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<document id="5506DB97D37483DDC85E1EC40AF5BC3F" ID-DOI="10.11646/zootaxa.5375.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10203164" ID-ZooBank="7DBBC861-FC05-4F54-A851-FEC9A5821802" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700816835735" checkinUser="plazi" docAuthor="Bekchanov, Norbek Kh., Nabozhenko, Maxim V. &amp; Bekchanova, Mokhira Kh." docDate="2023" docId="0142E9105351FFC013ACB6F5FF6DFEF8" docLanguage="en" docName="zootaxa.5375.4.6.pdf" docOrigin="Zootaxa 5375 (4)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5375.4.6/52345" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colposcelis (Turcmenicola) ustyurtensis Bekchanov &amp; Nabozhenko &amp; Bekchanova 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="557" masterDocId="FD7B91685355FFC7133BB318FFAAFFE5" masterDocTitle="Review of the subgenus Turcmenicola Bogatchev, 1952, the genus Colposcelis Dejean, 1834 (Coleoptera: Tenebrionidae: Tentyriini)" masterLastPageNumber="564" masterPageNumber="550" pageNumber="554" updateTime="1700817255677" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2F5185F9D03F5E3B523C3998A98F43B2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4C5707052A451B0FE6D2D38E7E052DB0">Review of the subgenus Turcmenicola Bogatchev, 1952, the genus Colposcelis Dejean, 1834 (Coleoptera: Tenebrionidae: Tentyriini)</mods:title>
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<mods:name id="59927CE846541430412669C356BC9DF8" type="personal">
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<mods:namePart id="A052DE3B8420F3CB4EF78D42A5083CC0">Bekchanov, Norbek Kh.</mods:namePart>
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<mods:namePart id="4554038EC03DC11B8E00963C551FB4C3">Nabozhenko, Maxim V.</mods:namePart>
<mods:affiliation id="54B76E2CB32F05372866C4261C6F69E9">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000, Russia. &amp; Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000, Russia</mods:affiliation>
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<mods:namePart id="D89B2DC4E914D9AE4891031D9AAE231C">Bekchanova, Mokhira Kh.</mods:namePart>
<mods:affiliation id="4B8B18731CEF21E327A9BCAF8286B146">Urgench State University, Kh. Alimdjan str., 14, Urgench 220100, Uzbekistan.</mods:affiliation>
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<mods:title id="86CFA439A9448F45A7C37E7EA8FDF5F9">Zootaxa</mods:title>
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<mods:part id="1381EA72FA19F6263C1B286C4CCFB98B">
<mods:date id="EC4B7D935FC29F0A5E5265FB68DBB52F">2023</mods:date>
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<treatment id="0142E9105351FFC013ACB6F5FF6DFEF8" LSID="urn:lsid:plazi:treatment:0142E9105351FFC013ACB6F5FF6DFEF8" httpUri="http://treatment.plazi.org/id/0142E9105351FFC013ACB6F5FF6DFEF8" lastPageId="7" lastPageNumber="557" pageId="4" pageNumber="554">
<subSubSection id="C1F10B8D5351FFC313ACB6F5FB88F9ED" box="[151,1058,1517,1544]" pageId="4" pageNumber="554" type="nomenclature">
<paragraph id="895458065351FFC313ACB6F5FB88F9ED" blockId="4.[151,1058,1517,1580]" box="[151,1058,1517,1544]" pageId="4" pageNumber="554">
<heading id="D21CEF6A5351FFC313ACB6F5FB88F9ED" bold="true" box="[151,1058,1517,1544]" fontSize="11" level="1" pageId="4" pageNumber="554" reason="1">
<emphasis id="BB9F84145351FFC313ACB6F5FB88F9ED" bold="true" box="[151,1058,1517,1544]" pageId="4" pageNumber="554">
<taxonomicName id="4EEB23855351FFC313ACB6F5FC79F9ED" authority="Bekchanov &amp; Nabozhenko" authorityName="Bekchanov &amp; Nabozhenko &amp; Bekchanova" authorityYear="2023" box="[151,979,1517,1544]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="554" phylum="Arthropoda" rank="species" species="ustyurtensis" status="sp. nov." subGenus="Turcmenicola">
