treatments-xml/data/03/EB/87/03EB87F8F17CFFA5FF31FB0D7D824E96.xml

<document id="F88EDA91F0622AFD9BF53BF05363C009" ID-CLB-Dataset="28034" ID-DOI="10.11646/zootaxa.4508.4.1" ID-GBIF-Dataset="a7ff59ca-84b3-4132-8a5d-367b14c84cee" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607312" ID-ZooBank="D268E210-6688-4B41-9A76-AE54C2FC07BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586215440" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2018" docId="03EB87F8F17CFFA5FF31FB0D7D824E96" docLanguage="en" docName="zootaxa.4508.4.1.pdf" docOrigin="Zootaxa 4508 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium besucheti Shavrin &amp; Smetana 2018, sp.n." docType="treatment" docVersion="8" lastPageNumber="467" masterDocId="FFD2FF80F173FFB5FFA6FFEC7C18480E" masterDocTitle="A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group" masterLastPageNumber="506" masterPageNumber="451" pageNumber="466" updateTime="1698713119729" updateUser="ExternalLinkService">
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<mods:title id="F68A16907ED979A5522774E0AE05A745">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). II. fusculum group, and two additional species of the nigrum group</mods:title>
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<mods:namePart id="6B54EB4AEAE1324F7381708DBA476431">Shavrin, Alexey V.</mods:namePart>
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<mods:namePart id="02B29E085225F34252D7EDDA668162FE">Smetana, Aleš</mods:namePart>
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<treatment id="03EB87F8F17CFFA5FF31FB0D7D824E96" ID-DOI="http://doi.org/10.5281/zenodo.6495083" ID-GBIF-Taxon="154496004" ID-Zenodo-Dep="6495083" LSID="urn:lsid:plazi:treatment:03EB87F8F17CFFA5FF31FB0D7D824E96" httpUri="http://treatment.plazi.org/id/03EB87F8F17CFFA5FF31FB0D7D824E96" lastPageId="16" lastPageNumber="467" pageId="15" pageNumber="466">
<subSubSection id="C3586565F17CFFBAFF31FB0D7EF94CF5" box="[151,737,1249,1275]" pageId="15" pageNumber="466" type="nomenclature">
<paragraph id="8BFD36EEF17CFFBAFF31FB0D7EF94CF5" blockId="15.[151,737,1249,1309]" box="[151,737,1249,1275]" pageId="15" pageNumber="466">
<heading id="D0B58182F17CFFBAFF31FB0D7EF94CF5" bold="true" box="[151,737,1249,1275]" fontSize="11" level="1" pageId="15" pageNumber="466" reason="1">
<emphasis id="B936EAFCF17CFFBAFF31FB0D7EF94CF5" bold="true" box="[151,737,1249,1275]" pageId="15" pageNumber="466">
<taxonomicName id="4C424D6DF17CFFBAFF31FB0D7E814CF5" ID-CoL="67CCS" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[151,665,1249,1275]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">
<emphasis id="B936EAFCF17CFFBAFF31FB0D7D814CF5" bold="true" box="[151,409,1249,1275]" italics="true" pageId="15" pageNumber="466">Anthobium besucheti</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
, 
<taxonomicNameLabel id="A2055787F17CFFBAFD00FB0D7EF94CF5" box="[678,737,1249,1275]" pageId="15" pageNumber="466" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586565F17CFFBAFF31FAE87D6C4D13" box="[151,372,1284,1309]" pageId="15" pageNumber="466" type="description">
<paragraph id="8BFD36EEF17CFFBAFF31FAE87D6C4D13" blockId="15.[151,737,1249,1309]" box="[151,372,1284,1309]" pageId="15" pageNumber="466">
(
<figureCitation id="13792A6BF17CFFBAFF39FAE87CF54D12" box="[159,237,1284,1309]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1628,1651]" captionTargetBox="[237,1323,200,1594]" captionTargetId="figure@5.[150,1436,195,1604]" captionTargetPageId="5" captionText="FIGURES 1–5. Habitus of Anthobium fusculum group: 1—A. fusculum (KAZAKHSTAN: Tian Shan), 2—A. altivagans (NEPAL: Telbrung Danda), 3—A. besucheti (paratype: Ghoropani Pass), 4—A. capitale (holotype), 5—A. cavicrus (NEPAL: Thare Pati). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607314/files/figure.png" pageId="15" pageNumber="466">Figs. 3</figureCitation>
