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<document id="E535440BF4F471BEC38CC8286AC89221" ID-DOI="10.11646/zootaxa.5019.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222728" ID-ZooBank="241751BA-E759-49B8-A394-271B2A4650AD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629385249404" checkinUser="plazi" docAuthor="Singh, Sudhir, Colombo, Wesley D., Shreevihar, Santhosh, Pandey, Vijay P. &amp; Tribull, Carly M." docDate="2021" docId="03ED87EE3B33446931EDFD79FC56C0C3" docLanguage="en" docName="zootaxa.5019.1.1.pdf" docOrigin="Zootaxa 5019 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5019.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Goniozus madrassippattanami Kurian 1955" docType="treatment" docVersion="5" lastPageNumber="16" masterDocId="FFD4FF963B3C4466317AFFAFFFCDC638" masterDocTitle="Rediscovery of Kurians types of Hymenoptera at Forest Research Institute, Dehradun, India with notes on Indian bethylid fauna" masterLastPageNumber="90" masterPageNumber="1" pageNumber="16" updateTime="1699070948635" updateUser="plazi">
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<mods:title id="C5B4071A7A9A8998DB32A1D9455C8A29">Rediscovery of Kurians types of Hymenoptera at Forest Research Institute, Dehradun, India with notes on Indian bethylid fauna</mods:title>
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<mods:namePart id="CF3BA59B9A9326407557DD8143A6BC24">Colombo, Wesley D.</mods:namePart>
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<mods:namePart id="AAABBF08F5D49ECE758ECD44B193BAD9">Pandey, Vijay P.</mods:namePart>
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<emphasis id="B930EAEA3B33446931EDFD79FD1DC4C8" bold="true" box="[151,720,726,753]" pageId="15" pageNumber="16">
6.
<taxonomicName id="4C444D7B3B33446931CFFD78FD1DC4C8" authority="Kurian, 1955" authorityName="Kurian" authorityYear="1955" box="[181,720,726,753]" class="Insecta" family="Bethylidae" genus="Goniozus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="madrassippattanami">
<emphasis id="B930EAEA3B33446931CFFD78FDE9C4C9" bold="true" box="[181,548,726,753]" italics="true" pageId="15" pageNumber="16">Goniozus madrassippattanami</emphasis>
<bibRefCitation id="EFD54B093B3344693351FD79FD1DC4C8" author="Kurian, C." box="[555,720,726,752]" pageId="15" pageNumber="16" pagination="67 - 155" refId="ref41065" refString="Kurian, C. (1955) Bethyloidea (Hymenoptera) from Indian. Agra University Journal of Research, 4, 67 - 155." type="journal article" year="1955">Kurian, 1955</bibRefCitation>
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</emphasis>
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<paragraph id="8BFB36F83B33446931EDFD54FE55C52D" blockId="15.[151,720,726,789]" box="[151,408,762,789]" pageId="15" pageNumber="16">
<figureCitation id="137F2A7D3B33446931EDFD54FF13C52D" box="[151,222,763,789]" captionStart="FIGURE 6" captionStartId="16.[151,250,1888,1912]" captionTargetBox="[168,1423,181,1860]" captionTargetId="figure-19@16.[164,1423,181,1864]" captionTargetPageId="16" captionText="FIGURE 6. AM. Goniozus madrassippattanami Kurian, 1955. AF. Holotype. A. Head, frontal view; B. Mesosoma, dorsal view; C. Antenna; D. Forewing; E. Legs (left to right) pro and metalegs; F. Metasoma, dorsal view. GK. Paratype.G. Mesosoma with forewing, lateral view; H. Mesosoma, dorsal view; I. Mesosoma, lateral view; J. Metapectal-propodeal disc, dorsal view; K. Metasoma, dorso-lateral view. L. Two slides of holotype. M. Labels of paratype." figureDoi="http://doi.org/10.5281/zenodo.5222740" httpUri="https://zenodo.org/record/5222740/files/figure.png" pageId="15" pageNumber="16">Figs 6</figureCitation>
, AM;
<figureCitation id="137F2A7D3B3344693049FD55FE9CC52D" box="[307,337,762,789]" captionStart="FIGURE 51" captionStartId="82.[151,250,1824,1848]" captionTargetBox="[329,1259,181,1800]" captionTargetId="figure-19@82.[329,1259,181,1800]" captionTargetPageId="82" captionText="FIGURE 51. AN. Male genitalia. AB. Goniozus madrassippattanami Kurian, 1955, paratype. A. dorsal view; B. ventral view; CD. Goniozus salvadorae Kurian, 1954b, paratype; C. dorsal view; D. ventral view; EF. Goniozus thalasodi Kurian, 1955, holotype; E. dorsal view; F. ventral view; GH. Epyris indicae (Kurian, 1955), holotype; G. dorsal view; H. ventral view; IJ. Epyris manii Kurian, 1955, holotype; I. dorsal view; J. ventral view; KL. Holepyris patnae (Kurian, 1955), holotype; K. dorsal view; L. ventral view; MN. Pristocera agraensis (Kurian, 1955), holotype; M. dorsal view; N. ventral view." figureDoi="http://doi.org/10.5281/zenodo.5222843" httpUri="https://zenodo.org/record/5222843/files/figure.png" pageId="15" pageNumber="16">51</figureCitation>
, AB.