<emphasis id="BB9F84145351FFC313ACB6F5FE8AF9ED" bold="true" box="[151,288,1517,1544]" italics="true" pageId="4" pageNumber="554">Colposcelis</emphasis>
(
<emphasis id="BB9F84145351FFC3120AB6F5FE73F9ED" bold="true" box="[305,473,1517,1544]" italics="true" pageId="4" pageNumber="554">Turcmenicola</emphasis>
)
<emphasis id="BB9F84145351FFC312D0B6F6FDD6F9ED" bold="true" box="[491,636,1518,1544]" italics="true" pageId="4" pageNumber="554">ustyurtensis</emphasis>
Bekchanov &amp; Nabozhenko
</taxonomicName>
,
<taxonomicNameLabel id="A0AC396F5351FFC310DBB6F6FB88F9ED" box="[992,1058,1518,1544]" pageId="4" pageNumber="554" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C1F10B8D5351FFC313ACB50AFF46F968" pageId="4" pageNumber="554" type="description">
<paragraph id="895458065351FFC313ACB50AFEA1F9C9" blockId="4.[151,1058,1517,1580]" box="[151,267,1554,1580]" pageId="4" pageNumber="554">
(
<figureCitation id="11D044835351FFC313A4B50AFF4DF9C9" box="[159,231,1554,1580]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="4" pageNumber="554">Figs 3</figureCitation>
,
<figureCitation id="11D044835351FFC313CFB50AFEA9F9C9" box="[244,259,1554,1580]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="4" pageNumber="554">4</figureCitation>
)
</paragraph>
<paragraph id="895458065351FFC313ACB54EFF46F968" blockId="4.[151,1436,1622,1677]" pageId="4" pageNumber="554">
<bibRefCitation id="ED7A25F75351FFC313ACB54EFEE1F98A" author="Skopin, N. G." box="[151,331,1622,1647]" pageId="4" pageNumber="554" pagination="277 - 296" refId="ref8121" refString="Skopin, N. G. (1964) Material to the fauna of tenebrionids (Coleoptera, Tenebrionidae) of Mangyshlak and North-Western Ustyurt. Trudy Kazakhskogo nauchno-issledovatel'skogo instituta zashchity rasteniy, 8, 277 - 296. [in Russian]" type="journal article" year="1964">Skopin 1964: 290</bibRefCitation>
(? as
<taxonomicName id="4EEB23855351FFC312BCB54EFD53F98B" baseAuthorityName="Bogatchev" baseAuthorityYear="1952" box="[391,761,1622,1647]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="554" phylum="Arthropoda" rank="species" species="jachontovi" subGenus="Scelocolpis">
<emphasis id="BB9F84145351FFC312BCB54EFE57F98B" box="[391,509,1622,1646]" italics="true" pageId="4" pageNumber="554">Colposcelis</emphasis>
(
<emphasis id="BB9F84145351FFC31137B54EFDD5F98B" box="[524,639,1622,1646]" italics="true" pageId="4" pageNumber="554">Scelocolpis</emphasis>
)
<emphasis id="BB9F84145351FFC311B5B54EFD53F98B" box="[654,761,1622,1646]" italics="true" pageId="4" pageNumber="554">jachontovi</emphasis>
</taxonomicName>
);
<bibRefCitation id="ED7A25F75351FFC31034B54EFB83F98B" author="Bekchanov, N. Kh. &amp; Nabozhenko, M. V. &amp; Bekchanov, Kh. U." box="[783,1065,1622,1647]" pageId="4" pageNumber="554" pagination="155 - 191" refId="ref7057" refString="Bekchanov, N. Kh., Nabozhenko, M. V. &amp; Bekchanov, Kh. U. (2023) Remarkable additions to the fauna of darkling beetles (Coleoptera: Tenebrionidae) of western Uzbekistan. Caucasian Entomological Bulletin, 19 (1), 155 - 191. https: // doi. org / 10.5281 / zenodo. 8199930" type="journal article" year="2023">
Bekchanov
<emphasis id="BB9F84145351FFC310B3B54FFC1EF98B" box="[904,948,1622,1646]" italics="true" pageId="4" pageNumber="554">et al</emphasis>
. 2023: 175
</bibRefCitation>
, fig. 23 (
<taxonomicName id="4EEB23855351FFC317BFB54EFA36F98A" box="[1156,1436,1622,1647]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="554" phylum="Arthropoda" rank="subGenus" subGenus="Turcmenicola">
<emphasis id="BB9F84145351FFC317BFB54EFB50F98B" box="[1156,1274,1622,1646]" italics="true" pageId="4" pageNumber="554">Colposcelis</emphasis>
(
<emphasis id="BB9F84145351FFC31632B54EFA39F98B" box="[1289,1427,1622,1646]" italics="true" pageId="4" pageNumber="554">Turcmenicola</emphasis>
)
</taxonomicName>
sp.)
