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<document id="A44E5FC29ECE31A5E270DEA65D782DF8" ID-DOI="10.1080/00222933.2018.1481235" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747141" ID-ZooBank="63A31AD2-F049-42CB-A45B-557014DC286E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670765052" checkinUser="carolina" docAuthor="Taylor, Paul D., Martha, Silviu O. &amp; Gordon, Dennis P." docDate="2018" docId="03EB8789FFE1447CA19571B7FE13FDBC" docLanguage="en" docName="JNATHIST.52.25-26.1657-1721.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222933.2018.1481235" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Aechmella Canu and Bassler 1917" docType="treatment" docVersion="4" lastPageNumber="1667" masterDocId="FFD2FFF1FFE84477A00C7447FF92FFBE" masterDocTitle="Synopsis of onychocellid cheilostome bryozoan genera" masterLastPageNumber="1721" masterPageNumber="1657" pageNumber="1665" updateTime="1698914560039" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3B96A3CFFF8015BB0F362F41B645DFAF">Synopsis of onychocellid cheilostome bryozoan genera</mods:title>
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<mods:namePart id="C93C7C2EF374EFBA975BE099A360F826">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="21A84B739437E49CFF5472A3E1E09AAE">Martha, Silviu O.</mods:namePart>
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<mods:title id="501AD2E57F61428DD06B54909EAF81FB">Journal of Natural History</mods:title>
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<mods:date id="79424322574FF9F3EC95A1BEDABEC768">2018</mods:date>
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<mods:number id="9EFF6D96E9EE37BD30E9B22764D8EF11">2018-07-03</mods:number>
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<paragraph id="8BFD369FFFE1447EA19571B7FC1BF9B7" blockId="9.[409,905,1520,1579]" box="[409,905,1520,1545]" pageId="9" pageNumber="1665">
<heading id="D0B581F3FFE1447EA19571B7FC1BF9B7" box="[409,905,1520,1545]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="1665" reason="2">
Genus
<taxonomicName id="4C424D1CFFE1447EA1E171B7FC1BF9B7" ID-CoL="858HJ" authority="Canu and Bassler, 1917" authorityName="Canu and Bassler" authorityYear="1917" box="[493,905,1520,1545]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="9" pageNumber="1665" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE1447EA1E171B7FDFEF9B7" bold="true" box="[493,620,1520,1545]" italics="true" pageId="9" pageNumber="1665">Aechmella</emphasis>
<bibRefCitation id="EFD34B6EFFE1447EA27971B7FC1BF9B7" author="Canu F &amp; Bassler RS" box="[629,905,1520,1545]" pageId="9" pageNumber="1665" pagination="1 - 87" refId="ref24368" refString="Canu F, Bassler RS. 1917. A synopsis of American Early Tertiary cheilostome Bryozoa. Bull U S Natl Mus. 96: 1 - 87." type="journal article" year="1917">Canu and Bassler, 1917</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3586514FFE1447EA2557255FD59F995" box="[601,715,1554,1579]" pageId="9" pageNumber="1665" type="description">
<paragraph id="8BFD369FFFE1447EA2557255FD59F995" blockId="9.[409,905,1520,1579]" box="[601,715,1554,1579]" pageId="9" pageNumber="1665">
(
<figureCitation id="13792A1AFFE1447EA26D7255FD50F995" box="[609,706,1554,1579]" captionStart="Figure 2" captionStartId="10.[160,225,932,955]" captionTargetBox="[163,1153,142,901]" captionTargetId="figure-209@10.[163,1153,142,901]" captionTargetPageId="10" captionText="Figure 2. Aechmella filimargo Canu and Bassler; Eocene, late Jacksonian (= Priabonian), Sepulga River west bank, Escambia County, Alabama, USA; syntypes USNM 62585. (a,b) Syntype 2 showing autozooids and four avicularia marked by arrows. (c) Early astogeny in syntype 2 with ancestrula arrowed. (d) Syntype 1 with broken ovicell arrowed. Scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747145" httpUri="https://zenodo.org/record/4747145/files/figure.png" pageId="9" pageNumber="1665">Figure 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFE1447EA0AC7226FCB4F901" pageId="9" pageNumber="1665" type="type_taxon">
<paragraph id="8BFD369FFFE1447EA0AC7226FEA6F9C4" blockId="9.[160,1156,1633,1727]" box="[160,308,1633,1658]" pageId="9" pageNumber="1665">
<emphasis id="B936EA8DFFE1447EA0AC7226FEA6F9C4" bold="true" box="[160,308,1633,1658]" italics="true" pageId="9" pageNumber="1665">
<typeStatus id="54F9883DFFE1447EA0AC7226FF4AF9C4" box="[160,216,1633,1658]" pageId="9" pageNumber="1665">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="8BFD369FFFE1447EA0AC72C2FCB4F901" blockId="9.[160,1156,1633,1727]" pageId="9" pageNumber="1665">
<taxonomicName id="4C424D1CFFE1447EA0AC72C2FE2EF923" ID-CoL="86D2C" box="[160,444,1668,1693]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="9" pageNumber="1665" phylum="Bryozoa" rank="species" species="filimargo">
<emphasis id="B936EA8DFFE1447EA0AC72C2FEE4F923" box="[160,374,1668,1693]" italics="true" pageId="9" pageNumber="1665">Aechmella filimargo</emphasis>
, 1917
</taxonomicName>
, by original designation.