</paragraph>
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<taxonomicName id="4C444D7B3B33446931EDFCEFFD91C56F" authority="Kurian, 1955" authorityName="Kurian" authorityYear="1955" box="[151,604,831,855]" class="Insecta" family="Bethylidae" genus="Goniozus" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="madrassippattanami">
<emphasis id="B930EAEA3B33446931EDFCEFFE00C56F" box="[151,461,831,855]" italics="true" pageId="15" pageNumber="16">Goniozus madrassippattanami</emphasis>
<bibRefCitation id="EFD54B093B33446930AEFC90FD91C56F" author="Kurian, C." box="[468,604,831,855]" pageId="15" pageNumber="16" pagination="67 - 155" refId="ref41065" refString="Kurian, C. (1955) Bethyloidea (Hymenoptera) from Indian. Agra University Journal of Research, 4, 67 - 155." type="journal article" year="1955">Kurian, 1955</bibRefCitation>
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, 4: 114, 122, 123, Figs: 206210.
</paragraph>
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<paragraph id="8BFB36F83B33446931EDFCF1FC98C54E" blockId="15.[151,944,831,886]" box="[151,853,862,886]" pageId="15" pageNumber="16">
<materialsCitation id="3B2C3CA53B33446931EDFCF1FC9CC54E" box="[151,849,862,886]" collectionCode="NFIC-FRI" country="India" location="Nilambur" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu" typeStatus="holotype">
Type locality:
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,
<collectingRegion id="4980F81A3B33446930EFFCF1FDDDC54E" box="[405,528,862,886]" country="India" name="Tamil Nadu" pageId="15" pageNumber="16">Tamil Nadu</collectingRegion>
,
<collectingCountry id="F35376683B3344693360FCF1FD83C54E" box="[538,590,862,886]" name="India" pageId="15" pageNumber="16">India</collectingCountry>
(
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<typeStatus id="54FF885A3B334469330CFCF1FD1CC54E" box="[630,721,862,886]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED55AE3D3B33446933A6FCF1FC86C54E" box="[732,843,862,886]" pageId="15" pageNumber="16">NFIC-FRI</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<subSubSection id="C35E65733B33446931EDFC0BFC41C31E" pageId="15" pageNumber="16" type="diagnosis">
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<emphasis id="B930EAEA3B33446931EDFC0BFEC7C586" bold="true" box="[151,266,932,958]" pageId="15" pageNumber="16">Diagnosis</emphasis>
. Female. Length 2.0 mm2.6 mm. General colour dark reddish-brown to black. Head as long as wide; frons polished; ocelli distinct; eye glabrous; median clypeal lobe conspicuous, anterior margin rounded, longer than lateral ones; median clypeal carina high in lateral view; antenna with sparse pubescence, scape 1.5 × longer than pedicel, pedicel as long as first flagellomere. Pronotum weakly coriaceous; mesonotum longer than pronotal dorsal area; notauli inconspicuous. Metapectal-propodeal disc wider than long; metapostnotal median carina absent; first abdominal spiracle dorsally placed. Tarsal claw simple. Wings hyaline; forewing with pterostigma dark brown, 2R1
<subScript id="17C034BD3B33446931BCFB21FF03C2A4" attach="right" box="[198,206,1166,1180]" fontSize="6" pageId="15" pageNumber="16">2</subScript>
c cell opened, Rs+M
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v vein present, 1M
<subScript id="17C034BD3B33446933D2FB21FD7DC2A4" attach="both" box="[680,688,1166,1180]" fontSize="6" pageId="15" pageNumber="16">2</subScript>
c opened. Metasoma polished. Male. Morphologically similar to female, except for genitalia: Harpe completely divided into dorsal and ventral arms, shorter than gonostipes in dorsal view, dorsal arm with apex rounded, weakly arched inward; gonostipes wide, covering aedeagal base; apex of cuspis anterior to apex of dorsal arm of harpe; digitus with apex aligned to apex of cuspis, its posterior margin smooth; aedeagus bottle-shaped, its apex aligned to apex of digitus.
