</emphasis>
= 3): BL 10.18-11.51, CL 5.16-5.57, CW 4.52-4.84, OL 4.43-5.66, OW 3.57-4.04; the number of cheliceral teeth varies from 12-13 (
<emphasis italics="true" pageId="0" pageNumber="17">N</emphasis>
= 3); there are 7 or 8 spinnerets. In addition, male palp and female genitalia also show intraspecific variations: in males, the left palp is slightly different from the right palp, e.g. the tegulum of left male palp with three teeth basally in ventral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3B</figureCitation>
), but they are missing in right male palp (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3E, F</figureCitation>
); the number of teeth located in middle portion of conductor also varies between left and right palp (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3C, E, H, I</figureCitation>
); in females, the relative position of the middle receptacular clusters situated on the dorsal side of the bursa copulatrix is slightly different (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genital anatomy of Songthela anhua Zhang &amp; Xu, sp. nov. A-C left palp A prolateral view B ventral view C retrolateral view D-G right palp D prolateral view E ventral view F retrolateral view G distal view H conductor of left palp I conductor of right palp J, K vulva dorsal view L, M vulva ventral view A-C, H XUX- 2021 - 007 (holotype) D-G, I XUX- 2021 - 018 J, L XUX- 2021 - 008 K, M XUX- 2021 - 012. Scale bars: 0.5 mm (A-G, J-M); 0.1 mm (H, I)." figureDoi="10.3897/zookeys.1154.98273.figure3" httpUri="https://binary.pensoft.net/fig/826620" pageId="0" pageNumber="17">3L, M</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="etymology">
<paragraph pageId="0" pageNumber="17">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="17">The species epithet, a noun in apposition, refers to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="distribution">
<paragraph pageId="0" pageNumber="17">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="17">Hunan (Anhua), China</paragraph>
</subSubSection>
</treatment>
</document>
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