</paragraph>
</subSubSection>
<subSubSection id="C1F10B8D5351FFC313ACB5A3FC24F8A4" pageId="4" pageNumber="554" type="materials_examined">
<paragraph id="895458065351FFC313ACB5A3FC24F8A4" blockId="4.[151,1437,1723,2038]" pageId="4" pageNumber="554">
<emphasis id="BB9F84145351FFC313ACB5A3FEE9F930" bold="true" box="[151,323,1723,1749]" pageId="4" pageNumber="554">Type material.</emphasis>
<materialsCitation id="3983525B5351FFC31273B5A3FB11F8F8" collectingDate="2023-04-23" collectionCode="ZIN, IZUZ, PCMN, PCNB" collectorName="M. V and S. V. Nabozhenko &amp; N. Kh. and Kh. U. Bekchanov &amp; U. E. Duschanov &amp; D. A. Yavkachev" country="Uzbekistan" latitude="43.085556" location="Karakalpakstan" longLatPrecision="19" longitude="58.540276" municipality="Kungrad District" pageId="4" pageNumber="554" specimenCount="12" specimenCount-female="11" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="5650E6A45351FFC31273B5A3FE19F930" box="[328,435,1723,1749]" pageId="4" pageNumber="554" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9FED938F5351FFC31286B5A4FE78F933" box="[445,466,1724,1750]" count="1" pageId="4" pageNumber="554" type="male">♂</specimenCount>
(
<collectionCode id="EFFAC0C35351FFC312E4B5A3FDBBF930" box="[479,529,1723,1749]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="554" type="Museum">ZIN</collectionCode>
) and
<typeStatus id="5650E6A45351FFC31170B5A4FD13F933" box="[587,697,1724,1750]" pageId="4" pageNumber="554" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9FED938F5351FFC311F9B5A3FCA7F933" box="[706,781,1723,1750]" count="11" pageId="4" pageNumber="554" type="female">11 ♀♀</specimenCount>
(
<collectionCode id="EFFAC0C35351FFC31021B5A3FCE7F930" box="[794,845,1723,1749]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="554" type="Museum">ZIN</collectionCode>
,
<collectionCode id="EFFAC0C35351FFC3106DB5A4FC33F933" box="[854,921,1724,1750]" pageId="4" pageNumber="554">IZUZ</collectionCode>
,
<collectionCode id="EFFAC0C35351FFC31099B5A3FC5EF930" box="[930,1012,1723,1749]" pageId="4" pageNumber="554">PCMN</collectionCode>
,
<collectionCode id="EFFAC0C35351FFC310C6B5A3FBE6F930" box="[1021,1100,1723,1749]" pageId="4" pageNumber="554">PCNB</collectionCode>
):
<collectingCountry id="F1FC18965351FFC31766B5A3FB74F930" box="[1117,1246,1723,1749]" name="Uzbekistan" pageId="4" pageNumber="554">Uzbekistan</collectingCountry>
,
<location id="8C340EDD5351FFC317DCB5A3FA32F930" LSID="urn:lsid:plazi:treatment:0142E9105351FFC013ACB6F5FF6DFEF8:8C340EDD5351FFC317DCB5A3FA32F930" box="[1255,1432,1723,1749]" country="Uzbekistan" latitude="43.085556" longLatPrecision="19" longitude="58.540276" municipality="Kungrad District" name="Karakalpakstan" pageId="4" pageNumber="554">Karakalpakstan</location>
,
<collectingMunicipality id="6930C27C5351FFC313ACB5C7FEFDF91C" box="[151,343,1759,1785]" pageId="4" pageNumber="554">Kungrad District</collectingMunicipality>
,
<locationDeviation id="98E641245351FFC3125AB5C7FCC2F91C" box="[353,872,1759,1786]" firstLocation="Elabad" pageId="4" pageNumber="554" secondLocation="Kungrad, Ustyurt Plateau">between Elabad and Kungrad, Ustyurt Plateau</locationDeviation>
, clay desert,
<geoCoordinate id="ECDF3EC15351FFC310C2B5F9FB2CF91F" box="[1017,1158,1760,1788]" degrees="43" direction="north" minutes="05" orientation="latitude" pageId="4" pageNumber="554" precision="15" seconds="08" value="43.085556">43°05ʹ08ʺ N</geoCoordinate>
,
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,
<date id="FD557EC65351FFC3161DB5C7FA36F91C" box="[1318,1436,1759,1785]" pageId="4" pageNumber="554" value="2023-04-23">
<collectingDate id="ED11872E5351FFC3161DB5C7FA36F91C" box="[1318,1436,1759,1785]" pageId="4" pageNumber="554" value="2023-04-23">23.iv.2023</collectingDate>
</date>
(
<collectorName id="241E3DD05351FFC313A4B41BFE7BF8F8" box="[159,465,1795,1821]" pageId="4" pageNumber="554">M.V and S.V. Nabozhenko</collectorName>
,
<collectorName id="241E3DD05351FFC312E5B41BFC8FF8F8" box="[478,805,1795,1821]" pageId="4" pageNumber="554">N.Kh. and Kh.U. Bekchanov</collectorName>
,
<collectorName id="241E3DD05351FFC3100BB41CFC40F8F8" box="[816,1002,1795,1822]" pageId="4" pageNumber="554">U.E. Duschanov</collectorName>
