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<document id="D015B375EED907AB95D5A1DEBF28A0AF" ID-CLB-Dataset="2406" ID-DOI="10.11646/zootaxa.5093.2.2" ID-GBIF-Dataset="ffbad455-ad57-4fa5-8efa-d18cc2d12e40" ID-ISSN="1175-5326" ID-Zenodo-Dep="5905141" ID-ZooBank="31913C5B-A8D5-4A30-8D9E-4C7C9E953E1E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643182084586" checkinUser="plazi" docAuthor="Antić, Dragan &amp; Mauriès, Jean-Paul" docDate="2022" docId="03E387B4FFC0CD0056C40AC6FF61FF66" docLanguage="en" docName="zootaxa.5093.2.2.pdf" docOrigin="Zootaxa 5093 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guipuzcosoma karinae Antić &amp; Mauriès 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="154" masterDocId="FFDAFFCCFFC7CD0C56530A51FFA7FFC8" masterDocTitle="Two new species of the previously monospecific genus Guipuzcosoma Vicente &amp; Mauriès, 1980 from Spain, with establishment of Guipuzcosomatidae fam. nov. (Diplopoda: Chordeumatida)" masterLastPageNumber="168" masterPageNumber="142" pageNumber="149" updateTime="1699304320109" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="05A55D70C5B3595507D82A40A4417D54">
<mods:title id="075B397A6874FA92794AA759E4311E77">Two new species of the previously monospecific genus Guipuzcosoma Vicente &amp; Mauriès, 1980 from Spain, with establishment of Guipuzcosomatidae fam. nov. (Diplopoda: Chordeumatida)</mods:title>
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<mods:namePart id="CC9B1530C7708619AF2C99F92832ED3A">Antić, Dragan</mods:namePart>
<mods:affiliation id="C8CE11BFCB81C74492FFA6E73E6AC983">University of Belgrade-Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia &amp; Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia</mods:affiliation>
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<mods:namePart id="ECD7E44D87A20EF4DF9A0DFF017452C3">Mauriès, Jean-Paul</mods:namePart>
<mods:nameIdentifier id="4C2998836D3C3D2E02A3290999681D5A" type="ORCID">0000-0002-0575-6937</mods:nameIdentifier>
<mods:affiliation id="5556148DDD27B0F9E7727A6D6AEDE95B">Muséum National d’Histoire Naturelle, Dpt. Origines &amp; Evolution, Section Arthropodes, 61 rue Buffon F- 75005 Paris, France mauries. jean-paul @ wanadoo. fr; https: // orcid. org / 0000 - 0002 - 0575 - 6937</mods:affiliation>
<mods:nameIdentifier id="085E49913CCDF2DA597C7E928F137932" type="email">mauries.jean-paul@wanadoo.fr</mods:nameIdentifier>
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<mods:title id="EA039E2DB005D329D716BFB0B1F11B34">Zootaxa</mods:title>
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<mods:part id="7C377B5EF56E8C5703F3A15D77C79F2F">
<mods:date id="835B90CBA92B7CBCB04CEE5EE7E0FE2B">2022</mods:date>
<mods:detail id="630DC267094DAC00440D41038B49F1F0" type="pubDate">
<mods:number id="EAFBCE8D67806B71EE9C27459EA85305">2022-01-26</mods:number>
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<mods:detail id="B3D26C50F28A3F508F8B8F75B82DEB92" type="volume">
<mods:number id="51BA93267F2D794DF8F312EE953E852B">5093</mods:number>
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<mods:number id="68D559989CCB25AFBCFBB9BBF559CB5B">2</mods:number>
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<mods:identifier id="404C3E478A4EEAEFCF55DF4952F0CF02" type="CLB-Dataset">2406</mods:identifier>
<mods:identifier id="2A198706832C1B306C89E8342ACF6913" type="DOI">10.11646/zootaxa.5093.2.2</mods:identifier>
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<treatment id="03E387B4FFC0CD0056C40AC6FF61FF66" ID-DOI="http://doi.org/10.5281/zenodo.5906928" ID-GBIF-Taxon="192626927" ID-Zenodo-Dep="5906928" LSID="urn:lsid:plazi:treatment:03E387B4FFC0CD0056C40AC6FF61FF66" httpUri="http://treatment.plazi.org/id/03E387B4FFC0CD0056C40AC6FF61FF66" lastPageId="12" lastPageNumber="154" pageId="7" pageNumber="149">
<subSubSection id="C3506529FFC0CD0B56C40AC6FDB3FF79" box="[151,532,151,178]" pageId="7" pageNumber="149" type="nomenclature">
<paragraph id="8BF536A2FFC0CD0B56C40AC6FDB3FF79" blockId="7.[151,532,151,213]" box="[151,532,151,178]" pageId="7" pageNumber="149">
<heading id="D0BD81CEFFC0CD0B56C40AC6FDB3FF79" bold="true" box="[151,532,151,178]" fontSize="11" level="1" pageId="7" pageNumber="149" reason="1">
<emphasis id="B93EEAB0FFC0CD0B56C40AC6FDB3FF79" bold="true" box="[151,532,151,178]" pageId="7" pageNumber="149">
<taxonomicName id="4C4A4D21FFC0CD0B56C40AC6FE08FF7A" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[151,431,151,178]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFC0CD0B56C40AC6FE08FF7A" bold="true" box="[151,431,151,178]" italics="true" pageId="7" pageNumber="149">Guipuzcosoma karinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57CBFFC0CD0B57E50AC6FDB3FF79" box="[438,532,151,177]" pageId="7" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFC0CD0B56C40AEAFE85FF1D" box="[151,290,187,213]" pageId="7" pageNumber="149" type="description">
<paragraph id="8BF536A2FFC0CD0B56C40AEAFE85FF1D" blockId="7.[151,532,151,213]" box="[151,290,187,213]" pageId="7" pageNumber="149">
<figureCitation id="13712A27FFC0CD0B56C40AEAFF5CFF1D" box="[151,251,187,213]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="9.[151,250,1943,1967]" captionStartId-1="10.[151,250,1673,1697]" captionStartId-2="11.[151,250,1847,1871]" captionStartId-3="12.[151,250,1761,1785]" captionStartId-4="13.[151,250,1757,1781]" captionTargetBox-0="[185,1403,181,1920]" captionTargetBox-1="[151,1436,181,1649]" captionTargetBox-2="[151,1435,181,1823]" captionTargetBox-3="[201,1387,199,1738]" captionTargetBox-4="[235,1358,194,1727]" captionTargetId-0="figure-19@9.[185,1403,181,1920]" captionTargetId-1="figure-17@10.[151,1436,181,1649]" captionTargetId-2="figure-19@11.[151,1435,181,1823]" captionTargetId-3="figure-18@12.[201,1387,199,1738]" captionTargetId-4="figure-19@13.[225,1363,181,1733]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 2. Guipuzcosoma karinae sp. nov., habitus (ZMBN). A. ♂ paratype, lateral view. B. ♂ paratype, anterior part of body, showing head and collum in dorsal view. C. ♂ paratype, anterior part of body, lateral view. D. ♀ paratype, midbody rings, dorsal view. E. ♂ paratype, lateral view. Scale bars: 1 mm." captionText-1="FIGURE 3. Guipuzcosoma karinae sp. nov., ♂ paratype, habitus (ZMBN). A. Head, anterior view. B. Head and collum, lateral view. C. Left antenna, anterior view. D. Field with ommatidia, lateral view. E, F. Body rings 15–17, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A–C, E, F), 0.02 mm (D)." captionText-2="FIGURE 4. Guipuzcosoma karinae sp. nov., legs and gonopods. A. ♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB). B–F. ♂ paratype (ZMBN). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Leg-pair 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2" captionText-3="FIGURE 5. Guipuzcosoma karinae sp. nov. A–H. ♂ paratype (ZMBN), I. ♀ paratype (NHMW MY10253). A–F. Anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G, H. Posterior gonopods, anterior and posterior views, respectively. Highlighed square in G shows a potential sperm. I. Vulvae, distal view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm." captionText-4="FIGURE 6. Guipuzcosoma karinae sp. nov. A–C. ♂ paratype (ZMBN), D–F. ♂ holotype (SMNG), G. ♀ paratype (NHMW MY10253). A–D.Anterior gonopods, anterior, lateral, posterior and postero-distal views, respectively. E, F. Posterior gonopods, anterior and lateral views, respectively. G. Vulvae, dital view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hairlike outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5905145" figureDoi-1="http://doi.org/10.5281/zenodo.5905147" figureDoi-2="http://doi.org/10.5281/zenodo.5905149" figureDoi-3="http://doi.org/10.5281/zenodo.5905151" figureDoi-4="http://doi.org/10.5281/zenodo.5905153" httpUri-0="https://zenodo.org/record/5905145/files/figure.png" httpUri-1="https://zenodo.org/record/5905147/files/figure.png" httpUri-2="https://zenodo.org/record/5905149/files/figure.png" httpUri-3="https://zenodo.org/record/5905151/files/figure.png" httpUri-4="https://zenodo.org/record/5905153/files/figure.png" pageId="7" pageNumber="149">Figs 2–6</figureCitation>
,
<figureCitation id="13712A27FFC0CD0B57550AEAFE85FF1D" box="[262,290,187,213]" captionStart="FIGURE 12" captionStartId="20.[151,250,1979,2003]" captionTargetBox="[151,1436,1101,1956]" captionTargetId="figure-470@20.[151,1436,1101,1956]" captionTargetPageId="20" captionText="FIGURE 12. Distribution of the genus Guipuzcosoma Vicente &amp; Mauriès (1980)." figureDoi="http://doi.org/10.5281/zenodo.5905165" httpUri="https://zenodo.org/record/5905165/files/figure.png" pageId="7" pageNumber="149">12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFC0CD0B56C40B52FC96FDAA" pageId="7" pageNumber="149" type="diagnosis">
<paragraph id="8BF536A2FFC0CD0B56C40B52FDD6FDD1" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="149">
<emphasis id="B93EEAB0FFC0CD0B56C40B52FEADFED5" bold="true" box="[151,266,259,285]" pageId="7" pageNumber="149">Diagnosis</emphasis>
. Differs from both congeners in the structure of the angiocoxites 1, which are almost the same width in their entire length, with an acuminate distal part bent laterad
<emphasis id="B93EEAB0FFC0CD0B55140B79FCC2FE89" box="[839,869,296,321]" italics="true" pageId="7" pageNumber="149">vs.</emphasis>
distal processes reduced, with lateral bumps in
<taxonomicName id="4C4A4D21FFC0CD0B53D20B79FF41FEAD" authorityName="Vicente &amp; Mauries" authorityYear="1980" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="comasi">
<emphasis id="B93EEAB0FFC0CD0B53D20B79FF41FEAD" italics="true" pageId="7" pageNumber="149">G. comasi</emphasis>
</taxonomicName>
, or long and sigmoid in
<taxonomicName id="4C4A4D21FFC0CD0B57AE0B1DFDF3FEAD" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[509,596,332,358]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFC0CD0B57AE0B1DFDF3FEAD" box="[509,596,332,358]" italics="true" pageId="7" pageNumber="149">G. reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC0CD0B54080B1AFD11FEAD" bold="true" box="[603,694,331,357]" pageId="7" pageNumber="149">
<taxonomicNameLabel id="A20D57CBFFC0CD0B54080B1AFD11FEAD" box="[603,694,331,357]" pageId="7" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The anterior part of the mesal bursal valve in
<taxonomicName id="4C4A4D21FFC0CD0B52910B1DFA9CFEAD" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[1218,1339,331,358]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFC0CD0B52910B1DFA9CFEAD" box="[1218,1339,331,358]" italics="true" pageId="7" pageNumber="149">G. karinae</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC0CD0B53110B1AFA3BFEAD" bold="true" box="[1346,1436,331,357]" pageId="7" pageNumber="149">
<taxonomicNameLabel id="A20D57CBFFC0CD0B53110B1AFA3BFEAD" box="[1346,1436,331,357]" pageId="7" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is subquadrangular with convex lateral margins,
<emphasis id="B93EEAB0FFC0CD0B54EB0B21FD70FE41" box="[696,727,368,393]" italics="true" pageId="7" pageNumber="149">vs.</emphasis>
heart-shaped in
<taxonomicName id="4C4A4D21FFC0CD0B55C20B21FBA5FE41" authorityName="Vicente &amp; Mauries" authorityYear="1980" box="[913,1026,368,394]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="comasi">
<emphasis id="B93EEAB0FFC0CD0B55C20B21FBA5FE41" box="[913,1026,368,394]" italics="true" pageId="7" pageNumber="149">G. comasi</emphasis>
</taxonomicName>
or quadrangular with straight lateral margins in
<taxonomicName id="4C4A4D21FFC0CD0B57450BC5FECAFE65" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[278,365,404,430]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFC0CD0B57450BC5FECAFE65" box="[278,365,404,430]" italics="true" pageId="7" pageNumber="149">G. reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC0CD0B57260BC2FE68FE65" bold="true" box="[373,463,403,429]" pageId="7" pageNumber="149">
<taxonomicNameLabel id="A20D57CBFFC0CD0B57260BC2FE68FE65" box="[373,463,403,429]" pageId="7" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Additionally,
<taxonomicName id="4C4A4D21FFC0CD0B54220BC5FD4DFE65" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[625,746,403,430]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFC0CD0B54220BC5FD4DFE65" box="[625,746,403,430]" italics="true" pageId="7" pageNumber="149">G. karinae</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC0CD0B54A20BC2FCECFE65" bold="true" box="[753,843,403,429]" pageId="7" pageNumber="149">
<taxonomicNameLabel id="A20D57CBFFC0CD0B54A20BC2FCECFE65" box="[753,843,403,429]" pageId="7" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C4A4D21FFC0CD0B55B20BC5FBF4FE65" authorityName="Vicente &amp; Mauries" authorityYear="1980" box="[993,1107,404,430]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="comasi">
<emphasis id="B93EEAB0FFC0CD0B55B20BC5FBF4FE65" box="[993,1107,404,430]" italics="true" pageId="7" pageNumber="149">G. comasi</emphasis>
</taxonomicName>
by subtriangular
<emphasis id="B93EEAB0FFC0CD0B53490BC5FA9EFE65" box="[1306,1337,404,429]" italics="true" pageId="7" pageNumber="149">vs.</emphasis>
rounded lateral parts of the colpocoxites in posterior view, or by less differentiated coxal processes on leg-pairs 7 and 11 compared to
<taxonomicName id="4C4A4D21FFC0CD0B577B0B8DFEDAFE3D" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[296,381,476,502]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="149" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFC0CD0B577B0B8DFEDAFE3D" box="[296,381,476,502]" italics="true" pageId="7" pageNumber="149">G. reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC0CD0B57D00B8AFE7BFE3D" bold="true" box="[387,476,475,501]" pageId="7" pageNumber="149">
<taxonomicNameLabel id="A20D57CBFFC0CD0B57D00B8AFE7BFE3D" box="[387,476,475,501]" pageId="7" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Significant differences also exist in the shape of the posterior part of the medial bursal structure of the vulvae in all three species.