, 
<figureCitation id="13792A6BF17CFFBAFF5CFAE87D5B4D13" box="[250,323,1284,1309]" captionStart="FIGURES 50–53" captionStartId="14.[151,264,1972,1995]" captionTargetBox="[386,1272,196,1956]" captionTargetId="figure@14.[231,1360,193,1960]" captionTargetPageId="14" captionText="FIGURES 50–53. Aedeagus of Anthobium altivagans (50–51: NEPAL: Ganesh Himal) and A. besucheti (52–53: NEPAL: Ghoropani Pass): 50, 52—parameral view, 51, 53—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607328/files/figure.png" pageId="15" pageNumber="466">52–53</figureCitation>
, 
<figureCitation id="13792A6BF17CFFBAFEE8FAE87D734D13" box="[334,363,1284,1309]" captionStart="FIGURE 54" captionStartId="15.[151,250,982,1004]" captionTargetBox="[179,1408,193,961]" captionTargetId="figure@15.[179,1408,193,961]" captionTargetPageId="15" captionText="FIGURE 54. Distribution of Anthobium altivagans (black circles), A. besucheti (black squares), A. rugosum (black triangles) and A. denticulatum (black star) in the Himalayan Region." httpUri="https://zenodo.org/record/2607330/files/figure.png" pageId="15" pageNumber="466">54</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3586565F17CFFBAFF31FAA079494E8A" pageId="15" pageNumber="466" type="materials_examined">
<paragraph id="8BFD36EEF17CFFBAFF31FAA07F6E4DA2" blockId="15.[151,1437,1356,2029]" pageId="15" pageNumber="466">
<emphasis id="B936EAFCF17CFFBAFF31FAA07DDE4D6B" bold="true" box="[151,454,1356,1381]" pageId="15" pageNumber="466">Type material examined:</emphasis>
<materialsCitation id="3B2A3CB3F17CFFBAFE69FAA07EFA4D86" ID-GBIF-Occurrence="2028139745" collectingDate="1983-10-07" collectorName="Distr. Ridge E &amp; Smetana &amp; Lobl'" country="Nepal" elevation="3100" location="Parbat" municipality="Ghoropani Pass" pageId="15" pageNumber="466" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54F9884CF17CFFBAFE69FAA07E2F4D6B" box="[463,567,1356,1381]" pageId="15" pageNumber="466" type="holotype">Holotype</typeStatus>
<specimenCount id="9D44FD67F17CFFBAFDE7FAA17E414D6A" box="[577,601,1357,1380]" pageId="15" pageNumber="466" type="male">♂</specimenCount>
: ‘ 
<collectingCountry id="F355767EF17CFFBAFDCDFAA07ED04D6A" box="[619,712,1356,1380]" name="Nepal" pageId="15" pageNumber="466">NEPAL</collectingCountry>
<location id="8E9D6035F17CFFBAFD74FAA07F024D6B" LSID="urn:lsid:plazi:treatment:03EB87F8F17CFFA5FF31FB0D7D824E96:8E9D6035F17CFFBAFD74FAA07F024D6B" box="[722,794,1356,1381]" country="Nepal" municipality="Ghoropani Pass" name="Parbat" pageId="15" pageNumber="466">Parbat</location>
| 
<collectorName id="26B75338F17CFFBAFC94FAA07FCE4D6A" box="[818,982,1356,1381]" pageId="15" pageNumber="466">Distr. Ridge E</collectorName>
| 
<collectingMunicipality id="6B99AC94F17CFFBAFC49FAA078BA4D6A" box="[1007,1186,1356,1381]" pageId="15" pageNumber="466">Ghoropani Pass</collectingMunicipality>
| 
<quantity id="4CBA9B0BF17CFFBAFB1CFAA079104D6A" box="[1210,1288,1356,1380]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="466" unit="m" value="3100.0">
<elevation id="006FD1DDF17CFFBAFB1CFAA079104D6A" box="[1210,1288,1356,1380]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="466" unit="m" value="3100.0">3100m</elevation>
</quantity>
<date id="FFFC102EF17CFFBAFAB4FAA179954D6A" box="[1298,1421,1356,1380]" pageId="15" pageNumber="466" value="1983-10-07">
<collectingDate id="EFB8E9C6F17CFFBAFAB4FAA179954D6A" box="[1298,1421,1356,1380]" pageId="15" pageNumber="466" value="1983-10-07">7. X. 1983</collectingDate>
</date>
| [A.] 
<collectorName id="26B75338F17CFFBAFF1BFA9C7D3D4D86" box="[189,293,1392,1416]" pageId="15" pageNumber="466">Smetana</collectorName>
&amp; [I.] 