</paragraph>
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<emphasis id="B930EAEA3B33446931BDFA9FFE7EC372" bold="true" box="[199,435,1328,1354]" pageId="15" pageNumber="16">Material examined.</emphasis>
<materialsCitation id="3B2C3CA53B33446930C6FA9FFD44C357" collectorName="Madras, A. H" country="India" location="Kerala" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="54FF885A3B33446930C6FA9FFDE4C372" box="[444,553,1328,1354]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
: [
<collectingCountry id="F35376683B3344693341FA9FFD5DC372" box="[571,656,1328,1354]" name="India" pageId="15" pageNumber="16">INDIA</collectingCountry>
,
<emphasis id="B930EAEA3B33446933E4FA9FFD21C372" box="[670,748,1328,1354]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="4980F81A3B33446933E4FA9FFD21C372" box="[670,748,1328,1354]" country="India" name="Kerala" pageId="15" pageNumber="16">Kerala</collectingRegion>
</emphasis>
,] Nilambur [
<emphasis id="B930EAEA3B33446932F9FA9FFBC5C372" box="[899,1032,1328,1354]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="4980F81A3B33446932F9FA9FFBC5C372" box="[899,1032,1328,1354]" country="India" name="Tamil Nadu" pageId="15" pageNumber="16">Tamil Nadu</collectingRegion>
</emphasis>
,]
<collectorName id="26B1532E3B334469355BFA9FFB7BC372" box="[1057,1206,1328,1354]" pageId="15" pageNumber="16">Madras, A.H</collectorName>
. Khan [coll.], 1939, parasitic
<collectedFrom id="385875053B3344693187FAFAFD98C357" box="[253,597,1364,1391]" pageId="15" pageNumber="16">
on
<taxonomicName id="4C444D7B3B334469305AFAFBFD98C357" authority=", RRD" authorityName="RRD" box="[288,597,1364,1391]" class="Insecta" family="Crambidae" genus="Hapalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="machaeralis">
<emphasis id="B930EAEA3B334469305AFAFBFDC0C356" box="[288,525,1364,1390]" italics="true" pageId="15" pageNumber="16">Hapalia machaeralis</emphasis>
, RRD
</taxonomicName>
</collectedFrom>
598
</materialsCitation>
.
<materialsCitation id="3B2C3CA53B33446933EEFAFAFBEFC356" box="[660,1058,1364,1391]" collectorName="Madras, A. H" country="India" location="Kerala" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="paratype">
<typeStatus id="54FF885A3B33446933EEFAFAFD37C357" box="[660,762,1365,1391]" pageId="15" pageNumber="16" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D42FD713B334469327FFAFAFCD1C357" box="[773,796,1365,1391]" pageId="15" pageNumber="16" type="male"></specimenCount>
, same data of
<typeStatus id="54FF885A3B33446932C6FAFBFBEFC356" box="[956,1058,1364,1390]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
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.
</paragraph>
<paragraph id="8BFB36F83B33446931BDFAD7FE28C052" blockId="15.[151,1437,932,1787]" pageId="15" pageNumber="16">
<emphasis id="B930EAEA3B33446931BDFAD7FE41C3AA" bold="true" box="[199,396,1400,1426]" pageId="15" pageNumber="16">Type conditions.</emphasis>
<typeStatus id="54FF885A3B33446930EEFAD7FE31C3AA" box="[404,508,1400,1426]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
completely dissected
<taxonomicName id="4C444D7B3B334469338DFAD6FC9DC3AA" box="[759,848,1400,1427]" class="Insecta" family="Bethylidae" genus="Odontepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="73" phylum="Arthropoda" rank="species" species="slide">on slide</taxonomicName>
Nº. 123/B.