</paragraph>
<paragraph id="8BF536A2FFC0CD0B56940872FC96FDAA" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="149">
<emphasis id="B93EEAB0FFC0CD0B56940872FEB3FDF5" bold="true" box="[199,276,547,573]" pageId="7" pageNumber="149">Name.</emphasis>
The new species is named in honour of colleague and friend, myriapodologist Karin Voigtländer (SMNG), one of the collectors. A noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFC0CD0B5694083AFDBEF9E5" pageId="7" pageNumber="149" type="materials_examined">
<paragraph id="8BF536A2FFC0CD0B5694083AFD4BFD4E" blockId="7.[151,1437,259,2014]" box="[199,748,619,646]" pageId="7" pageNumber="149">
<emphasis id="B93EEAB0FFC0CD0B5694083AFE16FD4D" bold="true" box="[199,433,619,645]" pageId="7" pageNumber="149">Material examined.</emphasis>
(
<specimenCount id="9D4CFD2BFFC0CD0B5793083AFDA5FD4E" box="[448,514,619,646]" pageId="7" pageNumber="149" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9D4CFD2BFFC0CD0B545D083DFDE9FD4E" box="[526,590,620,646]" pageId="7" pageNumber="149" type="female">7 ♀♀</specimenCount>
,
<specimenCount id="9D4CFD2BFFC0CD0B5408083AFD42FD4E" box="[603,741,619,646]" pageId="7" pageNumber="149" type="juvenile">13 juveniles</specimenCount>
)
</paragraph>
<paragraph id="8BF536A2FFC0CD0B569408DEFC70FD39" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="149">
<materialsCitation id="3B223CFFFFC0CD0B569408DEFC70FD39" ID-GBIF-Occurrence="3457637304" collectingDate="2009-04-20" collectionCode="SMNG" collectorName="K. Voigtlander" country="Spain" county="Leitza" elevation="550" latitude="43.0789" location="Gorriztaran" longLatPrecision="6" longitude="-1.8839" municipality="Ariz Mendiak" pageId="7" pageNumber="149" specimenCount="1" specimenCount-male="1" stateProvince="Navarra" typeStatus="holotype">
<emphasis id="B93EEAB0FFC0CD0B569408DEFE9AFD61" bold="true" box="[199,317,655,681]" pageId="7" pageNumber="149">
<typeStatus id="54F18800FFC0CD0B569408DEFE9FFD61" box="[199,312,655,681]" pageId="7" pageNumber="149" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F35D7632FFC0CD0B571508DEFE33FD61" box="[326,404,655,681]" name="Spain" pageId="7" pageNumber="149">SPAIN</collectingCountry>
●
<specimenCount id="9D4CFD2BFFC0CD0B57E508C1FE69FD62" box="[438,462,656,682]" pageId="7" pageNumber="149" type="male">♂</specimenCount>
;
<collectingRegion id="498EF840FFC0CD0B578F08C1FD03FD61" box="[476,676,655,682]" country="Spain" name="Navarra" pageId="7" pageNumber="149">Navarre province</collectingRegion>
,
<collectingCounty id="62944E2EFFC0CD0B54E308DEFD5EFD61" box="[688,761,655,681]" pageId="7" pageNumber="149">Leitza</collectingCounty>
,
<collectingMunicipality id="6B91ACD8FFC0CD0B555708DEFC03FD61" box="[772,932,655,681]" pageId="7" pageNumber="149">Ariz Mendiak</collectingMunicipality>
, between area “Ustarleku” and “Karobieta”,
<location id="8E956079FFC0CD0B56C408E2FEBFFD05" LSID="urn:lsid:plazi:treatment:03E387B4FFC0CD0056C40AC6FF61FF66:8E956079FFC0CD0B56C408E2FEBFFD05" box="[151,280,691,717]" country="Spain" county="Leitza" latitude="43.0789" longLatPrecision="6" longitude="-1.8839" municipality="Ariz Mendiak" name="Gorriztaran" pageId="7" pageNumber="149" stateProvince="Navarra">Gorriztaran</location>
/
<location id="8E956079FFC0CD0B577308E2FE5FFD05" LSID="urn:lsid:plazi:treatment:03E387B4FFC0CD0056C40AC6FF61FF66:8E956079FFC0CD0B577308E2FE5FFD05" box="[288,504,691,717]" country="Spain" county="Leitza" latitude="43.0789" longLatPrecision="6" longitude="-1.8839" municipality="Ariz Mendiak" name="Ibaia-stream valley" pageId="7" pageNumber="149" stateProvince="Navarra">Ibaia-stream valley</location>
,
<geoCoordinate id="EE7E5065FFC0CD0B545708E2FDC5FD05" box="[516,610,691,717]" degrees="43.0789" direction="north" orientation="latitude" pageId="7" pageNumber="149" precision="5" value="43.0789">43.0789</geoCoordinate>
,
<geoCoordinate id="EE7E5065FFC0CD0B543E08E2FD61FD05" box="[621,710,691,717]" degrees="1.8839" direction="west" orientation="longitude" pageId="7" pageNumber="149" precision="5" value="-1.8839">-1.8839</geoCoordinate>
,
<elevation id="0067D191FFC0CD0B548208E2FCF2FD05" box="[721,853,691,718]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="149" unit="m" value="550.0">
<quantity id="4CB29B47FFC0CD0B548208E2FCBEFD06" box="[721,793,691,718]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="149" unit="m" value="550.0">550 m</quantity>
a.s.l.