<collectorName id="26B75338F17CFFBAFEC1FA837DB44D86" box="[359,428,1391,1416]" pageId="15" pageNumber="466">Löbl’</collectorName>
&lt;rectangular label, printed
</materialsCitation>
&gt;, ‘ 
<materialsCitation id="3B2A3CB3F17CFFBAFCA7FA9C7F6A4DA2" ID-GBIF-Occurrence="2028139740" collectionCode="MHNG" collectorName="Shavrin A. &amp; Smetana A." country="Nepal" pageId="15" pageNumber="466" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F9884CF17CFFBAFCA7FA9C7F824D86" box="[769,922,1392,1416]" pageId="15" pageNumber="466" type="holotype">HOLOTYPE</typeStatus>
| 
<taxonomicName id="4C424D6DF17CFFBAFC16FA8378374D86" authorityName="Leach" authorityYear="1819" box="[944,1071,1391,1416]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="505" phylum="Arthropoda" rank="genus">Anthobium</taxonomicName>
| besucheti 
<emphasis id="B936EAFCF17CFFBAFB10FA8378F64D86" bold="true" box="[1206,1262,1391,1416]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="A2055787F17CFFBAFB10FA8378F64D86" box="[1206,1262,1391,1416]" pageId="15" pageNumber="466" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| 
<collectorName id="26B75338F17CFFBAFAA2FA8379664D86" box="[1284,1406,1391,1416]" pageId="15" pageNumber="466">Shavrin A.</collectorName>
&amp; 
<collectorName id="26B75338F17CFFBAFF31FA787D024DA2" box="[151,282,1428,1452]" pageId="15" pageNumber="466">Smetana A.</collectorName>
des. 2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="ED53AE2BF17CFFBAFCA8FA787F724DA2" box="[782,874,1428,1452]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="466" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFD36EEF17CFFBAFF61FA5479494E8A" blockId="15.[151,1437,1356,2029]" pageId="15" pageNumber="466">
<typeStatus id="54F9884CF17CFFBAFF61FA547D2C4DDE" box="[199,308,1464,1488]" pageId="15" pageNumber="466" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D44FD67F17CFFBAFEE1FA547DF24DDE" box="[327,490,1463,1488]" pageId="15" pageNumber="466" type="generic">23 specimens</specimenCount>
): 
<specimenCount id="9D44FD67F17CFFBAFE59FA5B7E5A4DC1" box="[511,578,1463,1488]" pageId="15" pageNumber="466" type="male">6 ♂♂</specimenCount>
[two specimens dissected], 
<materialsCitation id="3B2A3CB3F17CFFBAFC2BFA5B7DA34DFA" ID-GBIF-Occurrence="2028139751" collectionCode="CS" collectorName="Shavrin A. &amp; Smetana A." country="Nepal" pageId="15" pageNumber="466" specimenCount="18" specimenCount-female="12" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="9D44FD67F17CFFBAFC2BFA5B7FCA4DC1" box="[909,978,1463,1488]" pageId="15" pageNumber="466" type="female">6 ♀♀</specimenCount>
: same data as the holotype (
<specimenCount id="9D44FD67F17CFFBAFA80FA5B794F4DC1" box="[1318,1367,1463,1488]" pageId="15" pageNumber="466" type="male">5 ♂</specimenCount>
, 
<specimenCount id="9D44FD67F17CFFBAFAC0FA5B798E4DC1" box="[1382,1430,1463,1488]" pageId="15" pageNumber="466" type="female">5 ♀</specimenCount>
: MHNG; 
<specimenCount id="9D44FD67F17CFFBAFEA4FA307D2C4DFA" box="[258,308,1500,1524]" pageId="15" pageNumber="466" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D44FD67F17CFFBAFEE0FA307D614DFA" box="[326,377,1500,1524]" pageId="15" pageNumber="466" type="female">1 ♀</specimenCount>
: 
<collectionCode id="ED53AE2BF17CFFBAFE2AFA307DAA4DFA" box="[396,434,1500,1524]" country="France" name="Musee des Dinosaures d'Esperaza (Aude)" pageId="15" pageNumber="466">CS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2A3CB3F17CFFBAFE6BFA307ED64E16" ID-GBIF-Occurrence="2028139750" collectingDate="2013-09-21" collectionCode="NME" country="Nepal" elevation="3140" latitude="28.3825" location="Sundar Danda" longLatPrecision="20" longitude="84.34917" municipality="Lamjung" pageId="15" pageNumber="466" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D44FD67F17CFFBAFE6BFA307E184DFA" box="[461,512,1500,1524]" pageId="15" pageNumber="466" type="male">1 ♂</specimenCount>
: ‘ 
<collectingCountry id="F355767EF17CFFBAFDB1FA307E604DFA" box="[535,632,1500,1524]" name="Nepal" pageId="15" pageNumber="466">NEPAL</collectingCountry>