<taxonomicName id="4C444D7B3B33446932ADFAD7FBFBC3AA" box="[983,1078,1400,1426]" class="Insecta" family="Bethylidae" genus="Odontepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="slide">
<emphasis id="B930EAEA3B33446932ADFAD7FBFBC3AA" box="[983,1078,1400,1426]" italics="true" pageId="15" pageNumber="16">On slide</emphasis>
</taxonomicName>
: S1 C1—two legs, one without coxae; distal half of forewing; one antenna complete. S1 C2—one forewing and one hind wing. S2 C1—parts of six legs; two forewings. S2 C2—head; mesosoma; metasoma; one hind wing; one proleg without coxa and tarsi; one leg without coxae and femur.
<typeStatus id="54FF885A3B334469309FFA4AFD9FC3C7" box="[485,594,1509,1535]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
on two cards on a pin with labels 123/B.
<emphasis id="B930EAEA3B334469355CFA4BFBB8C3C6" box="[1062,1141,1508,1534]" italics="true" pageId="15" pageNumber="16">Pinned</emphasis>
: head missing; mesosoma with both pairs of wings; one leg and some coxae. Metasoma is mounted on a card on another pin, with the label “123/B,
<typeStatus id="54FF885A3B334469318DF983FE93C07E" box="[247,350,1580,1606]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
”.
<taxonomicName id="4C444D7B3B3344693009F983FE1FC07E" box="[371,466,1580,1606]" class="Insecta" family="Bethylidae" genus="Odontepyris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="slide">
<emphasis id="B930EAEA3B3344693009F983FE1FC07E" box="[371,466,1580,1606]" italics="true" pageId="15" pageNumber="16">On slide</emphasis>
</taxonomicName>
: S1 C3—genitalia was dissected out from the card mounted meatasoma and mounted under a rectangular coverslip.
</paragraph>
</subSubSection>
<subSubSection id="C35E65733B33446931BDF9DBFD10C0EE" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BFB36F83B33446931BDF9DBFD10C0EE" blockId="15.[151,1437,932,1787]" pageId="15" pageNumber="16">
<emphasis id="B930EAEA3B33446931BDF9DBFEF6C0B6" bold="true" box="[199,315,1652,1678]" pageId="15" pageNumber="16">Remarks.</emphasis>
<bibRefCitation id="EFD54B093B3344693039F9DBFE29C0B7" author="Kurian, C." box="[323,484,1652,1679]" pageId="15" pageNumber="16" pagination="67 - 155" refId="ref41065" refString="Kurian, C. (1955) Bethyloidea (Hymenoptera) from Indian. Agra University Journal of Research, 4, 67 - 155." type="journal article" year="1955">Kurian (1955)</bibRefCitation>
described the
<typeStatus id="54FF885A3B33446933F0F9DBFD21C0B6" box="[650,748,1652,1678]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
as male, and labeled the slides of the
<typeStatus id="54FF885A3B33446935DAF9DBFACFC0B6" box="[1184,1282,1652,1678]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
also as male, but the dissection of the metasoma confirmed it as female. The dissected ovipositor and last tergites of the
<typeStatus id="54FF885A3B3344693440F937FA51C08A" box="[1338,1436,1688,1714]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
are mounted on a small recangular cover slip on S1.
</paragraph>
</subSubSection>
<subSubSection id="C35E65733B33446931BDF94FFC56C0C3" box="[199,923,1760,1787]" pageId="15" pageNumber="16" type="biology_ecology">
<paragraph id="8BFB36F83B33446931BDF94FFC56C0C3" blockId="15.[151,1437,932,1787]" box="[199,923,1760,1787]" pageId="15" pageNumber="16">
<emphasis id="B930EAEA3B33446931BDF94FFEDCC0C2" bold="true" box="[199,273,1760,1786]" pageId="15" pageNumber="16">Hosts.</emphasis>
<taxonomicName id="4C444D7B3B3344693062F94FFE69C0C2" authorityName="Linnaeus" authorityYear="1758" box="[280,420,1760,1786]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
:
<taxonomicName id="4C444D7B3B33446930D4F94FFDFFC0C2" authorityName="Hampson" authorityYear="1903" box="[430,562,1760,1786]" class="Insecta" family="Hyblaeidae" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Hyblaeidae</taxonomicName>
:
<taxonomicName id="4C444D7B3B3344693346F94FFC5AC0C3" authority="(Walker)" baseAuthorityName="Walker" box="[572,919,1760,1787]" class="Insecta" family="Crambidae" genus="Hapalia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="machaeralis">
<emphasis id="B930EAEA3B3344693346F94FFCE4C0C2" box="[572,809,1760,1786]" italics="true" pageId="15" pageNumber="16">Hapalia machaeralis</emphasis>
(Walker)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>