</elevation>
,
<taxonomicName id="4C4A4D21FFC0CD0B553108E2FC05FD05" box="[866,930,691,717]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B553108E2FC05FD05" box="[866,930,691,717]" italics="true" pageId="7" pageNumber="149">Alnus</emphasis>
</taxonomicName>
wood along stream, on the lower edge of the slope, under leaves and stones;
<date id="FFF41062FFC0CD0B57AB0886FD30FD39" box="[504,663,727,753]" pageId="7" pageNumber="149" value="2009-04-20">
<collectingDate id="EFB0E98AFFC0CD0B57AB0886FD30FD39" box="[504,663,727,753]" pageId="7" pageNumber="149" value="2009-04-20">20 April 2009</collectingDate>
</date>
;
<collectorName id="26BF5374FFC0CD0B54F00889FCE1FD39" box="[675,838,727,754]" pageId="7" pageNumber="149">K. Voigtländer</collectorName>
leg;
<collectionCode id="ED5BAE67FFC0CD0B552D0886FC74FD39" box="[894,979,727,753]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="7" pageNumber="149">SMNG</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF536A2FFC0CD0B569408AAFDBEF9E5" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="149">
<emphasis id="B93EEAB0FFC0CD0B569408AAFEEFFCDD" bold="true" box="[199,328,763,789]" pageId="7" pageNumber="149">
<typeStatus id="54F18800FFC0CD0B569408AAFEE4FCDD" box="[199,323,763,789]" pageId="7" pageNumber="149" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F35D7632FFC0CD0B570208AAFE38FCDD" box="[337,415,763,789]" name="Spain" pageId="7" pageNumber="149">SPAIN</collectingCountry>
●
<specimenCount id="9D4CFD2BFFC0CD0B579308AAFE4AFCDE" box="[448,493,763,790]" pageId="7" pageNumber="149" type="female">1 ♀</specimenCount>
; same data as for
<typeStatus id="54F18800FFC0CD0B54EE08AAFC84FCDD" box="[701,803,763,789]" pageId="7" pageNumber="149" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED5BAE67FFC0CD0B557C08ADFCFBFCDE" box="[815,860,764,790]" pageId="7" pageNumber="149">IZB</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B552D08AAFC55FCDD" box="[894,1010,763,789]" pageId="7" pageNumber="149" type="juvenile">1 juvenile</specimenCount>
; same data as for
<typeStatus id="54F18800FFC0CD0B529308AAFA81FCDD" box="[1216,1318,763,789]" pageId="7" pageNumber="149" type="holotype">holotype</typeStatus>
;
<collectionCode id="ED5BAE67FFC0CD0B536108AAFA24FCDD" box="[1330,1411,763,789]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="7" pageNumber="149">SMNG</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B56C4094EFF63FCF2" box="[151,196,799,826]" pageId="7" pageNumber="149" type="female">1 ♀</specimenCount>
,
<collectingRegion id="498EF840FFC0CD0B56860971FE38FCF1" box="[213,415,799,826]" country="Spain" name="Navarra" pageId="7" pageNumber="149">Navarre province</collectingRegion>
, Leitza, Ariz Mendiak, between area “Ustarleku” and “Karobieta”, above side stream to Gorriztaran,
<geoCoordinate id="EE7E5065FFC0CD0B57740912FE22FC95" box="[295,389,835,861]" degrees="43.0778" direction="north" orientation="latitude" pageId="7" pageNumber="149" precision="5" value="43.0778">43.0778</geoCoordinate>
,
<geoCoordinate id="EE7E5065FFC0CD0B57C30912FE4EFC96" box="[400,489,835,862]" degrees="1.8775" direction="west" orientation="longitude" pageId="7" pageNumber="149" precision="5" value="-1.8775">-1.8775</geoCoordinate>
,
<quantity id="4CB29B47FFC0CD0B57A60912FD9BFC96" box="[501,572,835,862]" metricMagnitude="2" metricUnit="m" metricValue="6.15" pageId="7" pageNumber="149" unit="m" value="615.0">615 m</quantity>
a.s.l., grove of
<taxonomicName id="4C4A4D21FFC0CD0B54B80915FCF2FC96" box="[747,853,836,862]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B54B80915FCF2FC96" box="[747,853,836,862]" italics="true" pageId="7" pageNumber="149">Castanea</emphasis>
</taxonomicName>
, pollard trees on the slope with
<taxonomicName id="4C4A4D21FFC0CD0B52EC0912FA31FC95" box="[1215,1430,835,861]" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="species" species="ficaria">
<emphasis id="B93EEAB0FFC0CD0B52EC0912FA31FC95" box="[1215,1430,835,861]" italics="true" pageId="7" pageNumber="149">Ranunculus ficaria</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B56C40936FF57FC49" box="[151,240,871,897]" class="Magnoliopsida" family="Thymelaeaceae" genus="Daphne" kingdom="Plantae" order="Malvales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B56C40936FF57FC49" box="[151,240,871,897]" italics="true" pageId="7" pageNumber="149">Daphne</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B56AC0936FEDCFC49" box="[255,379,871,897]" class="Magnoliopsida" family="Ranunculaceae" genus="Helleborus" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B56AC0936FEDCFC49" box="[255,379,871,897]" italics="true" pageId="7" pageNumber="149">Helleborus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B57D90936FE77FC49" box="[394,464,871,897]" class="Magnoliopsida" family="Lamiaceae" genus="Salvia" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B57D90936FE77FC49" box="[394,464,871,897]" italics="true" pageId="7" pageNumber="149">Salvia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B578D0936FD2CFC49" authority=", Lathrea" authorityName="Lathrea" box="[478,651,871,898]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B578D0936FD83FC49" box="[478,548,871,897]" italics="true" pageId="7" pageNumber="149">Rubus</emphasis>
,
<emphasis id="B93EEAB0FFC0CD0B54600936FD2CFC49" box="[563,651,871,897]" italics="true" pageId="7" pageNumber="149">Lathrea</emphasis>
</taxonomicName>
, loamy and calcareous soil, under bark of thick rotten trunk;
<date id="FFF41062FFC0CD0B536C0936FF75FC6D" pageId="7" pageNumber="149" value="2009-04-20">20 April 2009</date>
; N. Lindner leg.;
<collectionCode id="ED5BAE67FFC0CD0B57C809DAFE4BFC6D" box="[411,492,907,933]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="7" pageNumber="149">SMNG</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B545F09DAFD90FC6E" box="[524,567,907,934]" pageId="7" pageNumber="149" type="male">1 ♂</specimenCount>
(used for
<collectionCode id="ED5BAE67FFC0CD0B54F809DAFD4EFC6D" box="[683,745,907,933]" pageId="7" pageNumber="149">SEM</collectionCode>
),
<specimenCount id="9D4CFD2BFFC0CD0B54AE09DAFC8EFC6E" box="[765,809,907,934]" pageId="7" pageNumber="149" type="female">1 ♀</specimenCount>
;
<collectingRegion id="498EF840FFC0CD0B556409DDFC59FC6D" box="[823,1022,907,934]" country="Spain" name="Navarra" pageId="7" pageNumber="149">Navarre province</collectingRegion>
, Leitza, Ariz Mendiak, between area “Ustarleku” and “Karobieta”, above side stream to Gorriztaran, north of chestnut-area,
<geoCoordinate id="EE7E5065FFC0CD0B52D209FEFB78FC02" box="[1153,1247,943,970]" degrees="43.0775" direction="north" orientation="latitude" pageId="7" pageNumber="149" precision="5" value="43.0775">43.0775</geoCoordinate>
,
<geoCoordinate id="EE7E5065FFC0CD0B52BF09FEFAE2FC01" box="[1260,1349,943,969]" degrees="1.8783" direction="west" orientation="longitude" pageId="7" pageNumber="149" precision="5" value="-1.8783">-1.8783</geoCoordinate>
,
<quantity id="4CB29B47FFC0CD0B530009FEFA3BFC02" box="[1363,1436,943,970]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="149" unit="m" value="600.0">600 m</quantity>
a.s.l., beach forest,
<taxonomicName id="4C4A4D21FFC0CD0B57230982FDA6FC25" box="[368,513,979,1005]" class="Magnoliopsida" family="Araliaceae" genus="Hedera" kingdom="Plantae" order="Apiales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="species" species="helix">
<emphasis id="B93EEAB0FFC0CD0B57230982FDA6FC25" box="[368,513,979,1005]" italics="true" pageId="7" pageNumber="149">Hedera helix</emphasis>
</taxonomicName>
, under vegetation,
<taxonomicName id="4C4A4D21FFC0CD0B54890982FC87FC25" box="[730,800,979,1005]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B54890982FC87FC25" box="[730,800,979,1005]" italics="true" pageId="7" pageNumber="149">Rubus</emphasis>
</taxonomicName>
, in dead leaves;
<date id="FFF41062FFC0CD0B558F0982FBDCFC25" box="[988,1147,979,1005]" pageId="7" pageNumber="149" value="2009-04-20">20 April 2009</date>
; P. Djursvoll leg.;
<collectionCode id="ED5BAE67FFC0CD0B53180985FA3BFC26" box="[1355,1436,980,1006]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="7" pageNumber="149">ZMBN</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B56E209A6FF7AFBDA" box="[177,221,1015,1042]" pageId="7" pageNumber="149" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D4CFD2BFFC0CD0B56BE09A6FECAFBD9" box="[237,365,1015,1041]" pageId="7" pageNumber="149" type="juvenile">3 juveniles</specimenCount>
;
<collectingRegion id="498EF840FFC0CD0B572909A9FDE5FBD9" box="[378,578,1015,1042]" country="Spain" name="Navarra" pageId="7" pageNumber="149">Navarre province</collectingRegion>
, Leitza, Ariz Mendiak, road NA-
<quantity id="4CB29B47FFC0CD0B559309A6FBA0FBD9" box="[960,1031,1015,1041]" metricMagnitude="0" metricUnit="m" metricValue="4.318" pageId="7" pageNumber="149" unit="in" value="170.0">170 in</quantity>
direction to Ezkurra, ca.
<quantity id="4CB29B47FFC0CD0B537E09A6FACFFBD9" box="[1325,1384,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="149" unit="km" value="3.0">3 km</quantity>
east of Leitza,
<geoCoordinate id="EE7E5065FFC0CD0B575E0E4AFECCFBFD" box="[269,363,1051,1077]" degrees="43.0900" direction="north" orientation="latitude" pageId="7" pageNumber="149" precision="5" value="43.09">43.0900</geoCoordinate>
,
<geoCoordinate id="EE7E5065FFC0CD0B572B0E4AFE76FBFE" box="[376,465,1051,1078]" degrees="1.8953" direction="west" orientation="longitude" pageId="7" pageNumber="149" precision="5" value="-1.8953">-1.8953</geoCoordinate>
,
<quantity id="4CB29B47FFC0CD0B578C0E4AFD80FBFE" box="[479,551,1051,1078]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="149" unit="m" value="580.0">580 m</quantity>
a.s.l., narrow, steep brook valley,
<taxonomicName id="4C4A4D21FFC0CD0B55E20E4AFC56FBFD" box="[945,1009,1051,1077]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B55E20E4AFC56FBFD" box="[945,1009,1051,1077]" italics="true" pageId="7" pageNumber="149">Alnus</emphasis>
</taxonomicName>
(young growth),
<taxonomicName id="4C4A4D21FFC0CD0B52E90E4AFAB4FBFD" authorityName="C.Linnaeus" authorityYear="1753" box="[1210,1299,1051,1077]" class="Magnoliopsida" family="Betulaceae" genus="Corylus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B52E90E4AFAB4FBFD" box="[1210,1299,1051,1077]" italics="true" pageId="7" pageNumber="149">Corylus</emphasis>
</taxonomicName>
etc., in leaf litter near stream;
<date id="FFF41062FFC0CD0B573E0E6EFDB6FB91" box="[365,529,1087,1113]" pageId="7" pageNumber="149" value="2009-04-20">20 April 2009</date>
; H. Reip leg.;
<collectionCode id="ED5BAE67FFC0CD0B54EC0E6EFCB7FB91" box="[703,784,1087,1113]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="7" pageNumber="149">SMNG</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B55660E6EFCC5FB92" box="[821,866,1087,1114]" pageId="7" pageNumber="149" type="female">1 ♀</specimenCount>
(used for
<collectionCode id="ED5BAE67FFC0CD0B558E0E6EFBBCFB91" box="[989,1051,1087,1113]" pageId="7" pageNumber="149">SEM</collectionCode>
), same data as previous;
<collectionCode id="ED5BAE67FFC0CD0B53120E11FA3BFB92" box="[1345,1436,1088,1114]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="149" type="Museum">NHMW</collectionCode>
MY10253 ●
<specimenCount id="9D4CFD2BFFC0CD0B577F0E32FEFEFBB6" box="[300,345,1123,1150]" pageId="7" pageNumber="149" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D4CFD2BFFC0CD0B57350E32FE42FBB5" box="[358,485,1123,1149]" pageId="7" pageNumber="149" type="juvenile">6 juveniles</specimenCount>
;
<collectingRegion id="498EF840FFC0CD0B57A20E35FD1EFBB5" box="[497,697,1123,1150]" country="Spain" name="Navarra" pageId="7" pageNumber="149">Navarre province</collectingRegion>
, Leitza, Ariz Mendiak, at area “Kornieta”,
<geoCoordinate id="EE7E5065FFC0CD0B52F10E32FAA7FBB5" box="[1186,1280,1123,1149]" degrees="43.0698" direction="north" orientation="latitude" pageId="7" pageNumber="149" precision="5" value="43.0698">43.0698</geoCoordinate>
,
<geoCoordinate id="EE7E5065FFC0CD0B535E0E32FAC2FBB5" box="[1293,1381,1123,1149]" degrees="1.8823" direction="west" orientation="longitude" pageId="7" pageNumber="149" precision="5" value="-1.8823">-1.8823</geoCoordinate>
,
<quantity id="4CB29B47FFC0CD0B53210E32FF0AFB6A" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="149" unit="m" value="910.0">910 m</quantity>
a.s.l.,