: 
<collectingMunicipality id="6B99AC94F17CFFBAFD2FFA307EE94DFB" box="[649,753,1500,1525]" pageId="15" pageNumber="466">Lamjung</collectingMunicipality>
, S- | 
<location id="8E9D6035F17CFFBAFC9AFA307FFD4DFB" LSID="urn:lsid:plazi:treatment:03EB87F8F17CFFA5FF31FB0D7D824E96:8E9D6035F17CFFBAFC9AFA307FFD4DFB" box="[828,997,1500,1525]" country="Nepal" latitude="28.3825" longLatPrecision="20" longitude="84.34917" municipality="Lamjung" name="Sundar Danda" pageId="15" pageNumber="466">Sundar Danda</location>
, 
<quantity id="4CBA9B0BF17CFFBAFC50FA30785F4DFA" box="[1014,1095,1500,1524]" metricMagnitude="3" metricUnit="m" metricValue="3.1399999999999997" pageId="15" pageNumber="466" unit="m" value="3140.0">
<elevation id="006FD1DDF17CFFBAFC50FA30785F4DFA" box="[1014,1095,1500,1524]" metricMagnitude="3" metricUnit="m" metricValue="3.1399999999999997" pageId="15" pageNumber="466" unit="m" value="3140.0">3140m</elevation>
</quantity>
, | 
<geoCoordinate id="EE765029F17CFFBAFBCAFA3078E84DFB" box="[1132,1264,1500,1525]" direction="north" orientation="latitude" pageId="15" pageNumber="466" precision="15" value="28.3825">28°22'57''N</geoCoordinate>
, 
<geoCoordinate id="EE765029F17CFFBAFAA5FA30799D4DFB" box="[1283,1413,1500,1525]" direction="east" orientation="longitude" pageId="15" pageNumber="466" precision="15" value="84.34917">84°20'57''E</geoCoordinate>
, | 
<date id="FFFC102EF17CFFBAFF31F9EC7D0F4E16" box="[151,279,1536,1560]" pageId="15" pageNumber="466" value="2013-09-21">
<collectingDate id="EFB8E9C6F17CFFBAFF31F9EC7D0F4E16" box="[151,279,1536,1560]" pageId="15" pageNumber="466" value="2013-09-21">21.IX.2013</collectingDate>
</date>
| leg. [J.]Hagge &amp; [J.]Schmidt’ (
<collectionCode id="ED53AE2BF17CFFBAFD24F9EC7EDE4E16" box="[642,710,1536,1560]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="15" pageNumber="466">NME</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B2A3CB3F17CFFBAFD7FF9EC7EF94E32" ID-GBIF-Occurrence="2028139741" collectingDate="1984-04-09" collectionCode="MHNG" country="Nepal" elevation="3600" location="Goru Dzure Dara" pageId="15" pageNumber="466" specimenCount="1" specimenCount-female="1" stateProvince="Kosi" typeStatus="paratype">
<specimenCount id="9D44FD67F17CFFBAFD7FF9EC7F1C4E19" box="[729,772,1536,1560]" pageId="15" pageNumber="466" type="female">1 ♀</specimenCount>
: ‘E. 
<collectingCountry id="F355767EF17CFFBAFC9EF9EC7F8C4E16" box="[824,916,1536,1560]" name="Nepal" pageId="15" pageNumber="466">NEPAL</collectingCountry>
: 
<collectingRegion id="4986F80CF17CFFBAFC38F9EC7FC74E16" box="[926,991,1536,1560]" country="Nepal" name="Kosi" pageId="15" pageNumber="466">KOSI</collectingRegion>
[Khandbari District] | 
<location id="8E9D6035F17CFFBAFB7DF9EC79834E19" LSID="urn:lsid:plazi:treatment:03EB87F8F17CFFA5FF31FB0D7D824E96:8E9D6035F17CFFBAFB7DF9EC79834E19" box="[1243,1435,1536,1560]" country="Nepal" name="Goru Dzure Dara" pageId="15" pageNumber="466" stateProvince="Kosi">Goru Dzure Dara</location>
| 
<quantity id="4CBA9B0BF17CFFBAFF03F9C87CE64E32" box="[165,254,1572,1597]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="15" pageNumber="466" unit="m" value="3600.0">
<elevation id="006FD1DDF17CFFBAFF03F9C87CE64E32" box="[165,254,1572,1597]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="15" pageNumber="466" unit="m" value="3600.0">3600 m</elevation>
</quantity>
, 
<date id="FFFC102EF17CFFBAFEADF9C87D454E32" box="[267,349,1572,1596]" pageId="15" pageNumber="466" value="1984-04-09">
<collectingDate id="EFB8E9C6F17CFFBAFEADF9C87D454E32" box="[267,349,1572,1596]" pageId="15" pageNumber="466" value="1984-04-09">9.IV.84</collectingDate>
</date>
| [I.]Löbl-[A.]Smetana’ (
<collectionCode id="ED53AE2BF17CFFBAFDDBF9C87EC14E32" box="[637,729,1572,1596]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="466" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
; 
<specimenCount id="9D44FD67F17CFFBAFD48F9C87F374E32" box="[750,815,1572,1596]" pageId="15" pageNumber="466" type="male">4 ♂♂</specimenCount>