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Eocene, late Jacksonian (= Priabonian),
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west bank,
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,
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,
<collectingCountry id="F355760FFFE1447EA2FD72E0FCB0F901" box="[753,802,1703,1727]" name="United States of America" pageId="9" pageNumber="1665">USA</collectingCountry>
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.
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<caption id="DF3D6617FFE2447DA0AC77E3FC1EFBAD" ID-DOI="http://doi.org/10.5281/zenodo.4747145" ID-Zenodo-Dep="4747145" httpUri="https://zenodo.org/record/4747145/files/figure.png" pageId="10" pageNumber="1666" startId="10.[160,225,932,955]" targetBox="[163,1153,142,901]" targetPageId="10">
<paragraph id="8BFD369FFFE2447DA0AC77E3FC1EFBAD" blockId="10.[160,1156,932,1043]" pageId="10" pageNumber="1666">
<emphasis id="B936EA8DFFE2447DA0AC77E3FF6CFC05" bold="true" box="[160,254,932,955]" pageId="10" pageNumber="1666">Figure 2.</emphasis>
<taxonomicName id="4C424D1CFFE2447DA10B77E3FD1BFC05" authority="Canu and Bassler" authorityName="Canu and Bassler" box="[263,649,932,955]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1666" phylum="Bryozoa" rank="species" species="filimargo">
<emphasis id="B936EA8DFFE2447DA10B77E3FE59FC05" box="[263,459,932,955]" italics="true" pageId="10" pageNumber="1666">Aechmella filimargo</emphasis>
Canu and Bassler
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; Eocene, late Jacksonian (= Priabonian), Sepulga River west bank, Escambia County, Alabama, USA; syntypes USNM 62585. (a,b) Syntype 2 showing autozooids and four avicularia marked by arrows. (c) Early astogeny in syntype 2 with ancestrula arrowed. (d) Syntype 1 with broken ovicell arrowed. Scale bars = 200 µm.
</paragraph>
</caption>
<subSubSection id="C3586514FFE2447DA0AC7009FD2EFAC2" pageId="10" pageNumber="1666" type="diagnosis">
<paragraph id="8BFD369FFFE2447DA0AC7009FE86FBD9" blockId="10.[160,1156,1102,1404]" box="[160,276,1102,1127]" pageId="10" pageNumber="1666">
<emphasis id="B936EA8DFFE2447DA0AC7009FE86FBD9" bold="true" box="[160,276,1102,1127]" italics="true" pageId="10" pageNumber="1666">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BFD369FFFE2447DA0AC7036FD2EFAC2" blockId="10.[160,1156,1102,1404]" pageId="10" pageNumber="1666">
Colony encrusting. Autozooids subhexagonal (
<figureCitation id="13792A1AFFE2447DA2B57036FC8AFB37" box="[697,792,1137,1161]" captionStart="Figure 2" captionStartId="10.[160,225,932,955]" captionTargetBox="[163,1153,142,901]" captionTargetId="figure-209@10.[163,1153,142,901]" captionTargetPageId="10" captionText="Figure 2. Aechmella filimargo Canu and Bassler; Eocene, late Jacksonian (= Priabonian), Sepulga River west bank, Escambia County, Alabama, USA; syntypes USNM 62585. (a,b) Syntype 2 showing autozooids and four avicularia marked by arrows. (c) Early astogeny in syntype 2 with ancestrula arrowed. (d) Syntype 1 with broken ovicell arrowed. Scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747145" httpUri="https://zenodo.org/record/4747145/files/figure.png" pageId="10" pageNumber="1666">Figure 2</figureCitation>