<taxonomicName id="4C4A4D21FFC0CD0B56A40ED9FE9AFB69" box="[247,317,1160,1185]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B56A40ED9FE9AFB69" box="[247,317,1160,1185]" italics="true" pageId="7" pageNumber="149">Fagus</emphasis>
</taxonomicName>
woodland, poor understorey, rocky, siliceous soil, mosses,
<taxonomicName id="4C4A4D21FFC0CD0B55840ED6FBF4FB69" box="[983,1107,1159,1185]" class="Magnoliopsida" family="Ranunculaceae" genus="Helleborus" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B55840ED6FBF4FB69" box="[983,1107,1159,1185]" italics="true" pageId="7" pageNumber="149">Helleborus</emphasis>
</taxonomicName>
, under bark and in leaf litter;
<date id="FFF41062FFC0CD0B56C40EFAFE9EFB0D" box="[151,313,1195,1221]" pageId="7" pageNumber="149" value="2009-04-20">20 April 2009</date>
; H. Reip leg.;
<collectionCode id="ED5BAE67FFC0CD0B57B20EFDFDA9FB0E" box="[481,526,1196,1222]" pageId="7" pageNumber="149">IZB</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B54630EFAFDF8FB0E" box="[560,607,1195,1222]" pageId="7" pageNumber="149" type="male">1 ♂</specimenCount>
;
<collectingRegion id="498EF840FFC0CD0B543F0EFDFC93FB0D" box="[620,820,1195,1222]" country="Spain" name="Navarra" pageId="7" pageNumber="149">Navarre province</collectingRegion>
, Sierra de Aralar, south Baraibar, on road NA 7510 to Santuario de San Migel, at area “Urkia”,
<geoCoordinate id="EE7E5065FFC0CD0B542A0E9EFD70FB21" box="[633,727,1231,1257]" degrees="42.9762" direction="north" orientation="latitude" pageId="7" pageNumber="149" precision="5" value="42.9762">42.9762</geoCoordinate>
,
<geoCoordinate id="EE7E5065FFC0CD0B54B50E9EFC99FB21" box="[742,830,1231,1257]" degrees="1.9318" direction="west" orientation="longitude" pageId="7" pageNumber="149" precision="5" value="-1.9318">-1.9318</geoCoordinate>
,
<quantity id="4CB29B47FFC0CD0B551E0E9EFC30FB22" box="[845,919,1231,1258]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="7" pageNumber="149" unit="m" value="710.0">710 m</quantity>
a.s.l., wet wood pasture,
<taxonomicName id="4C4A4D21FFC0CD0B52900E9EFABBFB21" authorityName="C.Linnaeus" authorityYear="1753" box="[1219,1308,1231,1257]" class="Magnoliopsida" family="Betulaceae" genus="Corylus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B52900E9EFABBFB21" box="[1219,1308,1231,1257]" italics="true" pageId="7" pageNumber="149">Corylus</emphasis>
</taxonomicName>
with some
<taxonomicName id="4C4A4D21FFC0CD0B56C40EA5FF52FAC6" box="[151,245,1268,1294]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B56C40EA5FF52FAC6" box="[151,245,1268,1294]" italics="true" pageId="7" pageNumber="149">Quercus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B57560EA2FE88FAC5" box="[261,303,1267,1293]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B57560EA2FE88FAC5" box="[261,303,1267,1293]" italics="true" pageId="7" pageNumber="149">Ilex</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B57130EA2FDBFFAC5" box="[320,536,1267,1293]" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="species" species="ficaria">
<emphasis id="B93EEAB0FFC0CD0B57130EA2FDBFFAC5" box="[320,536,1267,1293]" italics="true" pageId="7" pageNumber="149">Ranunculus ficaria</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B547B0EA2FD26FAC5" box="[552,641,1267,1293]" class="Magnoliopsida" family="Thymelaeaceae" genus="Daphne" kingdom="Plantae" order="Malvales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B547B0EA2FD26FAC5" box="[552,641,1267,1293]" italics="true" pageId="7" pageNumber="149">Daphne</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B54C20EA2FD7DFAC5" box="[657,730,1267,1293]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B54C20EA2FD7DFAC5" box="[657,730,1267,1293]" italics="true" pageId="7" pageNumber="149">Oxalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B54B80EA2FCC0FAC5" box="[747,871,1267,1293]" class="Magnoliopsida" family="Ranunculaceae" genus="Helleborus" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B54B80EA2FCC0FAC5" box="[747,871,1267,1293]" italics="true" pageId="7" pageNumber="149">Helleborus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B55240EA2FC48FAC5" box="[887,1007,1267,1293]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B55240EA2FC48FAC5" box="[887,1007,1267,1293]" italics="true" pageId="7" pageNumber="149">Euphorbia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B52530EA5FB6CFAC5" authority=", Lathre" authorityName="Lathre" box="[1024,1227,1267,1294]" class="Magnoliopsida" family="Geraniaceae" genus="Geranium" kingdom="Plantae" order="Geraniales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B52530EA5FBD5FAC6" box="[1024,1138,1268,1294]" italics="true" pageId="7" pageNumber="149">Geranium</emphasis>
,
<emphasis id="B93EEAB0FFC0CD0B52D20EA2FB6CFAC5" box="[1153,1227,1267,1293]" italics="true" pageId="7" pageNumber="149">Lathre</emphasis>
</taxonomicName>
a, in leaf litter;
<date id="FFF41062FFC0CD0B53D30EA2FEB3FAF9" pageId="7" pageNumber="149" value="2009-04-22">22 April 2009</date>
; P. Djursvoll leg.;
<collectionCode id="ED5BAE67FFC0CD0B57B10F49FD94FAFA" box="[482,563,1304,1330]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="7" pageNumber="149">ZMBN</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B54010F46FDDAFAFA" box="[594,637,1303,1330]" pageId="7" pageNumber="149" type="female">1 ♀</specimenCount>
; same as previous but wet wood pasture of
<emphasis id="B93EEAB0FFC0CD0B52360F49FB07FAF9" box="[1125,1184,1304,1329]" italics="true" pageId="7" pageNumber="149">Fagu</emphasis>
s,
<taxonomicName id="4C4A4D21FFC0CD0B52EA0F46FA32FAF9" box="[1209,1429,1303,1329]" class="Magnoliopsida" family="Ranunculaceae" genus="Anemone" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="species" species="nemorosa">
<emphasis id="B93EEAB0FFC0CD0B52EA0F46FA32FAF9" box="[1209,1429,1303,1329]" italics="true" pageId="7" pageNumber="149">Anemone nemorosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B56C40F6AFF72FA9D" box="[151,213,1339,1365]" class="Liliopsida" family="Araceae" genus="Arum" kingdom="Plantae" order="Alismatales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B56C40F6AFF72FA9D" box="[151,213,1339,1365]" italics="true" pageId="7" pageNumber="149">Arum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B56B10F6DFEF3FA9E" box="[226,340,1340,1366]" class="Magnoliopsida" family="Geraniaceae" genus="Geranium" kingdom="Plantae" order="Geraniales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B56B10F6DFEF3FA9E" box="[226,340,1340,1366]" italics="true" pageId="7" pageNumber="149">Geranium</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B57330F6AFE7BFA9D" box="[352,476,1339,1365]" class="Magnoliopsida" family="Ranunculaceae" genus="Helleborus" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B57330F6AFE7BFA9D" box="[352,476,1339,1365]" italics="true" pageId="7" pageNumber="149">Helleborus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4A4D21FFC0CD0B57BA0F6AFD41FA9D" authority=", Mercuriali" authorityName="Mercuriali" box="[489,742,1339,1366]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B57BA0F6AFDC6FA9D" box="[489,609,1339,1365]" italics="true" pageId="7" pageNumber="149">Euphorbia</emphasis>
,
<emphasis id="B93EEAB0FFC0CD0B543D0F6AFD41FA9D" box="[622,742,1339,1365]" italics="true" pageId="7" pageNumber="149">Mercuriali</emphasis>
</taxonomicName>
s,
<taxonomicName id="4C4A4D21FFC0CD0B54AD0F6AFC74FA9D" box="[766,979,1339,1365]" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" kingdom="Plantae" order="Ranunculales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="species" species="ficaria">
<emphasis id="B93EEAB0FFC0CD0B54AD0F6AFC74FA9D" box="[766,979,1339,1365]" italics="true" pageId="7" pageNumber="149">Ranunculus ficaria</emphasis>
</taxonomicName>
, under stones and logs;
<date id="FFF41062FFC0CD0B52880F6AFAD1FA9D" box="[1243,1398,1339,1365]" pageId="7" pageNumber="149" value="2009-04-22">22 April 2009</date>
; K. Voigtländer leg.;
<collectionCode id="ED5BAE67FFC0CD0B570A0F0EFE0DFAB1" box="[345,426,1375,1401]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="7" pageNumber="149">SMNG</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B57990F0EFE51FAB2" box="[458,502,1375,1402]" pageId="7" pageNumber="149" type="female">1 ♀</specimenCount>
;
<collectingRegion id="498EF840FFC0CD0B54570F31FD6CFAB1" box="[516,715,1375,1402]" country="Spain" name="Navarra" pageId="7" pageNumber="149">Navarre province</collectingRegion>
, Sierra de Aralar, south Baraibar, on road NA 7510 to Santuario de San Migel, at area “Izáin”,
<geoCoordinate id="EE7E5065FFC0CD0B57BA0FD2FDE0FA55" box="[489,583,1411,1437]" degrees="42.9714" direction="north" orientation="latitude" pageId="7" pageNumber="149" precision="5" value="42.9714">42.9714</geoCoordinate>
,
<geoCoordinate id="EE7E5065FFC0CD0B54010FD2FD0CFA55" box="[594,683,1411,1437]" degrees="1.9384" direction="west" orientation="longitude" pageId="7" pageNumber="149" precision="5" value="-1.9384">-1.9384</geoCoordinate>
,
<quantity id="4CB29B47FFC0CD0B54E50FD2FD5AFA56" box="[694,765,1411,1438]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="7" pageNumber="149" unit="m" value="790.0">790 m</quantity>
a.s.l., karst area with deep grykes and bare limestone rocks,
<taxonomicName id="4C4A4D21FFC0CD0B56C40FF9FF7AFA09" box="[151,221,1448,1473]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B56C40FF9FF7AFA09" box="[151,221,1448,1473]" italics="true" pageId="7" pageNumber="149">Fagus</emphasis>
</taxonomicName>
woodland &amp; some
<emphasis id="B93EEAB0FFC0CD0B57910FF9FD8EFA0A" box="[450,553,1448,1474]" italics="true" pageId="7" pageNumber="149">Crategus</emphasis>
bushes, in leaf litter;
<date id="FFF41062FFC0CD0B55740FF6FC6CFA09" box="[807,971,1447,1473]" pageId="7" pageNumber="149" value="2009-04-22">22 April 2009</date>
; P. Djursvoll leg.;
<collectionCode id="ED5BAE67FFC0CD0B52F00FF9FB53FA0A" box="[1187,1268,1448,1474]" country="Norway" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="7" pageNumber="149">ZMBN</collectionCode>
●
<specimenCount id="9D4CFD2BFFC0CD0B534B0FF6FA3FFA09" box="[1304,1432,1447,1473]" pageId="7" pageNumber="149" type="juvenile">3 juveniles</specimenCount>
; same as previous but
<geoCoordinate id="EE7E5065FFC0CD0B57DF0F9AFE4DFA2D" box="[396,490,1483,1509]" degrees="42.9724" direction="north" orientation="latitude" pageId="7" pageNumber="149" precision="5" value="42.9724">42.9724</geoCoordinate>
,
<geoCoordinate id="EE7E5065FFC0CD0B57A50F9AFDE8FA2D" box="[502,591,1483,1509]" degrees="1.9361" direction="west" orientation="longitude" pageId="7" pageNumber="149" precision="5" value="-1.9361">-1.9361</geoCoordinate>
,
<quantity id="4CB29B47FFC0CD0B540F0F9AFD03FA2E" box="[604,676,1483,1510]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="149" unit="m" value="730.0">730 m</quantity>
a.s.l., karst area with deep grykes and bare limestone rocks,
<taxonomicName id="4C4A4D21FFC0CD0B53050F9DFA3BFA2D" box="[1366,1436,1484,1509]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B53050F9DFA3BFA2D" box="[1366,1436,1484,1509]" italics="true" pageId="7" pageNumber="149">Fagus</emphasis>
</taxonomicName>
woodland &amp; some
<emphasis id="B93EEAB0FFC0CD0B57220FA1FE7FF9C2" box="[369,472,1520,1546]" italics="true" pageId="7" pageNumber="149">Crategus</emphasis>
bushes, ground of deep and collapsed gryke, sieved from
<taxonomicName id="4C4A4D21FFC0CD0B52250FA1FB1BF9C1" box="[1142,1212,1520,1545]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="149" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC0CD0B52250FA1FB1BF9C1" box="[1142,1212,1520,1545]" italics="true" pageId="7" pageNumber="149">Fagus</emphasis>
</taxonomicName>
leaf litter;
<date id="FFF41062FFC0CD0B536D0FBEFF75F9E5" pageId="7" pageNumber="149" value="2009-04-22">22 April 2009</date>
; K. Voigtländer leg.;
<collectionCode id="ED5BAE67FFC0CD0B57920C42FDB1F9E5" box="[449,534,1555,1581]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="7" pageNumber="149">SMNG</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFC0CD0456940C66FB4DF9BD" lastPageId="8" lastPageNumber="150" pageId="7" pageNumber="149" type="description">
<paragraph id="8BF536A2FFC0CD0B56940C66FE94F976" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="149">
<emphasis id="B93EEAB0FFC0CD0B56940C66FEFEF999" bold="true" box="[199,345,1591,1617]" pageId="7" pageNumber="149">Description.</emphasis>
SIZE AND NUMBER OF BODY RINGS. Body in adults with 30 rings (including telson).