[one specimen dissected], 
<materialsCitation id="3B2A3CB3F17CFFBAFBC2F9C878044E6E" ID-GBIF-Occurrence="2028139728" collectingDate="1983-10-07" collectionCode="MHNG" country="Nepal" elevation="3100" location="Crete Est Goropani" municipality="For." pageId="15" pageNumber="466" specimenCount="5" specimenCount-female="5" stateProvince="Kosi" typeStatus="paratype">
<specimenCount id="9D44FD67F17CFFBAFBC2F9C878BF4E32" box="[1124,1191,1572,1597]" pageId="15" pageNumber="466" type="female">5 ♀♀</specimenCount>
: ‘NEPAL—Goropani 
<collectingMunicipality id="6B99AC94F17CFFBAFF31F9A47CDC4E51" box="[151,196,1608,1631]" pageId="15" pageNumber="466">For.</collectingMunicipality>
| 
<location id="8E9D6035F17CFFBAFF7FF9A47DAB4E6E" LSID="urn:lsid:plazi:treatment:03EB87F8F17CFFA5FF31FB0D7D824E96:8E9D6035F17CFFBAFF7FF9A47DAB4E6E" box="[217,435,1607,1632]" country="Nepal" municipality="For." name="Crete Est Goropani" pageId="15" pageNumber="466" stateProvince="Kosi">Crête Est Goropani</location>
| 
<quantity id="4CBA9B0BF17CFFBAFE6FF9A47E0F4E6E" box="[457,535,1608,1632]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="466" unit="m" value="3100.0">
<elevation id="006FD1DDF17CFFBAFE6FF9A47E0F4E6E" box="[457,535,1608,1632]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="466" unit="m" value="3100.0">3100m</elevation>
</quantity>
<date id="FFFC102EF17CFFBAFDB8F9A47E9F4E6E" box="[542,647,1608,1632]" pageId="15" pageNumber="466" value="1983-10-07">
<collectingDate id="EFB8E9C6F17CFFBAFDB8F9A47E9F4E6E" box="[542,647,1608,1632]" pageId="15" pageNumber="466" value="1983-10-07">7.X.1983</collectingDate>
</date>
| [I.]Löbl—[A.]Smetana’ (
<collectionCode id="ED53AE2BF17CFFBAFC1EF9A4780C4E6E" box="[952,1044,1608,1632]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="466" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
. All 
<typeStatus id="54F9884CF17CFFBAFBF5F9A478A74E6E" box="[1107,1215,1608,1632]" pageId="15" pageNumber="466" type="paratype">paratypes</typeStatus>
with additional red rectangular printed label: ‘ 
<typeStatus id="54F9884CF17CFFBAFE1DF9807E534E8A" box="[443,587,1644,1668]" pageId="15" pageNumber="466" type="paratype">PARATYPE</typeStatus>
| 
<taxonomicName id="4C424D6DF17CFFBAFDF9F9807EC74E8B" authorityName="Leach" authorityYear="1819" box="[607,735,1644,1669]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="505" phylum="Arthropoda" rank="genus">Anthobium</taxonomicName>
| besucheti 
<emphasis id="B936EAFCF17CFFBAFCC5F9807F834E8B" bold="true" box="[867,923,1644,1669]" pageId="15" pageNumber="466">
<taxonomicNameLabel id="A2055787F17CFFBAFCC5F9807F834E8B" box="[867,923,1644,1669]" pageId="15" pageNumber="466" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. des. 2018’.
</paragraph>
</subSubSection>
<subSubSection id="C3586565F17CFFA5FF61F9637CCD4D5B" lastPageId="16" lastPageNumber="467" pageId="15" pageNumber="466" type="description">
<paragraph id="8BFD36EEF17CFFBAFF61F963791C4EFE" blockId="15.[151,1437,1356,2029]" pageId="15" pageNumber="466">
<emphasis id="B936EAFCF17CFFBAFF61F9637D494EA6" bold="true" box="[199,337,1679,1704]" pageId="15" pageNumber="466">Description</emphasis>
. Measurements (n=24): HW: 0.55–0.57, HL: 0.32–0.36; AL(
<typeStatus id="54F9884CF17CFFBAFBA3F96378744EA6" box="[1029,1132,1679,1704]" pageId="15" pageNumber="466" type="holotype">holotype</typeStatus>
): 0.93; OL: 0.12–0.14; PL: 0.41–0.48; PW: 0.81–0.95; ESL: 0.87–1.02; EW: 1.07–1.15; AW: 0.80–0.96; MTbL(
<typeStatus id="54F9884CF17CFFBAFB7AF958795A4EC3" box="[1244,1346,1716,1741]" pageId="15" pageNumber="466" type="holotype">holotype</typeStatus>
): 0.52; MTrL(
<typeStatus id="54F9884CF17CFFBAFF40F93B7D534EFE" box="[230,331,1751,1776]" pageId="15" pageNumber="466" type="holotype">holotype</typeStatus>
): 0.22 (MTrL 1–4: 0.14; MTrL 5: 0.08); AedL: 0.42; TL: 1.95–2.74 (
<typeStatus id="54F9884CF17CFFBAFBF6F93B78A04EFE" box="[1104,1208,1751,1776]" pageId="15" pageNumber="466" type="holotype">holotype</typeStatus>
: 2.40).