,
<collectorName id="241E3DD05351FFC310CDB41CFB1EF8F8" box="[1014,1204,1795,1822]" pageId="4" pageNumber="554">D.A. Yavkachev</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3983525B5351FFC317FDB41CFC21F8A4" collectingDate="2011-04-16" collectionCode="ZIN" collectorName="A. Abramov" country="Kazakhstan" location="Mangystau Region" municipality="Beyneu" pageId="4" pageNumber="554" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="5650E6A45351FFC317FDB41CFA81F8FB" box="[1222,1323,1796,1822]" pageId="4" pageNumber="554" type="paratype">Paratype</typeStatus>
,
<specimenCount id="9FED938F5351FFC3160CB41CFAE6F8FB" box="[1335,1356,1796,1822]" count="1" pageId="4" pageNumber="554" type="female">♀</specimenCount>
(
<collectionCode id="EFFAC0C35351FFC31660B41BFA3AF8F8" box="[1371,1424,1795,1821]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="554" type="Museum">ZIN</collectionCode>
):
<collectingCountry id="F1FC18965351FFC313ACB43FFEB7F8A4" box="[151,285,1831,1857]" name="Kazakhstan" pageId="4" pageNumber="554">Kazakhstan</collectingCountry>
,
<location id="8C340EDD5351FFC31213B430FE55F8A4" LSID="urn:lsid:plazi:treatment:0142E9105351FFC013ACB6F5FF6DFEF8:8C340EDD5351FFC31213B430FE55F8A4" box="[296,511,1831,1858]" country="Kazakhstan" municipality="Beyneu" name="Mangystau Region" pageId="4" pageNumber="554">Mangystau Region</location>
,
<collectingMunicipality id="6930C27C5351FFC31131B430FDC8F8A7" box="[522,610,1832,1858]" pageId="4" pageNumber="554">Beyneu</collectingMunicipality>
,
<date id="FD557EC65351FFC31155B43FFD48F8A4" box="[622,738,1831,1857]" pageId="4" pageNumber="554" value="2011-04-16">
<collectingDate id="ED11872E5351FFC31155B43FFD48F8A4" box="[622,738,1831,1857]" pageId="4" pageNumber="554" value="2011-04-16">16.iv.2011</collectingDate>
</date>
(
<collectorName id="241E3DD05351FFC311C9B43FFC29F8A4" box="[754,899,1831,1857]" pageId="4" pageNumber="554">A. Abramov</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C1F10B8D5351FFC113FCB453FDA2F8B3" lastPageId="6" lastPageNumber="556" pageId="4" pageNumber="554" type="description">
<paragraph id="895458065351FFC213FCB453FB2BFF1C" blockId="4.[151,1437,1723,2038]" lastBlockId="5.[151,1437,151,465]" lastPageId="5" lastPageNumber="555" pageId="4" pageNumber="554">
<emphasis id="BB9F84145351FFC313FCB453FEF3F880" bold="true" box="[199,345,1867,1893]" pageId="4" pageNumber="554">Description.</emphasis>
Male. Body length
<quantity id="4E13F5E35351FFC31100B453FDD5F883" box="[571,639,1867,1894]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="554" unit="mm" value="9.0">9 mm</quantity>
, width
<quantity id="4E13F5E35351FFC311E9B453FC86F883" box="[722,812,1867,1894]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="554" unit="mm" value="3.7">3.7 mm</quantity>
. Body robust, with long and slender legs and antennae, elytra much wider than pronotum, black, weakly shiny, legs black or dark-brown (
<figureCitation id="11D044835351FFC31708B477FB1EF86C" box="[1075,1204,1903,1929]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="4" pageNumber="554">Figs 3A–C</figureCitation>
). Head widest at eye level, where 1.07 times as wide as at genal level. Eyes moderately convex. Head at eye level 1.2 times as wide as interocular space. Lateral margin of eyes together with temples evenly rounded, not angulate (
<figureCitation id="11D044835351FFC3179EB4AFFB51F834" box="[1189,1275,1975,2001]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="4" pageNumber="554">Fig. 3E</figureCitation>
). Puncturation very fine and sparse, punctures 6–7 times as short as interpuncture distance. Keels inside from eyes moderately short, not reaching middle of genae. Temples ventrally with coarse punctures and longitudinal wrinkles, head surface between temples with coarse and sparse punctures (
<figureCitation id="11D044835350FFC21009B3A3FC20FF30" box="[818,906,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="5" pageNumber="555">Fig. 3C</figureCitation>
). Antennae long, with four distal antennomeres extending beyond base of pronotum. All antennomeres with fine and sparse puncturation.