<typeStatus id="54F18800FFC0CD0B52A60C66FAFAF999" box="[1269,1373,1591,1617]" pageId="7" pageNumber="149" type="holotype">Holotype</typeStatus>
male
<quantity id="4CB29B47FFC0CD0B56C40C0AFF57F9BE" box="[151,240,1627,1654]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="7" pageNumber="149" unit="mm" value="7.6">7.6 mm</quantity>
long, vertical diameter of the largest ring
<quantity id="4CB29B47FFC0CD0B548F0C0AFC9FF9BE" box="[732,824,1627,1654]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="149" unit="mm" value="0.7">0.7 mm</quantity>
.
<typeStatus id="54F18800FFC0CD0B55160C0DFC00F9BE" box="[837,935,1628,1654]" pageId="7" pageNumber="149" type="paratype">Paratype</typeStatus>
males
<quantity id="4CB29B47FFC0CD0B55A80C0AFBD7F9BE" box="[1019,1136,1627,1654]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="7" pageNumber="149" unit="mm" value="7.25" valueMax="7.5" valueMin="7.0">7–7.5 mm</quantity>
long, vertical diameter of the largest ring
<quantity id="4CB29B47FFC0CD0B57190C2EFE44F952" box="[330,483,1663,1690]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="7" pageNumber="149" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">0.65–0.7 mm</quantity>
, respectively.
<typeStatus id="54F18800FFC0CD0B54D70CD1FD41F952" box="[644,742,1664,1690]" pageId="7" pageNumber="149" type="paratype">Paratype</typeStatus>
females
<quantity id="4CB29B47FFC0CD0B551D0C2EFC65F952" box="[846,962,1663,1690]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" metricValueMax="9.0" metricValueMin="7.8" pageId="7" pageNumber="149" unit="mm" value="8.4" valueMax="9.0" valueMin="7.8">7.8–9 mm</quantity>
long, vertical diameter of the largest ring
<quantity id="4CB29B47FFC0CD0B56C40CF2FE88F976" box="[151,303,1699,1726]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="7" pageNumber="149" unit="mm" value="0.775" valueMax="0.8" valueMin="0.75">0.75–0.8 mm</quantity>
.
</paragraph>
<paragraph id="8BF536A2FFC0CD0B56940C96FE26F8CD" blockId="7.[151,1437,259,2014]" pageId="7" pageNumber="149">
COLORATION (
<figureCitation id="13712A27FFC0CD0B570E0C96FE06F929" box="[349,417,1735,1761]" captionStart="FIGURE 2" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[185,1403,181,1920]" captionTargetId="figure-19@9.[185,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 2. Guipuzcosoma karinae sp. nov., habitus (ZMBN). A. ♂ paratype, lateral view. B. ♂ paratype, anterior part of body, showing head and collum in dorsal view. C. ♂ paratype, anterior part of body, lateral view. D. ♀ paratype, midbody rings, dorsal view. E. ♂ paratype, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905145" httpUri="https://zenodo.org/record/5905145/files/figure.png" pageId="7" pageNumber="149">Fig. 2</figureCitation>
). Variable. From pale yellowish to brownish, with greyish patterns. Ventral halves lighter. Ommatidia blackish.
</paragraph>
<paragraph id="8BF536A2FFC0CD0456940D41FCD2FF1D" blockId="7.[151,1437,259,2014]" lastBlockId="8.[151,1437,151,1834]" lastPageId="8" lastPageNumber="150" pageId="7" pageNumber="149">
HEAD (
<figureCitation id="13712A27FFC0CD0B57470D5EFE14F8E2" box="[276,435,1807,1834]" captionStart="FIGURE 2" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[185,1403,181,1920]" captionTargetId="figure-19@9.[185,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 2. Guipuzcosoma karinae sp. nov., habitus (ZMBN). A. ♂ paratype, lateral view. B. ♂ paratype, anterior part of body, showing head and collum in dorsal view. C. ♂ paratype, anterior part of body, lateral view. D. ♀ paratype, midbody rings, dorsal view. E. ♂ paratype, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905145" httpUri="https://zenodo.org/record/5905145/files/figure.png" pageId="7" pageNumber="149">Figs 2B, C, E</figureCitation>
,
<figureCitation id="13712A27FFC0CD0B57930D5EFDA1F8E1" box="[448,518,1807,1833]" captionStart="FIGURE 3" captionStartId="10.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-17@10.[151,1436,181,1649]" captionTargetPageId="10" captionText="FIGURE 3. Guipuzcosoma karinae sp. nov., ♂ paratype, habitus (ZMBN). A. Head, anterior view. B. Head and collum, lateral view. C. Left antenna, anterior view. D. Field with ommatidia, lateral view. E, F. Body rings 15–17, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A–C, E, F), 0.02 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.5905147" httpUri="https://zenodo.org/record/5905147/files/figure.png" pageId="7" pageNumber="149">3A–D</figureCitation>
). Setose, frontal side slightly concave in males, convex in females. Labrum with three medial teeth and 4+4 labral and 2+2 supralabral setae (
<figureCitation id="13712A27FFC0CD0B55650D62FC28F885" box="[822,911,1843,1869]" captionStart="FIGURE 3" captionStartId="10.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-17@10.[151,1436,181,1649]" captionTargetPageId="10" captionText="FIGURE 3. Guipuzcosoma karinae sp. nov., ♂ paratype, habitus (ZMBN). A. Head, anterior view. B. Head and collum, lateral view. C. Left antenna, anterior view. D. Field with ommatidia, lateral view. E, F. Body rings 15–17, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A–C, E, F), 0.02 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.5905147" httpUri="https://zenodo.org/record/5905147/files/figure.png" pageId="7" pageNumber="149">Fig. 3A</figureCitation>
). Promentum triangular, without setae. Lingual plates with 4+4 setae arranged in row. Stipites with ca. 17 setae each. Antennae
<quantity id="4CB29B47FFC0CD0B526B0D06FB37F8BA" box="[1080,1168,1879,1906]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="149" unit="mm" value="1.2">1.2 mm</quantity>
long in
<typeStatus id="54F18800FFC0CD0B52A00D06FAF2F8B9" box="[1267,1365,1879,1905]" pageId="7" pageNumber="149" type="holotype">holotype</typeStatus>
male. Length of antennomeres (in mm): I (0.08), II (0.13), III (0.28), IV (0.18), V (0.25), VI (0.12), VII (0.12) and VIII (0.04). Length/breadth ratios of antennomeres I–VII: I (1), II (1.6), III (3.5), IV (2.6), V (2.3), VI (1.3) and VII (1.5). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum trichoideum, respectively (
<figureCitation id="13712A27FFC0CD0B536B0D92FA28F815" box="[1336,1423,1987,2013]" captionStart="FIGURE 3" captionStartId="10.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-17@10.[151,1436,181,1649]" captionTargetPageId="10" captionText="FIGURE 3. Guipuzcosoma karinae sp. nov., ♂ paratype, habitus (ZMBN). A. Head, anterior view. B. Head and collum, lateral view. C. Left antenna, anterior view. D. Field with ommatidia, lateral view. E, F. Body rings 15–17, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A–C, E, F), 0.02 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.5905147" httpUri="https://zenodo.org/record/5905147/files/figure.png" pageId="7" pageNumber="149">Fig. 3C</figureCitation>
). Lateral to antennal sockets a group of papillae-like outgrowths present (
<figureCitation id="13712A27FFCFCD04559D0AC6FB80FF79" box="[974,1063,151,177]" captionStart="FIGURE 3" captionStartId="10.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-17@10.[151,1436,181,1649]" captionTargetPageId="10" captionText="FIGURE 3. Guipuzcosoma karinae sp. nov., ♂ paratype, habitus (ZMBN). A. Head, anterior view. B. Head and collum, lateral view. C. Left antenna, anterior view. D. Field with ommatidia, lateral view. E, F. Body rings 15–17, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A–C, E, F), 0.02 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.5905147" httpUri="https://zenodo.org/record/5905147/files/figure.png" pageId="8" pageNumber="150">Fig. 3B</figureCitation>
). Number of ommatidia:
<quantity id="4CB29B47FFCFCD0453180AC6FA21FF79" box="[1355,1414,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="8" pageNumber="150" unit="in" value="13.0">13 in</quantity>
4 rows in both sexes, arranged in irregular subtriangle (
<figureCitation id="13712A27FFCFCD0454BD0AEAFCCEFF1D" box="[750,873,187,213]" captionStart="FIGURE 3" captionStartId="10.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-17@10.[151,1436,181,1649]" captionTargetPageId="10" captionText="FIGURE 3. Guipuzcosoma karinae sp. nov., ♂ paratype, habitus (ZMBN). A. Head, anterior view. B. Head and collum, lateral view. C. Left antenna, anterior view. D. Field with ommatidia, lateral view. E, F. Body rings 15–17, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A–C, E, F), 0.02 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.5905147" httpUri="https://zenodo.org/record/5905147/files/figure.png" pageId="8" pageNumber="150">Fig. 3B, D</figureCitation>
).
</paragraph>
<paragraph id="8BF536A2FFCFCD0456940A8EFE3AFED5" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">COLLUM. Narrower than head, with six macrochaetae, as all body rings. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<paragraph id="8BF536A2FFCFCD0456940B76FB81FE41" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">
BODY RINGS (
<figureCitation id="13712A27FFCFCD04570B0B76FE3AFE89" box="[344,413,295,321]" captionStart="FIGURE 2" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[185,1403,181,1920]" captionTargetId="figure-19@9.[185,1403,181,1920]" captionTargetPageId="9" captionText="FIGURE 2. Guipuzcosoma karinae sp. nov., habitus (ZMBN). A. ♂ paratype, lateral view. B. ♂ paratype, anterior part of body, showing head and collum in dorsal view. C. ♂ paratype, anterior part of body, lateral view. D. ♀ paratype, midbody rings, dorsal view. E. ♂ paratype, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905145" httpUri="https://zenodo.org/record/5905145/files/figure.png" pageId="8" pageNumber="150">
Fig.
<geoCoordinate id="EE7E5065FFCFCD0457DE0B76FE3AFE89" box="[397,413,295,321]" degrees="2, 3" direction="east" orientation="longitude" pageId="8" pageNumber="150" precision="5555" value="2.3">2</geoCoordinate>
</figureCitation>
,
<figureCitation id="13712A27FFCFCD0457F90B76FE4EFE89" box="[426,489,295,321]" captionStart="FIGURE 3" captionStartId="10.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-17@10.[151,1436,181,1649]" captionTargetPageId="10" captionText="FIGURE 3. Guipuzcosoma karinae sp. nov., ♂ paratype, habitus (ZMBN). A. Head, anterior view. B. Head and collum, lateral view. C. Left antenna, anterior view. D. Field with ommatidia, lateral view. E, F. Body rings 15–17, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A–C, E, F), 0.02 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.5905147" httpUri="https://zenodo.org/record/5905147/files/figure.png" pageId="8" pageNumber="150">3E, F</figureCitation>
). With lateral humps. Macrochaetae long and trichoid (
<figureCitation id="13712A27FFCFCD04520E0B76FB74FE89" box="[1117,1235,295,321]" captionStart="FIGURE 3" captionStartId="10.[151,250,1673,1697]" captionTargetBox="[151,1436,181,1649]" captionTargetId="figure-17@10.[151,1436,181,1649]" captionTargetPageId="10" captionText="FIGURE 3. Guipuzcosoma karinae sp. nov., ♂ paratype, habitus (ZMBN). A. Head, anterior view. B. Head and collum, lateral view. C. Left antenna, anterior view. D. Field with ommatidia, lateral view. E, F. Body rings 15–17, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A–C, E, F), 0.02 mm (D)." figureDoi="http://doi.org/10.5281/zenodo.5905147" httpUri="https://zenodo.org/record/5905147/files/figure.png" pageId="8" pageNumber="150">Fig. 3E, F</figureCitation>
). CIX (ring 15) = 1; MIX (ring15) ~ 2; PIX (ring15) = 0.4; MA (ring15) ~ 120˚. Prozonites with hexagonal tiles. Metazonites dorsally with scale-like structures, lateroventrally striated. Lateral parts of humps smooth.