</paragraph>
<paragraph id="8BFD36EEF17CFFA5FF61F9107DF4494E" blockId="15.[151,1437,1356,2029]" lastBlockId="16.[151,1437,151,1688]" lastPageId="16" lastPageNumber="467" pageId="15" pageNumber="466">
Body elongate, convex. Body yellow to yellow-brown, some specimens with reddish-brown head, antennomeres 7–11 and abdomen; mouthparts, antennomeres 1–6 and legs yellow. Posterior portions of infraorbital ridges and area between ocelli with slightly rugose transverse sculpture between punctures, neck and abdominal tergites with distinct isodiametric microsculpture. Head with irregular, small and deep punctation, denser on posterior portion; middle part of neck with irregular and sparse punctation; pronotum with irregular punctation, markedly denser, larger and coarser than that on posterior part of head, sparser on lateral portions and posterior third of middle elevation, some specimens with small impunctate area between mediobasal impression and/or on basolateral portions; visible part of scutellum impunctated or with several small punctures; punctation of elytra dense, distinctly larger and deeper than that on pronotum, smaller and denser on parascutellar area, each elytron with five to eight very vague and tangled longitudinal rows of punctures, replaced by confluent punctures in some 
<typeStatus id="54F9884CF163FFA5FF31FEE97D1E4913" box="[151,262,261,285]" pageId="16" pageNumber="467" type="paratype">paratypes</typeStatus>
; abdominal tergites without visible punctures or with small and dense punctation on abdominal tergites VII–VIII. Habitus as in 
<figureCitation id="13792A6BF163FFA5FE05FECB7DFF494E" box="[419,487,295,320]" captionStart="FIGURES 1–5" captionStartId="5.[151,264,1628,1651]" captionTargetBox="[237,1323,200,1594]" captionTargetId="figure@5.[150,1436,195,1604]" captionTargetPageId="5" captionText="FIGURES 1–5. Habitus of Anthobium fusculum group: 1—A. fusculum (KAZAKHSTAN: Tian Shan), 2—A. altivagans (NEPAL: Telbrung Danda), 3—A. besucheti (paratype: Ghoropani Pass), 4—A. capitale (holotype), 5—A. cavicrus (NEPAL: Thare Pati). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607314/files/figure.png" pageId="16" pageNumber="467">Fig. 3</figureCitation>
.
</paragraph>
<paragraph id="8BFD36EEF163FFA5FF61FEA07E404AA6" blockId="16.[151,1437,151,1688]" pageId="16" pageNumber="467">
Head 1.5–1.7 times as wide as long; middle portion of head between eyes slightly elevated, usually with middle transverse impression between eyes at posterior third, with indistinct to distinct small and narrow grooves in front of ocelli, sometimes reaching middle length of eyes; postocular ridges usually smooth; anterior portion between antennal insertion and anterior margin of eye with wide and shallow semicircular notch. Ocelli small, situated at level of postocular ridges; distance between ocelli one and half times as long as distance between ocellus and posterior margin of eye. Apical segment of maxillary palp about three times as long as preceding segment, widest in basal part and conically narrowed toward acute apex. Antenna moderately short, reaching shoulders of elytra when reclined; apical antennomeres widened, as wide as basal antennomere; length × width of antennomeres (
<typeStatus id="54F9884CF163FFA5FF39FD807D104A8B" box="[159,264,620,645]" pageId="16" pageNumber="467" type="holotype">holotype</typeStatus>
): 1: 0.13 × 0.05; 2: 0.07 × 0.04; 3–4: 0.07 × 0.03; 5–6: 0.08 × 0.03; 7: 0.07 × 0.03; 8: 0.07 × 0.04; 9: 0.07 × 0.05; 10: 0.07 × 0.05; 11: 0.15 × 0.05.
</paragraph>
<paragraph id="8BFD36EEF163FFA5FF61FD5979034B8E" blockId="16.[151,1437,151,1688]" pageId="16" pageNumber="467">Pronotum about twice as wide as long, 1.4–1.6 times as wide as head, widest in front of middle, indistinctly and evenly narrowed both posteriad and anteriad; apical margin slightly shorter than posterior margin, widely rounded on anterior margin; anterior angles widely rounded, slightly protruded anteriad, posterior angles obtuse; lateral edges of pronotum with irregular, small and smooth crenulation; pronotum with very wide, markedly elevated middle portion, with very indistinct to distinct and deep longitudinal impression on anterior half of middle elevation, with pair of distinct, slightly or moderately deep transverse impressions on mediobasal third.</paragraph>
<paragraph id="8BFD36EEF163FFA5FF61FC607FC04BE3" blockId="16.[151,1437,151,1688]" pageId="16" pageNumber="467">Elytra slightly broader than long, about twice as long as pronotum, slightly widened apicad, reaching apical margin of abdominal tergite IV or V; each elytron in middle with irregular, distinct elevations between punctures, forming indistinct two or three longitudinal elevations in some specimens.</paragraph>
<paragraph id="8BFD36EEF163FFA5FF61FC1B7EF04C1E" blockId="16.[151,1437,151,1688]" box="[199,744,1015,1040]" pageId="16" pageNumber="467">Abdomen with indistinctly elevated paratergites.</paragraph>
<paragraph id="8BFD36EEF163FFA5FF61FBF078234CAE" blockId="16.[151,1437,151,1688]" pageId="16" pageNumber="467">
Male. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII slightly sinuate. Aedeagus (
<figureCitation id="13792A6BF163FFA5FE47FBD37E214C56" box="[481,569,1087,1112]" captionStart="FIGURES 50–53" captionStartId="14.[151,264,1972,1995]" captionTargetBox="[386,1272,196,1956]" captionTargetId="figure@14.[231,1360,193,1960]" captionTargetPageId="14" captionText="FIGURES 50–53. Aedeagus of Anthobium altivagans (50–51: NEPAL: Ganesh Himal) and A. besucheti (52–53: NEPAL: Ghoropani Pass): 50, 52—parameral view, 51, 53—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607328/files/figure.png" pageId="16" pageNumber="467">Fig. 52</figureCitation>
) with median lobe moderately wide basally, gradually narrowed toward subtruncate apex; parameres wide, significantly exceeding apex of median lobe, with two small apical and preapical setae; internal sac very narrow and long. Aedeagus laterally as in 
<figureCitation id="13792A6BF163FFA5FC42FB6B782F4CAE" box="[996,1079,1159,1184]" captionStart="FIGURES 50–53" captionStartId="14.[151,264,1972,1995]" captionTargetBox="[386,1272,196,1956]" captionTargetId="figure@14.[231,1360,193,1960]" captionTargetPageId="14" captionText="FIGURES 50–53. Aedeagus of Anthobium altivagans (50–51: NEPAL: Ganesh Himal) and A. besucheti (52–53: NEPAL: Ghoropani Pass): 50, 52—parameral view, 51, 53—lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607328/files/figure.png" pageId="16" pageNumber="467">Fig. 53</figureCitation>
.