</paragraph>
<paragraph id="895458065350FFC213FCB21BFB78FE34" blockId="5.[151,1437,151,465]" pageId="5" pageNumber="555">
Prothorax (
<figureCitation id="11D044835350FFC2127CB21BFE44FEF8" box="[327,494,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="5" pageNumber="555">Figs 3A, B, D</figureCitation>
). Pronotum oval, transverse (1.13 times as wide as long), widest slightly ahead of middle, 1.13 times as wide as head. Anterior margin straight in middle, lateral margins evenly strongly rounded, base slightly rounded. Lateral margins and base completely bordered, while anterior margin bordered only on sides. Disc of pronotum strongly convex, very finely and sparsely punctured. Prosternum and prothoracic hypomera with almost invisible fine and sparse puncturation; each puncture with very short white seta (
<figureCitation id="11D044835350FFC2175FB28BFB14FE48" box="[1124,1214,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="5" pageNumber="555">Fig. 3D</figureCitation>
). Prosternal process widely depressed between procoxae and with weakly protruded narrowest middle part (
<figureCitation id="11D044835350FFC21750B2AFFB6FFE34" box="[1131,1221,439,465]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="5" pageNumber="555">Fig. 3D</figureCitation>
).
</paragraph>
<caption id="DD94088E5350FFC213ACB4BEFBA8F807" ID-DOI="http://doi.org/10.5281/zenodo.10203170" ID-Zenodo-Dep="10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="5" pageNumber="555" startId="5.[151,250,1958,1982]" targetBox="[154,1431,532,1934]" targetPageId="5" targetType="figure">
<paragraph id="895458065350FFC213ACB4BEFBA8F807" blockId="5.[151,1436,1958,2018]" pageId="5" pageNumber="555">
<emphasis id="BB9F84145350FFC213ACB4BEFEB9F85B" bold="true" box="[151,275,1958,1982]" pageId="5" pageNumber="555">FIGURE 3.</emphasis>
<taxonomicName id="4EEB23855350FFC21221B4BEFDBAF85B" authority="Bekchanov &amp; Nabozhenko &amp; Bekchanova, 2023" authorityName="Bekchanov &amp; Nabozhenko &amp; Bekchanova" authorityYear="2023" box="[282,528,1958,1982]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="555" phylum="Arthropoda" rank="species" species="ustyurtensis" status="sp. nov.">
<emphasis id="BB9F84145350FFC21221B4BEFDBAF85B" box="[282,528,1958,1982]" italics="true" pageId="5" pageNumber="555">Colposcelis ustyurtensis</emphasis>
</taxonomicName>
<emphasis id="BB9F84145350FFC2112CB4BEFDFAF85B" bold="true" box="[535,592,1958,1982]" pageId="5" pageNumber="555">
<taxonomicNameLabel id="A0AC396F5350FFC2112CB4BEFDFAF85B" box="[535,592,1958,1982]" pageId="5" pageNumber="555" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally.
</paragraph>
</caption>
<caption id="DD94088E5353FFC113ACB0C6FEB0FBD8" ID-DOI="http://doi.org/10.5281/zenodo.10203174" ID-Zenodo-Dep="10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="6" pageNumber="556" startId="6.[151,250,990,1014]" targetBox="[155,1432,181,963]" targetPageId="6" targetType="figure">
<paragraph id="895458065353FFC113ACB0C6FEB0FBD8" blockId="6.[151,1436,990,1086]" pageId="6" pageNumber="556">
<emphasis id="BB9F84145353FFC113ACB0C6FEB9FC13" bold="true" box="[151,275,990,1014]" pageId="6" pageNumber="556">FIGURE 4.</emphasis>
<taxonomicName id="4EEB23855353FFC11222B0C6FDA4FC13" authority="Bekchanov &amp; Nabozhenko &amp; Bekchanova, 2023" authorityName="Bekchanov &amp; Nabozhenko &amp; Bekchanova" authorityYear="2023" box="[281,526,990,1014]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="ustyurtensis" status="sp. nov.">
<emphasis id="BB9F84145353FFC11222B0C6FDA4FC13" box="[281,526,990,1014]" italics="true" pageId="6" pageNumber="556">Colposcelis ustyurtensis</emphasis>
</taxonomicName>
<emphasis id="BB9F84145353FFC1112FB0C6FDE7FC13" bold="true" box="[532,589,990,1014]" pageId="6" pageNumber="556">
<taxonomicNameLabel id="A0AC396F5353FFC1112FB0C6FDE7FC13" box="[532,589,990,1014]" pageId="6" pageNumber="556" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII.