</paragraph>
<paragraph id="8BF536A2FFCFCD0456940BC2FCE1FE19" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 distal setae. Paraprocts with 3+3 marginal setae in distal part.</paragraph>
<paragraph id="8BF536A2FFCFCD0456940B8AFCDBFDD1" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">LEG- PAIRS 1 AND 2. In both sexes with tarsal combs; femora with several long and robust setae; postfemora and tibiae each with several long and robust setae arranged in a group.</paragraph>
<paragraph id="8BF536A2FFCFCD0456940872FCE5FD61" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">
MALE SEXUAL CHARACTERS (
<figureCitation id="13712A27FFCFCD0457AD0872FDDEFDF5" box="[510,633,547,573]" captionStart="FIGURE 4" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[151,1435,181,1823]" captionTargetId="figure-19@11.[151,1435,181,1823]" captionTargetPageId="11" captionText="FIGURE 4. Guipuzcosoma karinae sp. nov., legs and gonopods. A. ♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB). B–F. ♂ paratype (ZMBN). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Leg-pair 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2" figureDoi="http://doi.org/10.5281/zenodo.5905149" httpUri="https://zenodo.org/record/5905149/files/figure.png" pageId="8" pageNumber="150">Fig. 4B–D</figureCitation>
). Leg-pair 2 with genital openings on coxae. Leg-pairs 3–7 enlarged compared to leg-pairs 1 and 2. Coxa 7 with mesal bump (
<figureCitation id="13712A27FFCFCD0455630816FC2CFDA9" box="[816,907,583,609]" captionStart="FIGURE 4" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[151,1435,181,1823]" captionTargetId="figure-19@11.[151,1435,181,1823]" captionTargetPageId="11" captionText="FIGURE 4. Guipuzcosoma karinae sp. nov., legs and gonopods. A. ♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB). B–F. ♂ paratype (ZMBN). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Leg-pair 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2" figureDoi="http://doi.org/10.5281/zenodo.5905149" httpUri="https://zenodo.org/record/5905149/files/figure.png" pageId="8" pageNumber="150">Fig. 4B</figureCitation>
). No other peculiarities on pregonopodal legs. Leg-pair 10 with coxal sacks and with short triangular trochanteral bumps (
<figureCitation id="13712A27FFCFCD0455B9083AFBE3FD4D" box="[1002,1092,619,645]" captionStart="FIGURE 4" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[151,1435,181,1823]" captionTargetId="figure-19@11.[151,1435,181,1823]" captionTargetPageId="11" captionText="FIGURE 4. Guipuzcosoma karinae sp. nov., legs and gonopods. A. ♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB). B–F. ♂ paratype (ZMBN). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Leg-pair 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2" figureDoi="http://doi.org/10.5281/zenodo.5905149" httpUri="https://zenodo.org/record/5905149/files/figure.png" pageId="8" pageNumber="150">Fig. 4C</figureCitation>
). Leg-pair 11 with coxal sacks and with short nippled subtriangular coxal process (
<figureCitation id="13712A27FFCFCD04548808DEFC91FD61" box="[731,822,655,681]" captionStart="FIGURE 4" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[151,1435,181,1823]" captionTargetId="figure-19@11.[151,1435,181,1823]" captionTargetPageId="11" captionText="FIGURE 4. Guipuzcosoma karinae sp. nov., legs and gonopods. A. ♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB). B–F. ♂ paratype (ZMBN). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Leg-pair 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2" figureDoi="http://doi.org/10.5281/zenodo.5905149" httpUri="https://zenodo.org/record/5905149/files/figure.png" pageId="8" pageNumber="150">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8BF536A2FFCFCD04569408E2FB2BFB69" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">
ANTERIOR GONOPODS (
<figureCitation id="13712A27FFCFCD04579108E2FD81FD05" box="[450,550,691,717]" captionStart="FIGURE 4" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[151,1435,181,1823]" captionTargetId="figure-19@11.[151,1435,181,1823]" captionTargetPageId="11" captionText="FIGURE 4. Guipuzcosoma karinae sp. nov., legs and gonopods. A. ♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB). B–F. ♂ paratype (ZMBN). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Leg-pair 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2" figureDoi="http://doi.org/10.5281/zenodo.5905149" httpUri="https://zenodo.org/record/5905149/files/figure.png" pageId="8" pageNumber="150">Figs 4E</figureCitation>
,
<figureCitation id="13712A27FFCFCD04546908E3FDDDFD05" box="[570,634,690,717]" captionStart="FIGURE 5" captionStartId="12.[151,250,1761,1785]" captionTargetBox="[201,1387,199,1738]" captionTargetId="figure-18@12.[201,1387,199,1738]" captionTargetPageId="12" captionText="FIGURE 5. Guipuzcosoma karinae sp. nov. A–H. ♂ paratype (ZMBN), I. ♀ paratype (NHMW MY10253). A–F. Anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G, H. Posterior gonopods, anterior and posterior views, respectively. Highlighed square in G shows a potential sperm. I. Vulvae, distal view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905151" httpUri="https://zenodo.org/record/5905151/files/figure.png" pageId="8" pageNumber="150">5A–F</figureCitation>
,
<figureCitation id="13712A27FFCFCD0454DD08E2FD74FD05" box="[654,723,691,717]" captionStart="FIGURE 6" captionStartId="13.[151,250,1757,1781]" captionTargetBox="[235,1358,194,1727]" captionTargetId="figure-19@13.[225,1363,181,1733]" captionTargetPageId="13" captionText="FIGURE 6. Guipuzcosoma karinae sp. nov. A–C. ♂ paratype (ZMBN), D–F. ♂ holotype (SMNG), G. ♀ paratype (NHMW MY10253). A–D.Anterior gonopods, anterior, lateral, posterior and postero-distal views, respectively. E, F. Posterior gonopods, anterior and lateral views, respectively. G. Vulvae, dital view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hairlike outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905153" httpUri="https://zenodo.org/record/5905153/files/figure.png" pageId="8" pageNumber="150">6A–D</figureCitation>
). Gonopodal sternum (
<emphasis id="B93EEAB0FFCFCD0455A108E3FBA5FD04" bold="true" box="[1010,1026,690,716]" pageId="8" pageNumber="150">S</emphasis>
) rather poorly developed, mainly membranous, distomedial parts forming round aperture and merging with gonopods. Coxites (
<emphasis id="B93EEAB0FFCFCD04528B0887FB5DFD38" bold="true" box="[1240,1274,726,752]" pageId="8" pageNumber="150">Cx</emphasis>
) more or less spoon-shaped, wide in lateral view, supporting two pairs of angiocoxites. Anteromedial pair (= angiocoxite 1) (
<emphasis id="B93EEAB0FFCFCD04532208ABFA34FCDC" bold="true" box="[1393,1427,762,788]" pageId="8" pageNumber="150">A1</emphasis>
) almost the same width in its entire length, with somewhat narrower proximal parts; distal process (
<emphasis id="B93EEAB0FFCFCD0452B4094FFABEFCF0" bold="true" box="[1255,1305,798,824]" pageId="8" pageNumber="150">dA1</emphasis>
) acuminate and bent laterad; posterior process (
<emphasis id="B93EEAB0FFCFCD0454700913FDF2FC94" bold="true" box="[547,597,834,860]" pageId="8" pageNumber="150">pA1</emphasis>
) well-developed, somewhat claw-like in lateral view, closely fitting against lateral sides of distal extension of colpocoxite (
<emphasis id="B93EEAB0FFCFCD0454900937FD54FC48" bold="true" box="[707,755,870,896]" pageId="8" pageNumber="150">deC</emphasis>
). Angiocoxites 2 (
<emphasis id="B93EEAB0FFCFCD04559C0937FC55FC48" bold="true" box="[975,1010,870,896]" pageId="8" pageNumber="150">A2</emphasis>
) divided into two branches, anterior branch (
<emphasis id="B93EEAB0FFCFCD0456A109DBFE84FC6C" bold="true" box="[242,291,906,932]" pageId="8" pageNumber="150">aA2</emphasis>
) consisting of several flagelloid processes with denticulated posterior margins distally; posterior branch (
<emphasis id="B93EEAB0FFCFCD0456F209FFFF74FC00" bold="true" box="[161,211,942,968]" pageId="8" pageNumber="150">pA2</emphasis>
) long, acuminate, with longitudinal striations, leans on flagelloid processes closing a canal between; both
<emphasis id="B93EEAB0FFCFCD04533909FFFA3AFC00" bold="true" box="[1386,1437,942,968]" pageId="8" pageNumber="150">pA1</emphasis>
and
<emphasis id="B93EEAB0FFCFCD04569A0983FF5CFC24" bold="true" box="[201,251,978,1004]" pageId="8" pageNumber="150">pA2</emphasis>
are directed mesad, and slightly posteriad with overlapping distal parts;
<emphasis id="B93EEAB0FFCFCD0452130983FBC4FC24" bold="true" box="[1088,1123,978,1004]" pageId="8" pageNumber="150">A2</emphasis>
outgrows all other parts of anterior gonopods. Colpocoxites (
<emphasis id="B93EEAB0FFCFCD04544B09A7FD8BFBD8" bold="true" box="[536,556,1014,1040]" pageId="8" pageNumber="150">C</emphasis>
) massive, strongly developed, chitinized; consist of smaller basal protrusions (
<emphasis id="B93EEAB0FFCFCD0456F20E4AFF1AFBFD" bold="true" box="[161,189,1051,1077]" pageId="8" pageNumber="150">bc</emphasis>
) roundish in lateral view and merged with main massive structures whose distal extensions (
<emphasis id="B93EEAB0FFCFCD0452970E4BFB53FBFC" bold="true" box="[1220,1268,1050,1076]" pageId="8" pageNumber="150">deC</emphasis>
) are in form of elongated horns strongly bent anteriad and closely fitting against mesal sides of
<emphasis id="B93EEAB0FFCFCD0452740E6FFBFEFB90" bold="true" box="[1063,1113,1086,1112]" pageId="8" pageNumber="150">pA1</emphasis>
; posterior mesal protrusions (
<emphasis id="B93EEAB0FFCFCD0456F20E33FF62FBB4" bold="true" box="[161,197,1122,1148]" pageId="8" pageNumber="150">pC</emphasis>
) rounded and laterally flattened with circular membranous remnants (
<emphasis id="B93EEAB0FFCFCD0455840E32FC5EFBB5" bold="true" box="[983,1017,1123,1149]" pageId="8" pageNumber="150">ms</emphasis>
) on lateral sides. Between
<emphasis id="B93EEAB0FFCFCD04537B0E33FAECFBB4" bold="true" box="[1320,1355,1122,1148]" pageId="8" pageNumber="150">A2</emphasis>
and
<emphasis id="B93EEAB0FFCFCD0453D20E33FA32FBB4" bold="true" box="[1409,1429,1122,1148]" pageId="8" pageNumber="150">C</emphasis>
, laterally, structure with large number of long, branched hair-like outgrowths (
<emphasis id="B93EEAB0FFCFCD0455A80ED6FBBEFB69" bold="true" box="[1019,1049,1159,1185]" pageId="8" pageNumber="150">ho</emphasis>
) presents.