</paragraph>
<paragraph id="8BFD36EEF163FFA5FF61FB4079914CCB" blockId="16.[151,1437,151,1688]" box="[199,1417,1196,1221]" pageId="16" pageNumber="467">Female. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded.</paragraph>
<paragraph id="8BFD36EEF163FFA5FF61FB237CCD4D5B" blockId="16.[151,1437,151,1688]" pageId="16" pageNumber="467">
<emphasis id="B936EAFCF163FFA5FF61FB237DAB4CE6" bold="true" box="[199,435,1231,1256]" pageId="16" pageNumber="467">Comparative notes.</emphasis>
Based on the body size, coloration, shape of the body with moderately short elytra, 
<taxonomicName id="4C424D6DF163FFA5FA22FB3C7DF34D02" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">
<emphasis id="B936EAFCF163FFA5FA22FB3C7D194D02" italics="true" pageId="16" pageNumber="467">A. besucheti</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
, 
<emphasis id="B936EAFCF163FFA5FE5FFB187E294D03" bold="true" box="[505,561,1268,1293]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="A2055787F163FFA5FE5FFB187E294D03" box="[505,561,1268,1293]" pageId="16" pageNumber="467" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is most similar to 
<taxonomicName id="4C424D6DF163FFA5FCADFB1878B14D02" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[779,1193,1268,1293]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
<emphasis id="B936EAFCF163FFA5FCADFB187FA64D02" box="[779,958,1268,1292]" italics="true" pageId="16" pageNumber="467">A. denticulatum</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
, 
<emphasis id="B936EAFCF163FFA5FB11FB1878F74D03" bold="true" box="[1207,1263,1268,1293]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="A2055787F163FFA5FB11FB1878F74D03" box="[1207,1263,1268,1293]" pageId="16" pageNumber="467" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, from which it differs by the slightly wider pronotum, by the narrower and smaller aedeagus with internal sac missing sclerotized teeth.
</paragraph>
</subSubSection>
<subSubSection id="C3586565F163FFA5FF61FAB37D7C4E5E" pageId="16" pageNumber="467" type="distribution">
<paragraph id="8BFD36EEF163FFA5FF61FAB3791F4D76" blockId="16.[151,1437,151,1688]" box="[199,1287,1375,1400]" pageId="16" pageNumber="467">
<emphasis id="B936EAFCF163FFA5FF61FAB37D784D76" bold="true" box="[199,352,1375,1400]" pageId="16" pageNumber="467">Distribution.</emphasis>
The species is known from several localities in central and eastern 
<collectingCountry id="F355767EF163FFA5FBF2FAB3788F4D76" box="[1108,1175,1375,1400]" name="Nepal" pageId="16" pageNumber="467">Nepal</collectingCountry>
(
<figureCitation id="13792A6BF163FFA5FB00FAB378E24D76" box="[1190,1274,1375,1400]" captionStart="FIGURE 54" captionStartId="15.[151,250,982,1004]" captionTargetBox="[179,1408,193,961]" captionTargetId="figure@15.[179,1408,193,961]" captionTargetPageId="15" captionText="FIGURE 54. Distribution of Anthobium altivagans (black circles), A. besucheti (black squares), A. rugosum (black triangles) and A. denticulatum (black star) in the Himalayan Region." httpUri="https://zenodo.org/record/2607330/files/figure.png" pageId="16" pageNumber="467">Fig. 54</figureCitation>
).