(ab)), rounded distally; zooidal boundaries raised with a medial groove. Cryptocyst extensive, seemingly granular. Gymnocyst lacking. Opesia terminal, occupying about one-third of frontal surface; bell shaped with lateral constrictions at approximately mid-length, lacking opesiular indentations, proximal edge concave. Ovicell hyperstomial, ooecium?cryptocyst-like (
<figureCitation id="13792A1AFFE2447DA42B70BBFB11FAAA" box="[1063,1155,1276,1300]" captionStart="Figure 2" captionStartId="10.[160,225,932,955]" captionTargetBox="[163,1153,142,901]" captionTargetId="figure-209@10.[163,1153,142,901]" captionTargetPageId="10" captionText="Figure 2. Aechmella filimargo Canu and Bassler; Eocene, late Jacksonian (= Priabonian), Sepulga River west bank, Escambia County, Alabama, USA; syntypes USNM 62585. (a,b) Syntype 2 showing autozooids and four avicularia marked by arrows. (c) Early astogeny in syntype 2 with ancestrula arrowed. (d) Syntype 1 with broken ovicell arrowed. Scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747145" httpUri="https://zenodo.org/record/4747145/files/figure.png" pageId="10" pageNumber="1666">Figure 2</figureCitation>
(d)). Avicularia interzooidal, symmetrical, about half the width of an autozooid; opesia pear shaped; rostrum acutely triangular (
<figureCitation id="13792A1AFFE2447DA27B7106FD47FAE7" box="[631,725,1345,1369]" captionStart="Figure 2" captionStartId="10.[160,225,932,955]" captionTargetBox="[163,1153,142,901]" captionTargetId="figure-209@10.[163,1153,142,901]" captionTargetPageId="10" captionText="Figure 2. Aechmella filimargo Canu and Bassler; Eocene, late Jacksonian (= Priabonian), Sepulga River west bank, Escambia County, Alabama, USA; syntypes USNM 62585. (a,b) Syntype 2 showing autozooids and four avicularia marked by arrows. (c) Early astogeny in syntype 2 with ancestrula arrowed. (d) Syntype 1 with broken ovicell arrowed. Scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747145" httpUri="https://zenodo.org/record/4747145/files/figure.png" pageId="10" pageNumber="1666">Figure 2</figureCitation>
(a)). Ancestrula similar in morphology to budded autozooids but smaller (
<figureCitation id="13792A1AFFE2447DA23A7123FD06FAC2" box="[566,660,1380,1404]" captionStart="Figure 2" captionStartId="10.[160,225,932,955]" captionTargetBox="[163,1153,142,901]" captionTargetId="figure-209@10.[163,1153,142,901]" captionTargetPageId="10" captionText="Figure 2. Aechmella filimargo Canu and Bassler; Eocene, late Jacksonian (= Priabonian), Sepulga River west bank, Escambia County, Alabama, USA; syntypes USNM 62585. (a,b) Syntype 2 showing autozooids and four avicularia marked by arrows. (c) Early astogeny in syntype 2 with ancestrula arrowed. (d) Syntype 1 with broken ovicell arrowed. Scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747145" httpUri="https://zenodo.org/record/4747145/files/figure.png" pageId="10" pageNumber="1666">Figure 2</figureCitation>
(c)).