</paragraph>
</caption>
<paragraph id="895458065353FFC113FCB774FC08FA67" blockId="6.[151,1437,1132,2023]" pageId="6" pageNumber="556">
Pterothorax (
<figureCitation id="11D044835353FFC11263B774FE73FB63" box="[344,473,1132,1158]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="6" pageNumber="556">Figs 3A–D</figureCitation>
). Elytra convex, wide, widest at middle (1.3 times as long as wide), 1.83 times as wide as head, 1.6 times as wide and 3.36 times as long as pronotum. Border of elytral base 1.2 times as wide as width of pronotum, interrupted in middle (not reaching scutellar shield) (
<figureCitation id="11D044835353FFC110ACB7ACFBB0FB2B" box="[919,1050,1204,1230]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="6" pageNumber="556">Figs 3A, B</figureCitation>
). Humeri widely rounded, poorly expressed. Puncturation of elytra very fine and sparse, almost invisible. Epipleura slightly depressed in basal part. Mesoventrite with simple (not rasp-like) concentric deep merged foveae in anterior half (
<figureCitation id="11D044835353FFC117AAB7E4FB47FAF3" box="[1169,1261,1276,1302]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="6" pageNumber="556">Fig. 3D</figureCitation>
). Posterior part of mesoventrite with intercoxal process, mesepimera, metepisterna and metepimera finely and sparsely punctured. Mesepisterna with sparse rasp-like puncturation. Metaventrite with very fine and sparse poorly visible puncturation in general and bracket-like coarse foveae along mesocoxae (
<figureCitation id="11D044835353FFC11000B670FC3FFA67" box="[827,917,1384,1410]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="6" pageNumber="556">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="895458065353FFC113FCB694FB7CF99B" blockId="6.[151,1437,1132,2023]" pageId="6" pageNumber="556">
Abdomen. Intercoxal process of abdominal ventrite 1 not bordered at apex; the border interruption with straight grooves before apex; surface along these grooves with bracket-like coarse foveae (
<figureCitation id="11D044835353FFC1175DB6A8FB6FFA2F" box="[1126,1221,1456,1482]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="6" pageNumber="556">Fig. 3D</figureCitation>
). Other surface of abdominal ventrite 1 and ventrites 2–5 with very sparse and fine punctures. Male terminalia and genitalia (
<figureCitation id="11D044835353FFC11672B6CCFA25FA0B" box="[1353,1423,1492,1518]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="6" pageNumber="556">Fig. 4</figureCitation>
). Male inner sternite VIII weakly sclerotized and apical part and more sclerotized in basal part, with deep emargination and acute apices (
<figureCitation id="11D044835353FFC11259B504FE10F9D3" box="[354,442,1564,1590]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="6" pageNumber="556">Fig. 4G</figureCitation>
). Spiculum gastrale with transverse triangular blades (
<figureCitation id="11D044835353FFC11712B504FB2AF9D3" box="[1065,1152,1564,1590]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="6" pageNumber="556">Fig. 4F</figureCitation>
). Apical piece of tegmen evenly roundly narrowed from base to apex; basal piece 1.38 times as long as apical piece, widest at apical third (
<figureCitation id="11D044835353FFC113A4B57CFF50F99B" box="[159,250,1636,1662]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="6" pageNumber="556">Fig. 4D</figureCitation>
). Median lobe very narrow, widest at basal stem, strongly sclerotized laterally (
<figureCitation id="11D044835353FFC11749B57CFB60F99B" box="[1138,1226,1636,1662]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="6" pageNumber="556">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="895458065353FFC113FCB591FCF3F923" blockId="6.[151,1437,1132,2023]" pageId="6" pageNumber="556">
Legs long and slender. Trochanters without setae (
<figureCitation id="11D044835353FFC11035B590FCC1F947" box="[782,875,1672,1698]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="6" pageNumber="556">Fig. 3D</figureCitation>
). Protibiae slightly S-shaped from extensor side, weakly bent along entire length from dorsal view (
<figureCitation id="11D044835353FFC111F6B5B4FCE1F923" box="[717,843,1708,1734]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="6" pageNumber="556">Figs 4A, B</figureCitation>
).