</paragraph>
<paragraph id="8BFD36EEF163FFA5FF61FA687D7C4E5E" blockId="16.[151,1437,151,1688]" pageId="16" pageNumber="467">
<emphasis id="B936EAFCF163FFA5FF61FA687D524D93" bold="true" box="[199,330,1412,1437]" pageId="16" pageNumber="467">Bionomics.</emphasis>
Specimens were collected at elevations from 
<quantity id="4CBA9B0BF163FFA5FCEDFA687FE34D92" box="[843,1019,1412,1437]" metricMagnitude="3" metricUnit="m" metricValue="3.35" metricValueMax="3.6" metricValueMin="3.1" pageId="16" pageNumber="467" unit="m" value="3350.0" valueMax="3600.0" valueMin="3100.0">3100 to 3600 m</quantity>
a.s.l. Specimens from Parbat District were taken by sifting of 
<taxonomicName id="4C424D6DF163FFA5FE01FA447DF24DB1" box="[423,490,1448,1471]" class="Magnoliopsida" family="Cyperaceae" genus="Carex" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="16" pageNumber="467" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFCF163FFA5FE01FA447DF24DB1" box="[423,490,1448,1471]" italics="true" pageId="16" pageNumber="467">Carex</emphasis>
</taxonomicName>
mixed with other vegetation along edges of the pond in an 
<taxonomicName id="4C424D6DF163FFA5FB24FA4478D84DCE" box="[1154,1216,1448,1472]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="16" pageNumber="467" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFCF163FFA5FB24FA4478D84DCE" box="[1154,1216,1448,1472]" italics="true" pageId="16" pageNumber="467">Abies</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C424D6DF163FFA5FB50FA4479834DCE" box="[1270,1435,1448,1472]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="16" pageNumber="467" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFCF163FFA5FB50FA4479834DCE" box="[1270,1435,1448,1472]" italics="true" pageId="16" pageNumber="467">Rhododendron</emphasis>
</taxonomicName>
forest; a male from Sundar Danda (
<collectingCountry id="F355767EF163FFA5FD8CFA207E684DEB" box="[554,624,1484,1509]" name="Nepal" pageId="16" pageNumber="467">Nepal</collectingCountry>
) was taken by sifting leaf litter in 
<emphasis id="B936EAFCF163FFA5FC5AFA2078F14DEA" box="[1020,1257,1484,1508]" italics="true" pageId="16" pageNumber="467">
<taxonomicName id="4C424D6DF163FFA5FC5AFA2078B94DEA" box="[1020,1185,1484,1508]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="16" pageNumber="467" phylum="Tracheophyta" rank="genus">Rhododendron</taxonomicName>
- 
<taxonomicName id="4C424D6DF163FFA5FB0CFA2078F14DEA" box="[1194,1257,1484,1508]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="16" pageNumber="467" phylum="Tracheophyta" rank="genus">Abies</taxonomicName>
</emphasis>
forest, together with 
<taxonomicName id="4C424D6DF163FFA5FF74FA1C7D7C4E06" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[210,356,1520,1544]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B936EAFCF163FFA5FF74FA1C7D7C4E06" box="[210,356,1520,1544]" italics="true" pageId="16" pageNumber="467">A. monticola</emphasis>
</taxonomicName>
(see below). Specimens from Goru Dzure Dara were collected together with 
<taxonomicName id="4C424D6DF163FFA5FB43FA1C797D4E09" baseAuthorityName="Champion" baseAuthorityYear="1925" box="[1253,1381,1520,1543]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="cavicrus">
<emphasis id="B936EAFCF163FFA5FB43FA1C797D4E09" box="[1253,1381,1520,1543]" italics="true" pageId="16" pageNumber="467">A. cavicrus</emphasis>
</taxonomicName>
(see below) and were taken by sifting of various debris, moss and 
<taxonomicName id="4C424D6DF163FFA5FCFFF9F87FE64E22" box="[857,1022,1556,1580]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" higherTaxonomySource="GBIF" kingdom="Plantae" order="Ericales" pageId="16" pageNumber="467" phylum="Tracheophyta" rank="genus">
<emphasis id="B936EAFCF163FFA5FCFFF9F87FE64E22" box="[857,1022,1556,1580]" italics="true" pageId="16" pageNumber="467">Rhododendron</emphasis>
</taxonomicName>
leaf litter in a small gorge with still remnants of snow.
</paragraph>
</subSubSection>
<subSubSection id="C3586565F163FFA5FF61F9B07D824E96" pageId="16" pageNumber="467" type="etymology">
<paragraph id="8BFD36EEF163FFA5FF61F9B07D824E96" blockId="16.[151,1437,151,1688]" pageId="16" pageNumber="467">
<emphasis id="B936EAFCF163FFA5FF61F9B07D564E7B" bold="true" box="[199,334,1628,1653]" pageId="16" pageNumber="467">Etymology.</emphasis>
Patronymic, the species is named to honour our colleague, coleopterologist Claude Besuchet (
<collectingRegion id="4986F80CF163FFA5FF39F96C7CE14E96" box="[159,249,1664,1688]" country="Switzerland" name="Geneve" pageId="16" pageNumber="467">Genève</collectingRegion>
, 
<collectingCountry id="F355767EF163FFA5FEA2F9937D974E96" box="[260,399,1663,1688]" name="Switzerland" pageId="16" pageNumber="467">Switzerland</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>