</paragraph>
<paragraph id="8BF536A2FFCFCD0456940EFAFAF9FAB1" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">
POSTERIOR GONOPODS (
<figureCitation id="13712A27FFCFCD0457EA0EFAFDB6FB0D" box="[441,529,1195,1221]" captionStart="FIGURE 4" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[151,1435,181,1823]" captionTargetId="figure-19@11.[151,1435,181,1823]" captionTargetPageId="11" captionText="FIGURE 4. Guipuzcosoma karinae sp. nov., legs and gonopods. A. ♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB). B–F. ♂ paratype (ZMBN). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Leg-pair 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2" figureDoi="http://doi.org/10.5281/zenodo.5905149" httpUri="https://zenodo.org/record/5905149/files/figure.png" pageId="8" pageNumber="150">Figs 4F</figureCitation>
,
<figureCitation id="13712A27FFCFCD04544E0EFBFDC2FB0D" box="[541,613,1194,1221]" captionStart="FIGURE 5" captionStartId="12.[151,250,1761,1785]" captionTargetBox="[201,1387,199,1738]" captionTargetId="figure-18@12.[201,1387,199,1738]" captionTargetPageId="12" captionText="FIGURE 5. Guipuzcosoma karinae sp. nov. A–H. ♂ paratype (ZMBN), I. ♀ paratype (NHMW MY10253). A–F. Anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G, H. Posterior gonopods, anterior and posterior views, respectively. Highlighed square in G shows a potential sperm. I. Vulvae, distal view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905151" httpUri="https://zenodo.org/record/5905151/files/figure.png" pageId="8" pageNumber="150">5G, H</figureCitation>
,
<figureCitation id="13712A27FFCFCD0454220EFAFD08FB0D" box="[625,687,1195,1221]" captionStart="FIGURE 6" captionStartId="13.[151,250,1757,1781]" captionTargetBox="[235,1358,194,1727]" captionTargetId="figure-19@13.[225,1363,181,1733]" captionTargetPageId="13" captionText="FIGURE 6. Guipuzcosoma karinae sp. nov. A–C. ♂ paratype (ZMBN), D–F. ♂ holotype (SMNG), G. ♀ paratype (NHMW MY10253). A–D.Anterior gonopods, anterior, lateral, posterior and postero-distal views, respectively. E, F. Posterior gonopods, anterior and lateral views, respectively. G. Vulvae, dital view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hairlike outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905153" httpUri="https://zenodo.org/record/5905153/files/figure.png" pageId="8" pageNumber="150">6E, F</figureCitation>
). Gonopodal sternum (
<emphasis id="B93EEAB0FFCFCD0455E40EFBFC60FB0C" bold="true" box="[951,967,1194,1220]" pageId="8" pageNumber="150">S</emphasis>
) poorly developed, membranous, leaving both gonopods almost free; only chitinized part is pair of anterior concave bulges (
<emphasis id="B93EEAB0FFCFCD0452380E9EFB20FB21" bold="true" box="[1131,1159,1231,1257]" pageId="8" pageNumber="150">cb</emphasis>
) for accommodation of proximal (basal) parts of posterior gonopods. Coxotelopodites (
<emphasis id="B93EEAB0FFCFCD04550F0EA3FCDDFAC4" bold="true" box="[860,890,1266,1292]" pageId="8" pageNumber="150">Ct</emphasis>
) massive, without articulation, distally with long setae, while mesally, in upper half, with two striking acuminate horns (
<emphasis id="B93EEAB0FFCFCD04559D0F46FC79FAF9" bold="true" box="[974,990,1303,1329]" pageId="8" pageNumber="150">h</emphasis>
); basomesally with partially chitinized colpocoxites (
<emphasis id="B93EEAB0FFCFCD0457690F6BFEE9FA9C" bold="true" box="[314,334,1338,1364]" pageId="8" pageNumber="150">C</emphasis>
) both mainly with one long distal seta and slightly serrated mesal margins. Coxotelopodites subdistally with brownish or blackish pigment remnants (
<emphasis id="B93EEAB0FFCFCD04554F0F0EFC9FFAB1" bold="true" box="[796,824,1375,1401]" pageId="8" pageNumber="150">pr</emphasis>
). Distal half of
<emphasis id="B93EEAB0FFCFCD0455B90F0FFBAFFAB0" bold="true" box="[1002,1032,1374,1400]" pageId="8" pageNumber="150">Ct</emphasis>
slightly curved posteromesad.
</paragraph>
<paragraph id="8BF536A2FFCFCD0456940FD2FCEDFA55" blockId="8.[151,1437,151,1834]" box="[199,842,1411,1437]" pageId="8" pageNumber="150">
LEG- PAIR 3 IN FEMALE (
<figureCitation id="13712A27FFCFCD0457ED0FD2FDB2FA55" box="[446,533,1411,1437]" captionStart="FIGURE 4" captionStartId="11.[151,250,1847,1871]" captionTargetBox="[151,1435,181,1823]" captionTargetId="figure-19@11.[151,1435,181,1823]" captionTargetPageId="11" captionText="FIGURE 4. Guipuzcosoma karinae sp. nov., legs and gonopods. A. ♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB). B–F. ♂ paratype (ZMBN). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Leg-pair 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. Anterior gonopods, anterior view. F. Posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2" figureDoi="http://doi.org/10.5281/zenodo.5905149" httpUri="https://zenodo.org/record/5905149/files/figure.png" pageId="8" pageNumber="150">Fig. 4A</figureCitation>
). With lateral sternal lobes.
</paragraph>
<paragraph id="8BF536A2FFCFCD0456940FF6FB4DF9BD" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">
VULVAE (
<figureCitation id="13712A27FFCFCD0457600FF6FE2FFA09" box="[307,392,1446,1473]" captionStart="FIGURE 5" captionStartId="12.[151,250,1761,1785]" captionTargetBox="[201,1387,199,1738]" captionTargetId="figure-18@12.[201,1387,199,1738]" captionTargetPageId="12" captionText="FIGURE 5. Guipuzcosoma karinae sp. nov. A–H. ♂ paratype (ZMBN), I. ♀ paratype (NHMW MY10253). A–F. Anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G, H. Posterior gonopods, anterior and posterior views, respectively. Highlighed square in G shows a potential sperm. I. Vulvae, distal view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905151" httpUri="https://zenodo.org/record/5905151/files/figure.png" pageId="8" pageNumber="150">Fig. 5I</figureCitation>
,
<figureCitation id="13712A27FFCFCD0457C90FF6FE1AFA09" box="[410,445,1447,1473]" captionStart="FIGURE 6" captionStartId="13.[151,250,1757,1781]" captionTargetBox="[235,1358,194,1727]" captionTargetId="figure-19@13.[225,1363,181,1733]" captionTargetPageId="13" captionText="FIGURE 6. Guipuzcosoma karinae sp. nov. A–C. ♂ paratype (ZMBN), D–F. ♂ holotype (SMNG), G. ♀ paratype (NHMW MY10253). A–D.Anterior gonopods, anterior, lateral, posterior and postero-distal views, respectively. E, F. Posterior gonopods, anterior and lateral views, respectively. G. Vulvae, dital view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hairlike outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905153" httpUri="https://zenodo.org/record/5905153/files/figure.png" pageId="8" pageNumber="150">6G</figureCitation>
). With well-developed wrinkled postgenital membranous structure (
<emphasis id="B93EEAB0FFCFCD0452BF0FF6FA85FA09" bold="true" box="[1260,1314,1447,1473]" pageId="8" pageNumber="150">Pgm</emphasis>
) covering posterior and partially lateral parts of bursae (
<emphasis id="B93EEAB0FFCFCD0454C20F9AFD03FA2D" bold="true" box="[657,676,1483,1509]" pageId="8" pageNumber="150">B</emphasis>
). Lateral valves of
<emphasis id="B93EEAB0FFCFCD0455280F9AFC29FA2D" bold="true" box="[891,910,1483,1509]" pageId="8" pageNumber="150">B</emphasis>
with sharp denticles postriorly and spicule-like outgrowths in distoposterior half; distoanteriorly wrinkled with ca seven long setae. Mesal valves of
<emphasis id="B93EEAB0FFCFCD0453550FBEFABEF9C1" bold="true" box="[1286,1305,1519,1545]" pageId="8" pageNumber="150">B</emphasis>
completely merged, only with anterior fissure as evidence of former separation; anterior ⅔ wrinkled distally with long setae and convex lateral margins; posterior ⅓ separated from rest by shallow depression and represents striking unpaired medial structure (
<emphasis id="B93EEAB0FFCFCD04570E0C0AFE28F9BD" bold="true" box="[349,399,1627,1653]" pageId="8" pageNumber="150">mbs</emphasis>
) with two lateral lobes clearly separated by groove. Operculum (
<emphasis id="B93EEAB0FFCFCD04523A0C0BFBD8F9BC" bold="true" box="[1129,1151,1626,1652]" pageId="8" pageNumber="150">O</emphasis>
) bilobed.
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFCFCD0456940C2EFEE0F92A" pageId="8" pageNumber="150" type="discussion">
<paragraph id="8BF536A2FFCFCD0456940C2EFEE0F92A" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">
<emphasis id="B93EEAB0FFCFCD0456940C2EFE9CF951" bold="true" box="[199,315,1663,1689]" pageId="8" pageNumber="150">Remarks.</emphasis>
The structure that we think could be a sperm was observed in the base of the posterior gonopods (
<figureCitation id="13712A27FFCFCD04533C0C2EFF1BF975" captionStart="FIGURE 5" captionStartId="12.[151,250,1761,1785]" captionTargetBox="[201,1387,199,1738]" captionTargetId="figure-18@12.[201,1387,199,1738]" captionTargetPageId="12" captionText="FIGURE 5. Guipuzcosoma karinae sp. nov. A–H. ♂ paratype (ZMBN), I. ♀ paratype (NHMW MY10253). A–F. Anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G, H. Posterior gonopods, anterior and posterior views, respectively. Highlighed square in G shows a potential sperm. I. Vulvae, distal view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905151" httpUri="https://zenodo.org/record/5905151/files/figure.png" pageId="8" pageNumber="150">Fig. 5G</figureCitation>
, highlighted square). Such structures were observed on the vulvae of
<taxonomicName id="4C4A4D21FFCFCD0455910CF2FBB0F974" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[962,1047,1699,1725]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="8" pageNumber="150" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFCFCD0455910CF2FBB0F974" box="[962,1047,1699,1725]" italics="true" pageId="8" pageNumber="150">G. reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFCFCD04524D0CF2FBD0F975" bold="true" box="[1054,1143,1699,1725]" pageId="8" pageNumber="150">
<taxonomicNameLabel id="A20D57CBFFCFCD04524D0CF2FBD0F975" box="[1054,1143,1699,1725]" pageId="8" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(see below under remarks of this species).
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFCFCD0056940CBBFF61FF66" lastPageId="12" lastPageNumber="154" pageId="8" pageNumber="150" type="distribution">
<paragraph id="8BF536A2FFCFCD0456940CBBFDDCF8E2" blockId="8.[151,1437,151,1834]" pageId="8" pageNumber="150">
<emphasis id="B93EEAB0FFCFCD0456940CBBFEC6F8CC" bold="true" box="[199,353,1770,1796]" pageId="8" pageNumber="150">Distribution.</emphasis>
So far, known only from a few localities in the
<collectingRegion id="498EF840FFCFCD0455F40CBAFBD4F8CD" box="[935,1139,1771,1797]" country="Spain" name="Navarra" pageId="8" pageNumber="150">Navarre Province</collectingRegion>
, close to the border with Gipuzkoa Province (
<figureCitation id="13712A27FFCFCD04572D0D5EFE77F8E1" box="[382,464,1807,1833]" captionStart="FIGURE 12" captionStartId="20.[151,250,1979,2003]" captionTargetBox="[151,1436,1101,1956]" captionTargetId="figure-470@20.[151,1436,1101,1956]" captionTargetPageId="20" captionText="FIGURE 12. Distribution of the genus Guipuzcosoma Vicente &amp; Mauriès (1980)." figureDoi="http://doi.org/10.5281/zenodo.5905165" httpUri="https://zenodo.org/record/5905165/files/figure.png" pageId="8" pageNumber="150">Fig. 12</figureCitation>
, green circles).