</paragraph>
<paragraph id="895458065353FFC113FCB5C8FDA2F8B3" blockId="6.[151,1437,1132,2023]" pageId="6" pageNumber="556">
Female (
<figureCitation id="11D044835353FFC11211B5C8FE03F90F" box="[298,425,1744,1770]" captionStart="FIGURE 3" captionStartId="5.[151,250,1958,1982]" captionTargetBox="[154,1431,532,1934]" captionTargetId="figure-172@5.[151,1436,525,1934]" captionTargetPageId="5" captionText="FIGURE 3. Colposcelis ustyurtensis sp. n., habitus, details of structure. A = male, holotype, dorsally; B = female, dorsally; C = female, ventrally; D = thorax and abdominal ventrite 1, ventrally; E = head dorsally." figureDoi="http://doi.org/10.5281/zenodo.10203170" httpUri="https://zenodo.org/record/10203170/files/figure.png" pageId="6" pageNumber="556">Figs 3B, C</figureCitation>
). Body length
<quantity id="4E13F5E35353FFC11169B5C8FD76F90E" box="[594,732,1744,1771]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="9.7" metricValueMin="9.5" pageId="6" pageNumber="556" unit="mm" value="9.6" valueMax="9.7" valueMin="9.5">9.5–9.7 mm</quantity>
, width
<quantity id="4E13F5E35353FFC1100BB5C8FC16F90E" box="[816,956,1744,1771]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="6" pageNumber="556" unit="mm" value="4.35" valueMax="4.4" valueMin="4.3">4.3–4.4 mm</quantity>
. Body more robust, antennae shorter, only with three apical antennomeres extending beyond base of pronotum. Pronotum widest at middle, 1.18–1.2 times as wide as long. Elytra 2–2.1 times as wide as head and 1.7–1.75 times as wide as pronotum. Legs shorter, protibiae not S-shaped from extensor side.
</paragraph>
</subSubSection>
<subSubSection id="C1F10B8D5353FFC013FCB478FF6DFEF8" lastPageId="7" lastPageNumber="557" pageId="6" pageNumber="556" type="etymology">
<paragraph id="895458065353FFC113FCB478FC40F89F" blockId="6.[151,1437,1132,2023]" box="[199,1002,1888,1915]" pageId="6" pageNumber="556">
<emphasis id="BB9F84145353FFC113FCB478FEE4F89F" bold="true" box="[199,334,1888,1914]" pageId="6" pageNumber="556">Etymology.</emphasis>
The name derives from the Ustyurt plateau in Middle Asia.
</paragraph>
<paragraph id="895458065353FFC013FCB49CFA3BFF30" blockId="6.[151,1437,1132,2023]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="557" pageId="6" pageNumber="556">
<emphasis id="BB9F84145353FFC113FCB49CFE4BF87B" bold="true" box="[199,481,1924,1950]" pageId="6" pageNumber="556">Comparative diagnosis.</emphasis>
The new species is very similar to
<taxonomicName id="4EEB23855353FFC11057B49DFBA9F87B" baseAuthorityName="Bogatchev" baseAuthorityYear="1952" box="[876,1027,1924,1951]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="jachontovi">
<emphasis id="BB9F84145353FFC11057B49DFBA9F87B" box="[876,1027,1924,1951]" italics="true" pageId="6" pageNumber="556">C. jachontovi</emphasis>
</taxonomicName>
, but clearly differs from the latter in characters presented in a key below. In addition,
<taxonomicName id="4EEB23855353FFC1118BB4B1FCFCF827" authority="Bekchanov &amp; Nabozhenko &amp; Bekchanova, 2023" authorityName="Bekchanov &amp; Nabozhenko &amp; Bekchanova" authorityYear="2023" box="[688,854,1961,1987]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="ustyurtensis" status="sp. n.">
<emphasis id="BB9F84145353FFC1118BB4B1FCFCF827" box="[688,854,1961,1987]" italics="true" pageId="6" pageNumber="556">C. ustyurtensis</emphasis>
</taxonomicName>
<emphasis id="BB9F84145353FFC11060B4B0FC30F827" bold="true" box="[859,922,1960,1986]" pageId="6" pageNumber="556">
<taxonomicNameLabel id="A0AC396F5353FFC11060B4B0FC30F827" box="[859,922,1960,1986]" pageId="6" pageNumber="556" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4EEB23855353FFC117F8B4B1FAF3F827" baseAuthorityName="Bogatchev" baseAuthorityYear="1952" box="[1219,1369,1960,1987]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="jachontovi">
<emphasis id="BB9F84145353FFC117F8B4B1FAF3F827" box="[1219,1369,1960,1987]" italics="true" pageId="6" pageNumber="556">C. jachontovi</emphasis>
</taxonomicName>
in the black body with black or dark-brown legs (brown body with brown legs in
<taxonomicName id="4EEB23855353FFC110EFB4D5FBC3F803" baseAuthorityName="Bogatchev" baseAuthorityYear="1952" box="[980,1129,1996,2023]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="556" phylum="Arthropoda" rank="species" species="jachontovi">
<emphasis id="BB9F84145353FFC110EFB4D5FBC3F803" box="[980,1129,1996,2023]" italics="true" pageId="6" pageNumber="556">C. jachontovi</emphasis>