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFE2447CA0AC71F3FE13FDBC" lastPageId="11" lastPageNumber="1667" pageId="10" pageNumber="1666" type="discussion">
<paragraph id="8BFD369FFFE2447DA0AC71F3FE94FA73" blockId="10.[160,1156,1460,1727]" box="[160,262,1460,1485]" pageId="10" pageNumber="1666">
<emphasis id="B936EA8DFFE2447DA0AC71F3FE94FA73" bold="true" box="[160,262,1460,1485]" italics="true" pageId="10" pageNumber="1666">Remarks</emphasis>
</paragraph>
<paragraph id="8BFD369FFFE2447CA0AC7190FC3CFE27" blockId="10.[160,1156,1460,1727]" lastBlockId="11.[160,1155,142,409]" lastPageId="11" lastPageNumber="1667" pageId="10" pageNumber="1666">
The
<typeStatus id="54F9883DFFE2447DA0C17190FF6FFA51" box="[205,253,1495,1519]" pageId="10" pageNumber="1666">type</typeStatus>
species of
<emphasis id="B936EA8DFFE2447DA17C7190FDCEFA51" box="[368,604,1495,1519]" italics="true" pageId="10" pageNumber="1666">
<taxonomicName id="4C424D1CFFE2447DA17C7190FE4CFA51" authorityName="Canu and Bassler" authorityYear="1917" box="[368,478,1495,1519]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1666" phylum="Bryozoa" rank="genus">Aechmella</taxonomicName>
,
<taxonomicName id="4C424D1CFFE2447DA1EA7190FDCEFA51" box="[486,604,1495,1519]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1666" phylum="Bryozoa" rank="species" species="filimargo">A. filimargo</taxonomicName>
</emphasis>
, was described as rare by
<bibRefCitation id="EFD34B6EFFE2447DA3637190FBF7FA51" author="Canu F &amp; Bassler RS" box="[879,1125,1495,1519]" pageId="10" pageNumber="1666" pagination="1 - 87" refId="ref24368" refString="Canu F, Bassler RS. 1917. A synopsis of American Early Tertiary cheilostome Bryozoa. Bull U S Natl Mus. 96: 1 - 87." type="journal article" year="1917">Canu and Bassler (1917</bibRefCitation>
, p. 30), who gave the range of the genus as Cenomanian to Miocene. Scanning electron micrographs of the
<typeStatus id="54F9883DFFE2447DA188725AFE26F98B" box="[388,436,1565,1589]" pageId="10" pageNumber="1666">type</typeStatus>
material of this species were kindly prepared for us by JoAnn Sanner of the USNM. The genus was named for the lance-like shape of the avicularia which, in contrast to the
<typeStatus id="54F9883DFFE2447DA16F7225FE01F9C4" box="[355,403,1634,1658]" pageId="10" pageNumber="1666">type</typeStatus>
and most other species of
<taxonomicName id="4C424D1CFFE2447DA2B77225FCA8F9C4" authorityName="Jullien" authorityYear="1882" box="[699,826,1634,1658]" class="Gymnolaemata" family="Onychocellidae" genus="Onychocella" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1666" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE2447DA2B77225FCA8F9C4" box="[699,826,1634,1658]" italics="true" pageId="10" pageNumber="1666">Onychocella</emphasis>
</taxonomicName>
, is symmetrical. Subsequently,
<bibRefCitation id="EFD34B6EFFE2447DA0AC72C2FE0BF923" author="Canu F &amp; Bassler RS" box="[160,409,1669,1693]" pageId="10" pageNumber="1666" pagination="1 - 879" refId="ref24397" refString="Canu F, Bassler RS. 1920. North American Early Tertiary Bryozoa. Bull U S Natl Mus. 106: 1 - 879." type="journal article" year="1920">Canu and Bassler (1920</bibRefCitation>
, p. 233) redescribed
<taxonomicName id="4C424D1CFFE2447DA27472C2FD7DF923" box="[632,751,1668,1693]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1666" phylum="Bryozoa" rank="species" species="filimargo">
<emphasis id="B936EA8DFFE2447DA27472C2FD7DF923" box="[632,751,1668,1693]" italics="true" pageId="10" pageNumber="1666">A. filimargo</emphasis>
</taxonomicName>
, introduced another species from the Eocene
<taxonomicName id="4C424D1CFFE2447DA10472E0FD0CF901" authority="Canu and Bassler, 1920" authorityName="Canu and Bassler" authorityYear="1920" box="[264,670,1703,1727]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="1666" phylum="Bryozoa" rank="species" species="crassimargo">
<emphasis id="B936EA8DFFE2447DA10472E0FE32F901" box="[264,416,1703,1727]" italics="true" pageId="10" pageNumber="1666">A. crassimargo</emphasis>
<bibRefCitation id="EFD34B6EFFE2447DA1AB72E0FD0CF901" author="Canu F &amp; Bassler RS" box="[423,670,1703,1727]" pageId="10" pageNumber="1666" pagination="1 - 879" refId="ref24397" refString="Canu F, Bassler RS. 1920. North American Early Tertiary Bryozoa. Bull U S Natl Mus. 106: 1 - 879." type="journal article" year="1920">Canu and Bassler, 1920</bibRefCitation>
</taxonomicName>
and referred a further nine existing species to
<taxonomicName id="4C424D1CFFE3447CA0CC74C9FEBEFF18" authorityName="Canu and Bassler" authorityYear="1917" box="[192,300,142,166]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" pageNumber="1667" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE3447CA0CC74C9FEBEFF18" box="[192,300,142,166]" italics="true" pageId="11" pageNumber="1667">Aechmella</emphasis>
</taxonomicName>
, the majority of which were originally described from the Late Cretaceous of Europe. Indeed, many of the species subsequently assigned to
<taxonomicName id="4C424D1CFFE3447CA33074F6FC3AFF77" authorityName="Canu and Bassler" authorityYear="1917" box="[828,936,177,201]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" pageNumber="1667" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE3447CA33074F6FC3AFF77" box="[828,936,177,201]" italics="true" pageId="11" pageNumber="1667">Aechmella</emphasis>
</taxonomicName>
are of Cretaceous or Paleocene age. There has been a tendency to use the genus for encrusting onychocellids with relatively small opesiae, wider than long, often almost crescent-shaped and sometimes with opesiular indentations but no lateral constrictions, and possessing small, symmetrical avicularia (e.g.