</paragraph>
<caption id="DF35662AFFCECD0556C40DC6FD08F83F" ID-DOI="http://doi.org/10.5281/zenodo.5905145" ID-Zenodo-Dep="5905145" httpUri="https://zenodo.org/record/5905145/files/figure.png" pageId="9" pageNumber="151" startId="9.[151,250,1943,1967]" targetBox="[185,1403,181,1920]" targetPageId="9">
<paragraph id="8BF536A2FFCECD0556C40DC6FD08F83F" blockId="9.[151,1436,1943,2040]" pageId="9" pageNumber="151">
<emphasis id="B93EEAB0FFCECD0556C40DC6FEB5F867" bold="true" box="[151,274,1943,1967]" pageId="9" pageNumber="151">FIGURE 2.</emphasis>
<taxonomicName id="4C4A4D21FFCECD05574B0DC9FDA6F878" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[280,513,1944,1968]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="9" pageNumber="151" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFCECD05574B0DC9FDA6F878" box="[280,513,1944,1968]" italics="true" pageId="9" pageNumber="151">Guipuzcosoma karinae</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFCECD0554540DC9FDF1F878" bold="true" box="[519,598,1944,1968]" pageId="9" pageNumber="151">
<taxonomicNameLabel id="A20D57CBFFCECD0554540DC9FDF1F878" box="[519,598,1944,1968]" pageId="9" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus (ZMBN).
<emphasis id="B93EEAB0FFCECD0555440DC6FC97F867" bold="true" box="[791,816,1943,1967]" pageId="9" pageNumber="151">A.</emphasis>
♂ paratype, lateral view.
<emphasis id="B93EEAB0FFCECD0552620DC9FBEFF878" bold="true" box="[1073,1096,1944,1968]" pageId="9" pageNumber="151">B.</emphasis>
♂ paratype, anterior part of body, showing head and collum in dorsal view.
<emphasis id="B93EEAB0FFCECD0554660DEAFDE9F81B" bold="true" box="[565,590,1979,2003]" pageId="9" pageNumber="151">C.</emphasis>
♂ paratype, anterior part of body, lateral view.
<emphasis id="B93EEAB0FFCECD0552740DEDFBE7F81C" bold="true" box="[1063,1088,1980,2004]" pageId="9" pageNumber="151">D.</emphasis>
♀ paratype, midbody rings, dorsal view.
<emphasis id="B93EEAB0FFCECD0556800DB1FF4CF830" bold="true" box="[211,235,2016,2040]" pageId="9" pageNumber="151">E.</emphasis>
♂ paratype, lateral view.
<emphasis id="B93EEAB0FFCECD0557A30D8EFDC0F830" bold="true" box="[496,615,2015,2040]" pageId="9" pageNumber="151">Scale bars:</emphasis>
1 mm.
</paragraph>
</caption>
<caption id="DF35662AFFCDCD0656C40CD8FCA7F921" ID-DOI="http://doi.org/10.5281/zenodo.5905147" ID-Zenodo-Dep="5905147" httpUri="https://zenodo.org/record/5905147/files/figure.png" pageId="10" pageNumber="152" startId="10.[151,250,1673,1697]" targetBox="[151,1436,181,1649]" targetPageId="10">
<paragraph id="8BF536A2FFCDCD0656C40CD8FCA7F921" blockId="10.[151,1436,1673,1769]" pageId="10" pageNumber="152">
<emphasis id="B93EEAB0FFCDCD0656C40CD8FEB5F969" bold="true" box="[151,274,1673,1697]" pageId="10" pageNumber="152">FIGURE 3.</emphasis>
<taxonomicName id="4C4A4D21FFCDCD06574B0CD8FDA5F969" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[280,514,1673,1697]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="10" pageNumber="152" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFCDCD06574B0CD8FDA5F969" box="[280,514,1673,1697]" italics="true" pageId="10" pageNumber="152">Guipuzcosoma karinae</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFCDCD06545B0CD8FDFFF969" bold="true" box="[520,600,1673,1697]" pageId="10" pageNumber="152">
<taxonomicNameLabel id="A20D57CBFFCDCD06545B0CD8FDFFF969" box="[520,600,1673,1697]" pageId="10" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ paratype, habitus (ZMBN).
<emphasis id="B93EEAB0FFCDCD0655C40CD8FC08F969" bold="true" box="[919,943,1673,1697]" pageId="10" pageNumber="152">A.</emphasis>
Head, anterior view.
<emphasis id="B93EEAB0FFCDCD0652D40CD8FB38F969" bold="true" box="[1159,1183,1673,1697]" pageId="10" pageNumber="152">B.</emphasis>
Head and collum, lateral view.
<emphasis id="B93EEAB0FFCDCD0656870CFCFF4AF90D" bold="true" box="[212,237,1709,1733]" pageId="10" pageNumber="152">C.</emphasis>
Left antenna, anterior view.
<emphasis id="B93EEAB0FFCDCD0654450CFCFD88F90D" bold="true" box="[534,559,1709,1733]" pageId="10" pageNumber="152">D.</emphasis>
Field with ommatidia, lateral view.
<emphasis id="B93EEAB0FFCDCD0655F70CFCFC12F90D" bold="true" box="[932,949,1709,1733]" pageId="10" pageNumber="152">E</emphasis>
,
<emphasis id="B93EEAB0FFCDCD0655900CFCFC71F90D" bold="true" box="[963,982,1709,1733]" pageId="10" pageNumber="152">F.</emphasis>
Body rings 15–17, dorsal and lateral views, respectively.
<emphasis id="B93EEAB0FFCDCD06574E0C80FE33F921" bold="true" box="[285,404,1745,1769]" pageId="10" pageNumber="152">Scale bars:</emphasis>
0.1 mm (
<emphasis id="B93EEAB0FFCDCD0657A50C80FD80F921" bold="true" box="[502,551,1745,1769]" pageId="10" pageNumber="152">A–C</emphasis>
,
<emphasis id="B93EEAB0FFCDCD0654670C80FDE2F921" bold="true" box="[564,581,1745,1769]" pageId="10" pageNumber="152">E</emphasis>
,
<emphasis id="B93EEAB0FFCDCD0654010C80FDC6F921" bold="true" box="[594,609,1745,1769]" pageId="10" pageNumber="152">F</emphasis>
), 0.02 mm (
<emphasis id="B93EEAB0FFCDCD06548C0C80FD56F921" bold="true" box="[735,753,1745,1769]" pageId="10" pageNumber="152">D</emphasis>
).
</paragraph>
</caption>
<caption id="DF35662AFFCCCD0756C40D66FA3AF817" ID-DOI="http://doi.org/10.5281/zenodo.5905149" ID-Zenodo-Dep="5905149" httpUri="https://zenodo.org/record/5905149/files/figure.png" pageId="11" pageNumber="153" startId="11.[151,250,1847,1871]" targetBox="[151,1435,181,1823]" targetPageId="11">
<paragraph id="8BF536A2FFCCCD0756C40D66FA3AF817" blockId="11.[151,1437,1847,2015]" pageId="11" pageNumber="153">
<emphasis id="B93EEAB0FFCCCD0756C40D66FEB5F887" bold="true" box="[151,274,1847,1871]" pageId="11" pageNumber="153">FIGURE 4.</emphasis>
<taxonomicName id="4C4A4D21FFCCCD0757440D69FDA7F887" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[279,512,1847,1871]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="153" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFCCCD0757440D69FDA7F887" box="[279,512,1847,1871]" italics="true" pageId="11" pageNumber="153">Guipuzcosoma karinae</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFCCCD0754550D66FDF1F887" bold="true" box="[518,598,1847,1871]" pageId="11" pageNumber="153">
<taxonomicNameLabel id="A20D57CBFFCCCD0754550D66FDF1F887" box="[518,598,1847,1871]" pageId="11" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, legs and gonopods.
<emphasis id="B93EEAB0FFCCCD07557B0D66FCE6F887" bold="true" box="[808,833,1847,1871]" pageId="11" pageNumber="153">A.</emphasis>
♀ paratype, leg-pair 3, anterior view, arrow indicates lateral sternal lobe (IZB).
<emphasis id="B93EEAB0FFCCCD07570C0D0AFE37F8BB" bold="true" box="[351,400,1883,1907]" pageId="11" pageNumber="153">B–F.</emphasis>
♂ paratype (ZMBN).
<emphasis id="B93EEAB0FFCCCD0754280D0AFD35F8BB" bold="true" box="[635,658,1883,1907]" pageId="11" pageNumber="153">B.</emphasis>
Leg-pair 7, posterior view, arrow indicates coxal mesal bump.
<emphasis id="B93EEAB0FFCCCD0753770D0AFA9AF8BB" bold="true" box="[1316,1341,1883,1907]" pageId="11" pageNumber="153">C.</emphasis>
Leg-pair 10, anterior view, arrow indicates trochanteral bump.
<emphasis id="B93EEAB0FFCCCD0754E60D2EFD69F85F" bold="true" box="[693,718,1919,1943]" pageId="11" pageNumber="153">D.</emphasis>
Leg-pair 11, posterior view, arrow indicates coxal process.
<emphasis id="B93EEAB0FFCCCD0753790D2EFAE5F85F" bold="true" box="[1322,1346,1919,1943]" pageId="11" pageNumber="153">E.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B93EEAB0FFCCCD0757F10DF2FE12F873" bold="true" box="[418,437,1955,1979]" pageId="11" pageNumber="153">F.</emphasis>
Posterior gonopods, posterior view.
<emphasis id="B93EEAB0FFCCCD0755680DF2FBABF873" bold="true" box="[827,1036,1955,1979]" pageId="11" pageNumber="153">Abbreviations: A1:</emphasis>
angiocoxite 1;
<emphasis id="B93EEAB0FFCCCD0752E00DF2FB7DF873" bold="true" box="[1203,1242,1955,1979]" pageId="11" pageNumber="153">A2:</emphasis>
angiocoxite 2;
<emphasis id="B93EEAB0FFCCCD0753D20DF2FA3AF873" bold="true" box="[1409,1437,1955,1979]" pageId="11" pageNumber="153">C:</emphasis>
colpocoxite;
<emphasis id="B93EEAB0FFCCCD07574E0D96FEE7F817" bold="true" box="[285,320,1991,2015]" pageId="11" pageNumber="153">Ct:</emphasis>
coxotelopodite;
<emphasis id="B93EEAB0FFCCCD0757BC0D96FDB1F817" bold="true" box="[495,534,1991,2015]" pageId="11" pageNumber="153">Cx:</emphasis>
coxite;
<emphasis id="B93EEAB0FFCCCD07543F0D96FD24F817" bold="true" box="[620,643,1991,2015]" pageId="11" pageNumber="153">h:</emphasis>
mesal horns;
<emphasis id="B93EEAB0FFCCCD0755450D96FC9FF817" bold="true" box="[790,824,1991,2015]" pageId="11" pageNumber="153">pr:</emphasis>
pigment remnants;
<emphasis id="B93EEAB0FFCCCD07525B0D96FBB8F817" bold="true" box="[1032,1055,1991,2015]" pageId="11" pageNumber="153">S:</emphasis>
gonopodal sternum.
<emphasis id="B93EEAB0FFCCCD0752A80D96FAD3F817" bold="true" box="[1275,1396,1991,2015]" pageId="11" pageNumber="153">Scale bars:</emphasis>
0.2
</paragraph>
</caption>
<paragraph id="8BF536A2FFCBCD0056C40AC6FF61FF66" blockId="12.[151,198,151,174]" box="[151,198,151,174]" pageId="12" pageNumber="154">mm.</paragraph>
</subSubSection>
</treatment>
</document>
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