</taxonomicName>
) and the structure of blades in spiculum gastrale which are transverse and triangular in the new species (
<figureCitation id="11D044835352FFC010D9B38FFB9DFF54" box="[994,1079,151,177]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="7" pageNumber="557">Fig. 4F</figureCitation>
), but longitudinal and oval in
<taxonomicName id="4EEB23855352FFC016B8B38FFEA7FF30" baseAuthorityName="Bogatchev" baseAuthorityYear="1952" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="557" phylum="Arthropoda" rank="species" species="jachontovi">
<emphasis id="BB9F84145352FFC016B8B38FFEA7FF30" italics="true" pageId="7" pageNumber="557">C. jachontovi</emphasis>
</taxonomicName>
(
<figureCitation id="11D044835352FFC01226B3A3FED8FF30" box="[285,370,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1376,1400]" captionTargetBox="[152,1434,187,1344]" captionTargetId="figure-20@4.[151,1436,181,1351]" captionTargetPageId="4" captionText="FIGURE 2. Colposcelis jachontovi, details of structure. A = head dorsally; B = apical antennomeres, female; C = tegmen of aedeagus; D = median lobe of aedeagus; E = male inner sternite and tergite VIII; F = spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.10203168" httpUri="https://zenodo.org/record/10203168/files/figure.png" pageId="7" pageNumber="557">Fig. 2F</figureCitation>
). Both species are well different in the structure of aedeagus (compare
<figureCitation id="11D044835352FFC017B5B3A3FAA2FF30" box="[1166,1288,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1376,1400]" captionTargetBox="[152,1434,187,1344]" captionTargetId="figure-20@4.[151,1436,181,1351]" captionTargetPageId="4" captionText="FIGURE 2. Colposcelis jachontovi, details of structure. A = head dorsally; B = apical antennomeres, female; C = tegmen of aedeagus; D = median lobe of aedeagus; E = male inner sternite and tergite VIII; F = spiculum gastrale." figureDoi="http://doi.org/10.5281/zenodo.10203168" httpUri="https://zenodo.org/record/10203168/files/figure.png" pageId="7" pageNumber="557">Figs 2C, D</figureCitation>
and
<figureCitation id="11D044835352FFC01604B3A3FA2EFF30" box="[1343,1412,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,990,1014]" captionTargetBox="[155,1432,181,963]" captionTargetId="figure-20@6.[151,1436,181,965]" captionTargetPageId="6" captionText="FIGURE 4. Colposcelis ustyurtensis sp. n., details of structure. A = male protibia from the extensor side; B = ditto, dorsally; C = apical antennomeres, female; D = tegmen of aedeagus; E = median lobe of aedeagus; F = spiculum gastrale; G = male inner sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.10203174" httpUri="https://zenodo.org/record/10203174/files/figure.png" pageId="7" pageNumber="557">4D, E</figureCitation>
).
</paragraph>
<paragraph id="895458065352FFC013FCB3C6FF6DFEF8" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="557">
<emphasis id="BB9F84145352FFC013FCB3C6FEE9FF1D" bold="true" box="[199,323,222,248]" pageId="7" pageNumber="557">Bionomics</emphasis>
and photographs of habitats are given in
<bibRefCitation id="ED7A25F75352FFC0101BB3C7FB9EFF1C" author="Bekchanov, N. Kh. &amp; Nabozhenko, M. V. &amp; Bekchanov, Kh. U." box="[800,1076,223,249]" pageId="7" pageNumber="557" pagination="155 - 191" refId="ref7057" refString="Bekchanov, N. Kh., Nabozhenko, M. V. &amp; Bekchanov, Kh. U. (2023) Remarkable additions to the fauna of darkling beetles (Coleoptera: Tenebrionidae) of western Uzbekistan. Caucasian Entomological Bulletin, 19 (1), 155 - 191. https: // doi. org / 10.5281 / zenodo. 8199930" type="journal article" year="2023">
Bekchanov
<emphasis id="BB9F84145352FFC0109CB3C7FC73FF1C" box="[935,985,223,249]" italics="true" pageId="7" pageNumber="557">et al</emphasis>
. (2023)
</bibRefCitation>
(as
<taxonomicName id="4EEB23855352FFC0175EB3C7FA36FF1C" box="[1125,1436,223,249]" class="Insecta" family="Chrysomelidae" genus="Colposcelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="557" phylum="Arthropoda" rank="subGenus" subGenus="Turcmenicola">
<emphasis id="BB9F84145352FFC0175EB3C7FB42FF1C" box="[1125,1256,223,249]" italics="true" pageId="7" pageNumber="557">Colposcelis</emphasis>
(
<emphasis id="BB9F84145352FFC017C1B3C7FA39FF1C" box="[1274,1427,223,249]" italics="true" pageId="7" pageNumber="557">Turcmenicola</emphasis>
)
</taxonomicName>
sp.).
</paragraph>
</subSubSection>
</treatment>
</document>
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