<bibRefCitation id="EFD34B6EFFE3447CA10A757BFEE9FEEA" author="Voigt E" box="[262,379,316,340]" pageId="11" pageNumber="1667" pagination="1 - 80" refId="ref28800" refString="Voigt E. 1959. Revision der von F. v. Hagenow 1838 - 1850 aus der Schreibkreide von Rugen veroffentlichten Bryozoen. Geol. 8: 1 - 80." type="journal article" year="1959">Voigt 1959</bibRefCitation>
;
<bibRefCitation id="EFD34B6EFFE3447CA18A757BFDA1FEEA" author="Berthelsen O" box="[390,563,316,340]" pageId="11" pageNumber="1667" refId="ref23703" refString="Berthelsen O. 1962. Cheilostome Bryozoa in the Danian deposits of East Denmark. KObenhavn: C. A. Reitzel." type="book" year="1962">Berthelsen 1962</bibRefCitation>
;
<bibRefCitation id="EFD34B6EFFE3447CA232757BFCC9FEEA" author="Taylor PD &amp; McKinney FK" box="[574,859,316,340]" pageId="11" pageNumber="1667" pagination="1 - 346" refId="ref28471" refString="Taylor PD, McKinney FK. 2006. Cretaceous Bryozoa from the Campanian and Maastrichtian of the Atlantic and Gulf Coastal Plains, United States. Scripta Geol. 132: 1 - 346." type="journal article" year="2006">Taylor and McKinney 2006</bibRefCitation>
;
<bibRefCitation id="EFD34B6EFFE3447CA36A757BFC71FEEA" author="Taylor PD" box="[870,995,316,340]" pageId="11" pageNumber="1667" pagination="823 - 834" refId="ref28263" refString="Taylor PD. 2008. Late Cretaceous cheilostome bryozoans from California and Baja California. J Paleontol. 82: 823 - 834." type="journal article" year="2008">Taylor 2008</bibRefCitation>
). Based on the morphology of the
<typeStatus id="54F9883DFFE3447CA1797519FE37FEC8" box="[373,421,350,374]" pageId="11" pageNumber="1667">type</typeStatus>
species, the generic identities of these species must be revised and we introduce the new genus
<taxonomicName id="4C424D1CFFE3447CA1D375C6FDCCFE27" authorityName="Taylor &amp; Martha &amp; Gordon" authorityYear="2018" box="[479,606,385,409]" class="Gymnolaemata" family="Onychocellidae" genus="Aechmellina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="11" pageNumber="1667" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFE3447CA1D375C6FDCCFE27" box="[479,606,385,409]" italics="true" pageId="11" pageNumber="1667">Aechmellina</emphasis>
</taxonomicName>
below to accommodate them.
</paragraph>
<paragraph id="8BFD369FFFE3447CA0AC7581FF79FE61" blockId="11.[160,385,454,514]" box="[160,235,454,479]" pageId="11" pageNumber="1667">
<emphasis id="B936EA8DFFE3447CA0AC7581FF79FE61" bold="true" box="[160,235,454,479]" italics="true" pageId="11" pageNumber="1667">Range</emphasis>
</paragraph>
<paragraph id="8BFD369FFFE3447CA0AC75ADFE13FDBC" blockId="11.[160,385,454,514]" box="[160,385,490,514]" pageId="11" pageNumber="1667">Eocene to Miocene.</paragraph>
</subSubSection>
</treatment>
</document>